Fl or a of t h e A z or e s
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- Daisy Lee
- 5 years ago
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1 Fl Fl ora a of of t th he e A Az z ore es s A A Fi Fi el el d d G ui G ui d e d e SSeS es ces ce eoc nc ono d on d nnd ed ed enl eenl ar nl ar nl ar g ar geg eg d eg d eed d ed diti e ee diti oono no on nn MMaM a rm ar gr ag rg af rg af raf Paf PuP u Pbli u u sbli shs h es he rhe rs re s sr s
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3 Fl or a of t h e A z or e s A Fi el d G ui d e H a n n o S c h äf er S e c o n d e nl ar g e d e diti o n wit h d et er mi n ati o n k e y t o all g e n er a M a r g r af P u bli s h e r s
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5 Fl or a of t h e A z or e s A Fi el d G ui d e H a n n o S c h äf er M a r g r af P u bli s h e r s
6 S c h äf er, H a n n o Fl or a of t h e A z or e s A Fi el d G ui d e S e c o n d e nl ar g e d e diti o n Bi bli o gr afi s c h e I nf or m ati o n D er D e ut s c h e n Bi bli ot h e k Di e D e ut s c h e Bi bli ot h e k v er z ei c h n et di e s e P u bli k ati o n i n d er D e ut s c h e n N ati o n al bi bli o gr afi e; d et ailli ert e bi bli o gr afi s c h e D at e n si n d i m I nt er n et ü b er htt p:// d n b. d d b. d e a br uf b ar. C o v er s h o w s R u b u s h o c h st ett er or u m (l eft) a n d M yrt u s c o m m u ni s P h ot o o n t h e i n n er c o v er s h o w s S p ar a xi s b ul bif er a all p h ot o s b y H a n n o S c h äf er e x c e pt P er s e a i n di c a ( p a g e 7 5) F o e ni c ul u m v ul g ar e ( p a g e 1 4 3) C e dr o n ell a c a n ari e n si s ( p a g e 1 6 3) b y P et er S c h ö nf el d er wit h ki n d p er mi s si o n A b util o n t h e o p hr a sti ( p a g e 1 7 5) L o ni c er a j a p o ni c a ( p a g e 2 4 5) M oll u g o v erti cill at a ( p a g e 1 1 5) P u er ari a l o b at a ( p a g e 1 4 3) b y D a n T e n a gli a wit h ki n d p er mi s si o n. Pr o of r e a di n g Fr e d R u m s e y, L o n d o n Sil k e Bl et z er, N e w Y or k Pri nti n g & Bi n di n g t y p o dr u c k r o s s d orf , M ar gr af P u bli s h er s W ei k er s h ei m w w w. m ar gr af- v erl a g. d e All ri g ht s r e s er v e d. N o p art of t hi s p u bli c ati o n m a y b e r e pr o d u c e d or tr a n s mitt e d, i n a n y f or m or b y a n y m e a n s, wit h o ut writt e n p er mi s si o n b y t h e p u bli s h er. M a r g r af P u bli s h e r s C o- p u bli s h e d wit h B a c k h u y s P u bli s h er s
7 T a bl e of C o nt e nt I ntr o d u cti o n G e o gr a p h y a n d G e ol o g y Cli m at e Hi st or y A gri c ult ur e V e g et ati o n Fl or a: A z or e a n E n d e mi c s Fl or a: Pl a nt I n v a d er s A b br e vi ati o n s a n d L e g e n d Artifi ci al K e y t o t h e G e n er a of t h e A z or e a n Fl or a S p e ci e s i n s y st e m ati c or d er Gl o s s ar y R ef er e n c e s I n d e x... of s ci e ntifi c n a m e s of P ort u g u e s e n a m e s v
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9 I N T R O D U C TI O N G e o gr a p h y a n d G e ol o g y T h e A z or e a n Isl a n ds ar e a gr o u p of t hr e e ar c hi p el a g os c o m p os e d of ni n e i n h a bit e d isl a n ds i n t h e mi d dl e of t h e Atl a nti c O c e a n b et w e e n a n d n ort h er n l atit u d e, a n d b et w e e n a n d w est er n l o n git u d e. T h e s h ort est dist a n c e t o t h e E ur o p e a n ( P ort u g u es e) c o ast is a b o ut 1, k m, w h er e as t h e dist a n c e t o N e w F o u n dl a n d is a p pr o xi m at el y 1, k m. T h e e ast er n ar c hip el a g o c o nsists of C or v o t h e isl a n ds of S a nt a Fl or es M ari a, S ã o Mi g u el, a n d t h e u ni n h a bit e d F or mi g as r e ef. T h e c e ntr al ar c hi p el a g o c o nsists of t h e isl a n ds of Ter c eir a, S ã o J or g e, Gr a ci os a, Pi c o, a n d F ai al. T h e t w o s m all er isl a n ds, Il h a d as Fl or es a n d C or v o, m a k e u p t h e w est er n ar c hi p el a g o. T h e dist a n c e b et w e e n t h e e aster n- a n d w est er n m ost isl a n ds, S a nt a M ari a a n d Il h a d as Fl or es, r es p e cti v el y, is a b o ut k m. T h e hi g h est p e a ks ar e «M o nt a n h a d o Pi c o» ( Pi c o Isl a n d, 2, m), «Pi c o d a Var a» ( S ã o Mi g u el, 1, m) a n d «Pi c o d a Es p er a n ç a» ( S ã o J or g e, A ntill es 1, m). Alt o g et h er, t h e A z or e a n Isl a n ds c o v er m or e t h a n 2, k m 2 of l a n d. T o g et h er wit h t h e S el v a- g e ns, t h e M a d eir a gr o u p, t h e C a n ar y Isl a n ds, t h e C a p e Ver d e Isl a n ds, a n d p arts of S o ut h- w est er n M or o c c o, t h e y f or m t h e fl or al r e gi o n of M a c ar o n esi a. T h e A z or es ar e of v ol c a ni c ori gi n wit h o ut a n y p arts of c o nti n e nt al cr ust. T h e y ar e l o c at e d cl os e t o t h e mi d- Atl a nti c ri d g e, a sl o w m o vi n g s e a fl o or-s pr e a di n g z o n e, r e a c hi n g fr o m J a n M a y e n a n d I c el a n d i n t h e n ort h t o Trist a n d a C u n h a i n t h e s o ut h. F ai al B er m u d a B a h a m as Gr a ci os a S a o J or g e Pi c o T er c eir a k m A z or es S a o Mi g u el S a nt a M ari a M a d eir a C a n ari es C a p e V er d es k m
10 I n t h e A z or es r e gi o n, t h e E ur asi a n, t h e Afri c a n a n d t h e A m eri c a n pl at e ar e drifti n g a w a y fr o m e a c h ot h er, c a usi n g c o nst a nt t e ct o ni c dist ur b a n c es. M a g m a r e a c h es t h e u p p er p arts of t h e cr ust a n d es c a p es, f or mi n g v ol c a n o es. T o d a y, 1 2 a cti v e v ol c a n o es, fi v e of t h e m s u b m ari n e, c a n b e f o u n d i n t h e A z or es. E x a mi n ati o ns of r o c ks a n d f ossils r e v e al e d i m p ort a nt diff er e n c es i n a g e b et w e e n t h e isl a n ds of t h e ar e a. S a nt a M ari a, t h e e ast er n m ost isl a n d d e v el o p e d pr o b a bl y i n t h e Mi o c e n e, a b o ut 5 t o 1 4 milli o n y e ars a g o. T h e n ort h- e ast er n p arts of S ã o Mi g u el ar e s ai d t o b e 4 milli o n y e ars ol d, w h er e as t h e w est er n p arts s e e m t o h a v e e xist e d f or l ess t h a n 2 milli o n y e ars. I n t h e c e ntr al ar c hi p el a g o, Ter c eir a is pr o b a- bl y 3. 5 milli o n y e ars ol d, m u c h ol d er t h a n S ã o J or g e, F ai al, Pi c o, or Gr a ci os a. T h e l a v as of F ai al, S ã o J or g e, a n d Gr a ci os a w er e pr o d u c e d l ess t h a n 7 5 0, y e ars a g o. Wit h its 4 0, y e ars, Pi c o, t h e hi g h est m o u n- t ai n of P ort u g al, is t h e y o u n g est isl a n d of t h e gr o u p. I n g e n er al, t h e A z or es ar e m u c h y o u n g er t h a n m ost of t h e ot h er F ur n a s, o n e of t h e t o uri sti c c e nt er s o n S ã o Mi g u el, i s f a m o u s f or it s h ot s pri n g s 2
11 V ol c a ni c cr at er s a n d t o d a y i m p ort a nt n at ur e r e s er v e s: C al d eir a s o n Il h a d a s Fl or e s Atl a nti c isl a n ds: t h e M a d eir a gr o u p is s ai d t o b e 6 0 t o 7 0 milli o n y e ars ol d, a n d t h e a g e of t h e C a n ar y Isl a n ds r a n g es b et w e e n 3 a n d 2 5 milli o n y e ars. Wit hi n t h e e ast er n a n d c e ntr al ar c hip el a g o of t h e A z or es, v ol c a ni c a cti vit y m o v es c o nst a ntl y t o t h e w est. T o d a y, h ot s pri n gs ( e. g. «F ur n as d o E n x ofr e» o n Ter c eir a, «F ur n as» at S ã o Mi g u el) a n d e v e n er u pti o ns ar e c o nst a nt pr o ofs f or t h eir v ol c a ni c ori gi n. T h e r o c ks of t h e A z or es b el o n g t o t h e t hr e e m aj or gr o u ps: b as alt, a n d esit e, a n d tr a c h yt e. All t hr e e ar e v ol c a ni c er u pti v a a n d si mil ar i n c h e mi c al c o n- stit uti o n. A n al ys es s h o w as t h e m ai n c o m p o n e nts Si O 2, Al 2 O 3, a n d F e- O xi d es. T h e s oils ar e cl assifi e d as A n d os ol es. T his ki n d of s oil d e v el o ps fr o m air b or n e v ol c a ni c d e p osits, es p e- ci all y b asi c as h es. T h e y ar e ri c h i n a m or p h o us w e at h eri n g pr o d u cts a n d v ol c a ni c gl ass a n d t h er ef or e fi n el y c o ars e d, wit h l o w b ul k d e nsit y, b ut n e v ert h el ess, t h e y ar e us u all y q uit e sti c k y. T h e i o n e x c h a n g e c a p a cit y is r at h er hi g h, w h at m a k es t h es e s oils v er y f ertil e. T h e y ar e i nt e nsi v el y c ultiv at e d, w h e n o c c uri n g o n l ess st e e p ar e as. H o w e v er, s us c e pti bilit y t o er o- si o n, es p e ci all y o n sl o p es a n d i n c o ast al ar e as is r at h er hi g h. 3
12 Artifi ci al K e y t o t h e G e n er a of t h e A z or e a n Fl or a T his di c h ot o m o us k e y t o t h e g e n er a of t h e A z or e a n fl or a c o nt ai ns str o n gl y m o difi e d p arts of t h e k e ys t o t h e fl or a of M a d eir a ( Pr ess & S h ort ), t h e fl or a of t h e C a n ari es ( H o h e n est er & Welss ), a n d t h e fl or a of P ort u g al ( Fr a n c o ). T h e f a mil y c o n c e pt f oll o ws St e v e ns, P. F. ( ). A n gi os p er m P h yl o g e n y We bsit e. Versi o n 6, M a y [ a n d m or e or l ess c o n- ti n u o usl y u p d at e d si n c e] at htt p:// w w w. m o b ot. or g/ M O B O T/r es e ar c h/ A P w e b/. T o d et er mi n e t h e g e n us of a c ert ai n pl a nt, o n e h as t o c h o os e t a b. A (f er ns or f er n alli es) or B (s e e d pl a nts), t h e n c h o os e b et w e e n t w o alt er n ati v es ( e. g.: 1 or 1 *) a n d c o nti n u e at t h e i n di c at e d n u m b er u ntil g etti n g t o a g e n us. I n t h e c as e of Is o ët es a z ori c a, o n e w o ul d c h o os e t a b. A, t h e n Nr. 1 *, 2 *, a n d fi n all y arri v e at Nr Pl a nt r e pr o d u ci n g b y s p or es. M ostl y h er bs (f er ns, l y c o p o ds, h ors et ails) b ut als o t h e tr e e f er n S p h a er o pt eris c o o p eri. Ta b. A: F e r ns, h o rs et ails, cl u b m oss es, s pi k e m oss es a n d q uill w o rts S * Pl a nt wit h (s o m eti m es v er y s m all) s e e ds. H er bs, s hr u bs, tr e es a n d cli m b ers. Ta b. B: S e e d pl a nts S. 2 0 Ta b. A: F er n s, h or s et ail s, cl u b m o s s e s, s pi k e m o s s e s a n d q uill w ort s 1 St e ms j oi nt e d, h oll o w, disti n ctl y ri d g e d. L e a v es r e d u c e d t o s m all s c al es, i n w h orls at e a c h n o d e. - E q uis et u m ( E q uis et a c e a e) 1 * St e ms n ot j oi nt e d. L e a v es n ot i n w h orls. 2 2 L e a v es l o n g- p eti ol at e, wit h 4 l e afl ets. M arsil e a az ori c a ( M arsil e a c e a e) 2 * L e a v es n ot wit h 4 l e afl ets. 3 3 L e a v es fr o n d-li k e, diff er e nti at e d i nt o disti n ct l a mi n a a n d p eti ol e. 8 3 * L e a v es n ot diff er e nti at e d i nt o l a mi n a a n d p eti ol e. 4 4 Pl a nt a q u ati c, us u all y s u b m er g e d. L e a v es filif or m, gr ass-li k e, i n b as al r os ett e. Pl a nt fr a g m e nts oft e n a c c u m ul at e d o n t h e s h or es of s m all l a k es. Is o ët es az ori c a (Is o ët a c e a e) 4 * Pl a nt us u all y t err estri al. 5 5 L e a v es t hi n, fr a gil e, m m, wit h b as al li g ul e. Pl a nt m oss-li k e. S p or a n gi a i n c o ns pi c u o us. - S el a gi n ell a kr a ussi a n a ( S el a gi n ell a c e a e) 5 * L e a v es t hi c k, s c al e-li k e, m ostl y > 4 m m, li g ul e a bs e nt. S p or a n gi a oft e n i n l ar g e s pi k es
13 6 Pl a nt er e ct, r e g ul arl y di c h ot o m o usl y br a n c hi n g, f ertil e l e a v es n ot i n disti n ct, t er mi n al s pi k es. - H u p erzi a ( Ly c o p o di a c e a e) 6 * Pl a nt cr e e pi n g, irr e g ul arl y br a n c h e d, wit h as c e n di n g l at er al br a n c h es a n d t er mi n al f ertil e s pi k es. 7 7 L e a v es a p p e ari n g o p p osit e a n d d e c uss at e o n m ai n br a n c h es. - Di p h asi astr u m m a d eir e ns e ( Ly c o p o di a c e a e) 7 * L e a v es s pir all y, c ur v e d u p w ar ds. - Ly c o p o di ell a ( Ly c o p o di a c e a e) 8 ( 3) L e a v es fl es h y, di vi d e d i nt o a fl att e n e d, o v at e, e ntir e or pi n n atifi d, st eril e s e g m e nt a n d a l o n g, s pi k e-li k e s p or a n gi a- b e ari n g f ertil e s e g m e nt. Pl a nt < 2 0 c m. 9 8 * L e a v es n ot fl es h y, n ot wit h s pi k e-li k e f ertil e s e g m e nt St eril e s e g m e nt of l e af e ntir e, f ertil e s pi k e si m pl e. - O p hi o gl oss u m ( O p hi o gl oss a c e a e) 9 * St eril e s e g m e nt of l e af pi n n atifi d, wit h 3-9 p airs of pi n n a e. F ertil e s e g m e nt c o m p o u n d. - B otr y c hi u m l u n ari a ( O p hi o gl oss a c e a e) 1 0 ( 8) L e a v es 2- pi n n at e, bri g ht gr e e n. C e ntr al l e a v es wit h t er mi n al s p or a n gi a r e pl a ci n g t h e l a mi n a, y ell o w, t ur ni n g r e d dis h- br o w n. - Os m u n d a r e g alis ( Os m u n d a c e a e). 1 0 * C e ntr al l e a v es n ot wit h t er mi n al y ell o w t o br o w n, s p or a n gi a- b e ari n g p art F ertil e a n d st eril e l e a v es dissi mil ar * F ertil e a n d st eril e l e a v es si mil ar ( or diff eri n g o nl y i n si z e) L e a v es e ntir e, v el v et y. F ertil e l e a v es c ur v e d, s p or a n gi a c o v eri n g c o m pl et e l o w er s urf a c e. E pi p h yt e of t h e cl o u d f or est z o n e. - El a p h o gl oss u m s e mi c yli n dri c u m ( L o m ari o psi d a c e a e) 1 2 * L e a v es pi n n at e L e a v es 1- pi n n at e, t h e st eril e o n es ± p at e nt, t h e f ertil e o n es er e ct * L e a v es 2- t o 4- pi n n at e, t hi n. All l e a v es ± er e ct Pi n n a e e ntir e or cr e n at e. S ori li n e ar, m ar gi n al. - Bl e c h n u m s pi c a nt ( Bl e c h n a c e a e). 1 4 * Pi n n a e fi n el y s err at e. S ori o bl o n g, n ot m ar gi n al. - D o o di a c a u d at a ( Bl e c h n a c e a e) 1 5 ( 1 3) L e a v es 2- pi n n at e- pi n n atis e ct. A n n u al pl a nt of s h a d y pl a c es. - A n o gr a m m a l e pt o p h yll a ( A di a nt a c e a e) 1 5 * L e a v es 4- pi n n at e. P er e n ni al. - O n y c hi u m j a p o ni c u m ( Cr y pt o gr a m m a c e a e) 1 6 ( 1 1) L e a v es m e m br a n o us, fr a gil e, tr a nsl u c e nt, v er y t hi n. S p or a n gi a i n p o c k ets o n m ar gi n of t h e l e af. R hi z o m e v er y t hi n, l o n g- cr e e pi n g * L e a v es n ot m e m br a n o us. S p or a n gi a n ot i n p o c k ets. R hi z o m e t hi c k, cr e e pi n g or er e ct
14 1 7 L e a v es < 1 0 c m l o n g, pi n n at e. S p or a n gi a i n 2- v al v e d, m ar gi n al p o c k ets. E pi p h yt es of t h e cl o u d f or est z o n e. - H y m e n o p h yll u m ( H y m e n o p h yll a c e a e) 1 7 * L e a v es u p t o 5 0 c m l o n g, 2- t o 3- pi n n at e, tri a n g ul ar. S p or a n gi a i n fl as k-s h a p e d, m ar gi n al p o c k ets. Terr estri al f er n of r a vi n es a n d l a ur el f or est. - Tri c h o m a n es s p e ci os u m ( H y m e n o p h yll a c e a e) 1 8 ( 1 6) L e a v es e ntir e or p al m at el y l o b e d * L e a v es pi n n at e or pi n n atifi d L e a v es p al m at el y 3-5(- 7) l o b e d. - As pl e ni u m h e mi o nit es ( As pl e ni a c e a e) 1 9 * L e a v es e ntir e C o m m o n t err estri al f er n. L e a v es l a n c e ol at e, u p t o 6 0 c m l o n g, gl a br o us. S ori p air e d, li n e ar. - As pl e ni u m s c ol o p e n dri u m ( As pl e ni a c e a e) 2 0 * E xtr e m el y r ar e e pi p h yt es of t h e cl o u d f or est z o n e. L e a v es u p t o 1 0 c m l o n g. S ori or bi c ul ar L e af m ar gi n bl a c k, wit h gl a n d ul ar h airs. - Gr a m mitis az ori c a ( Gr a m miti d a c e a e) 2 1 * L e af m ar gi n gr e e n, gl a br o us or wit h si m pl e h airs. - C er a d e ni a j u n g er m a n ni oi d es ( Gr a m miti d a c e a e) 2 2 ( 1 8) L e a v es pi n n atifi d * L e a v es pi n n at e Yo u n g l e a v es bri g ht gr e e n, ± er e ct. S ori r o u n d e d. Pl a nt oft e n e pi p h yti c. - P ol y p o di u m az ori c u m ( P ol y p o di a c e a e) 2 3 * Yo u n g l e a v es or a n g e t o r e d, p e n d e nt. Pl a nt al w a ys t err estri al. - Bl e c h n u m a p p e n di c ul at u m ( Bl e c h n a c e a e) 2 4 ( 2 2) R hi z o m e wit h s m all b ul bs. - N e p hr ol e pis c or dif oli a ( Ol e a n dr a c e a e) 2 4 * R hi z o m e wit h o ut b ul bs S ori c o v er e d b y d efl e x e d m ar gi n of l e af * S ori n ot c o v er e d b y m ar gi n of l e af L e af-s e g m e nts l o n g, li n e ar-l a n c e ol at e. - Pt eris ( Pt eri d a c e a e) 2 6 * L e af-s e g m e nts n ot li n e ar-l a n c e ol at e R hi z o m e l o n g, s u bt err a n e a n. We e d of p ast ur es, gr o wi n g i n l ar g e st a n ds. - Pt eri di u m a q uili n u m ( H y p ol e pi d a c e a e) 2 7 * R hi z o m e s h ort, s u p erfi ci al. S m all pl a nts of w alls a n d r o c k cr e vi c es L e af-s e g m e nts fl a b ell at e, ± t o ot h e d. - A di a nt u m ( A di a nt a c e a e) 2 8 * L e af-s e g m e nts n ot fl a b ell at e, ± e ntir e L e a v es gl a br o us. - P ell a e a viri dis ( A di a nt a c e a e) 2 9 * L e a v es gl a n d ul ar or f ari n os e b e n e at h L e a v es wit h f e w gl a n d ul ar h airs or s c al es b e n e at h. Pl a nt of dr y b as alt w alls. S m elli n g of c o u m ari n. - C h eil a nt h es m a d er e nsis ( A di a nt a c e a e) 1 8
15 Pi ct ur e s h o ws Eri c a az ori c a S p e ci e s i n s y st e m ati c or d er
16 L y c o p o di a c e a e H u p erzi a d e nt at a ( H ert er) H ol u b 1 S y n.: H. s el a g o ( L.) B er n h. e x S c hr a n k & M art. ss p. d e nt at a ( H ert er) Val e nti n e c m. St. er e ct, di c h ot o m o usl y br a n c h e d. L vs. c a m m, li n e ar-l a n c e ol at e, si m pl e, a c ut e, m ar gi ns wit h u p t o 5 0 c o ns pi c u o us t e et h. S p or o p h ylls al o n g t h e br a n c h es, wit h s h ortl y st al k e d s p or a n gi a. G e m m a e a bs e nt. S p o. t etr a h e dr al. P. E : S c att er e d o n st e e p, oft e n s a n d y sl o p es a b o v e m. D : E n d e mi c t o M a d eir a & A z or es ( all e x c e pt M A R, G R A). H u p erzi a s u b ere ct a ( L o w e) Tar di e u 2 S y n.: H. s el a g o ( L.) B er n h. e x S c hr a n k & M art. ss p. s u b ere ct a ( L o w e) Fr a n c o & Vas c c m. St. er e ct, di c h ot o m o usl y br a n c h e d. L vs. c a m m, li n e ar-l a n c e ol at e, si m- pl e, a c ut e, m ar gi n us u all y e ntir e. A xils of u p p er l vs. oft e n b e ari n g g e m m a e. S p o. t etr a h e d- r al. P. E : R ar e i n h u mi d r e gi o ns a b o v e m. S o m eti m es e pi p h yti c o n ol d j u ni p er tr u n ks. D : E n d e mi c t o M a d eir a & A z or es ( all e x c e pt M A R, G R A). Ly c o p o di ell a c er n u a ( L.) Pi c. S er m. [ Pi n h eiri n h o, M us g o- d o- m at o] 3 S y n.: P al hi n h a e a c er n u a ( L.) Fr a n c o & Vas c.; L e pi d otis c er n u a ( L.) B e a u v. U p t o 5 0 c m. St. w o o d y at b as e, cr e e pi n g, oft e n s u bt err a n e a n, m u c h br a n c h e d wit h s o m e er e ct, tr e e-li k e s h o ots. L vs. c a m m, si m pl e, e ntir e a n d a c ut e. S p or o p h ylls d e nt at e, i n t er mi n al, p e n d e nt, o v oi d s pi k es of c m. S p o. t etr a h e dr al. P. E : L o c all y c o m m o n ar o u n d h ot s pri n gs, i n ol d Cr y pt o m eri a pl a nt ati o ns a n d o n h u mi d sl o p es us u all y a b o v e m. D : MI G, T E R, PI C, F AI, F L O. Wi d el y distri b ut e d t hr o u g h o ut t h e tr o pi cs a n d s u btr o pi cs. Ly c o p o di ell a i n u n d at a ( L.) H ol u b 4 S y n.: L e pi d otis i n u n d at a ( L.) O pi z c m. St. h er b a c e o us, us u all y s h ort a n d cr e e pi n g a b o v e t h e gr o u n d wit h f e w br a n c h es a n d si n gl e, er e ct s p or e b e ari n g s h o ots. L vs. c a m m, e ntir e, c ur v e d, a c ut e. S p or o p h ylls i n a si n gl e, t er mi n al s pi k e of c m. P. E : R ar e, i n c o ns pi c u o us pl a nt of m o orl a n ds, w et h e at hs a n d m ar gi ns of l a k es a b o v e m. D : MI G, PI C, F L O. Wi d el y distri b ut e d t hr o u g h o ut E ur o p e. 5 6
17
18 L y c o p o di a c e a e Di p h asi astr u m m a d eire ns e (J. H. Wil c e) H ol u b [ F et o- vi v a z] 1 S y n.: Di p h asi u m m a d eire ns e (J. H. Wil c e) R ot h m c m. St. cr e e pi n g, wit h as c e n di n g or er e ct, m u c h- br a n c h e d s h o ots. St eril e br a n c h es fl att e n e d. L vs. l a c e ol at e, a pi c es a c ut e t o r o u n d e d. F ertil e s h o ots wit h 3-6 er e ct s pi k es o n f or k e d p e d u n cl es of u p t o 8 c m. S p or a n gi a r e nif or m wit h tril et e s p o. P. E : R ar e s p e ci es of m o orl a n ds, j u ni p er f or ests a n d w et h e at hs us u all y a b o v e m. D : E n d e mi c t o M a d eir a & A z or es ( all e x c e pt M A R, G R A). S el a gi n ell a c e a e S el a gi n ell a kr a ussi a n a ( K u n z e) A. Br a u n 2 S y n.: S. az ori c a B a k er c m l o n g. St. fl att e n e d, cr e e pi n g, m u c h- br a n c h e d a n d m at-f or mi n g, a p p e ari n g arti c u- l at e d. L vs m m, di m or p hi c, d e nti c ul at e, a c ut e, as y m m etri c al, wit h li g ul e o n a d a xi al si d e. S p or o p h ylls i n t er mi n al, s essil e str o bili. S p or a n gi a h et er os p or o us, t h e l o w est wit h m e g as p or es. Mi cr os p or a n gi a n e ar t h e a p e x, mi cr os p or es y ell o w. P. E : Ver y c o m m o n i n w et a n d s h a d y pl a c es e x c e pt c o ast al r e gi o ns. T h o u g ht t o b e a n es c a p e d or n a m e nt al b ut r e c e ntl y, its s p or es w er e dis c o v er e d i n y e ar ol d A z or e a n l a k e s e di m e nts. D : All isl a n ds. S. d e nti c ul at a ( L.) S pri n g wit h st. u p t o 1 0 c m l o n g, wit h o ut arti c ul ati o ns, l at er al l vs. u p t o 2. 5 m m, is c o m m o n o n M a d eir a. Is o ët a c e a e Is o ët es az ori c a D uri e u e x Mil d e c m. A q u ati c, gr ass-li k e f er n. St. s h ort. L vs. i n a r os ett e, sl e n d er, wit h a l o n g, s u b ul at e li g ul e n e ar t h e s p at h ul at e l e af- b as e. S p or a n gi a s olit ar y, at t h e b as e of t h e l vs., p arti all y c o- v er e d wit h tiss u e. H et er os p or o us. M e g as p or es r eti c ul at e, t etr a h e dr al, µ m. Mi cr os p or es br o w n, s pi n os e. P.. E : R ar e a n d e n d a n g er e d s p e ci es of cr at er l a k es a b o v e m, w h er e it gr o ws i n s h all o w w at er t o a d e pt h of 3 m. It is r estri ct e d t o oli g otr o p hi c w at ers a n d t h us t hr e at e n e d b y a gri c ult ur al d e v el o p m e nt, es p e ci all y b y i ntr o d u cti o n of e x oti c fis h s p e ci es. D : E n d e mi c t o t h e A z or es ( e x c e pt M A R, MI G, G R A). Pr o b a bl y a cl os e r el ati v e of t h e N A m eri c a n I. t u c k er m a nii A. Br. 5 8
19 1 2 3
20 E q uis et a c e a e E q uis et u m ar v e ns e L c m. Er e ct, r hi z o m at o us h er b. St eril e st e ms gr e e n, m u c h- br a n c h e d wit h s u b er e ct br a n c h es a n d s m all, s c al y l e a v es. F ertil e st e ms si m pl e, c m, p al e br o w n wit h t er min al s p or e- b e ari n g str o bili, 1-4 c m. P. E: R ar e i n p ast ur es, o n r o a dsi d es a n d w ast e pl a c es. D: MI G. N ati v e t o E ur asi a, N A m eri c a. Si mil ar: t h e m u c h- br a n c h e d E. r a m osissi m u m D esf. wit h a pi c ul at e c o n es a n d c e ntr al h oll o w > h alf st. Ø ( M A R, MI G, T E R). E q uis et u m t el m at ei a E hr h. [ C a v ali n h a] c m. R hi z o m e cr e e pi n g, s u bt err a n e a n. St. er e ct, gr o o v e d a n d r e g ul arl y j oi nt e d. Br a n c h es of st eril e st. i n w h orls, c a. 2 0 c m l o n g, wit h 4 bi- a n gl e d ri d g es b e ari n g s a w-li k e t e et h, u nit e d i nt o a b as al s h e at h wit h t e et h. F ertil e st. a p p e ari n g b ef or e t h e st eril e, s h ort er, u n br a n c h e d, p al e br o w n. C o n es 4-7 c m, c o m p os e d of s p or a n gi o p h or es, b or n e at t h e e n ds of t h e m ai n st. H o m os p or o us, s p o. s p h eri c al. P. E : L o c all y c o m m o n o n ri v er b a n ks, al o n g t h e m ar gi ns of l a k es a n d o n w ast e pl a c es b et w e e n m. D : All e x c e pt G R A, PI C, C O R. N ati v e t o M A C, E ur asi a, N Afri c a, N A m eri c a. O p hi o gl oss a c e a e O p hi o gl oss u m az ori c u m C. Pr esl [ L ín g u a- d a- c o br a] c m. R hi z o m e s h ort, n ot cr e e pi n g. Fr o n ds 1-2(- 3) p er pl a nt, c o nsisti n g of u n di vi d e d, fl es h y l a mi n a a n d f ertil e s pi k e. L a mi n a u p t o 5 3 c m, s h ortl y st al k e d ( u p t o 5 c m), a c ut e t o o bt us e, n et- v ei n e d wit h fr e e v ei n- e n di n gs. S pi k e li n e ar, wit h 2 r o ws of s p or a n gi a. H o m os p or o us, s p o. t u b er c ul at e. P. Fr o n ds us u all y a p p e ari n g i n l at e s pri n g a n d s u m m er. E : Rare i n heat hla n d, a m o n gst Eric a s hr u bs a n d al o n g le va das bet wee n m. D : M A R, T E R, F AI, F L O. N ati v e t o M A C, W & C E ur o p e. O p hi o gl oss u m l usit a ni c u m L c m. R hi z o m e fl es h y, v er y s h ort. Fr o n ds 1-3(- 5) p er pl a nt, c o nsisti n g of u n di vi d e d fl es h y l a mi n a a n d f ertil e s pi k e. L a mi n a c m, s h ortl y st al k e d ( u p t o 2 c m), l a n c e ol at e, o bt us e, n et- v ei n e d wit h o ut fr e e v ei n- e n di n gs. S pi k e li n e ar, wit h 2 r o ws of s p or a n gi a. H o m os p or o us, s p o. ± s m o ot h. P. Fr o n ds us u all y a p p e ari n g i n wi nt er a n d e arl y s pri n g. E : L o c all y c o m m o n a n d i n gr e at n u m b ers i n s e as o n all y w et p ast ur es a n d o n w ast e gr o u n d es p e ci all y b el o w m. D : M A R, MI G, T E R, F AI, F L O. N ati v e t o M E D & W E ur o p e. 6 0
21
22 T This This This first T his first el el fi d el fi d gel d gui d gui d dg ui e gd uie ui dof de of de tof e th of of e th te htwil he he wil e wil dfl dfl wil dfl o odfl woers wers iw in ers tin th in e th n te hta he hz Ae e zor AAz or es Az or zes or or es pr es pr es opr ovi pr pr ovi do vi od vies ddes sdes sh es sh ort sh ort sh ort hd ort es d es dcri des dcri es es cri p- p- cri p- p- p- ti tio tio ns ti ti ons ons fo ns for ns for for f6 or or of 050 of 0 tof th of e th te hta he ha be e oab out ab out bo ut 1o ut 12 ut s 0 0s p 0 es pci se psci pe es pe ci es eci es ciof es es of of vof as of v as vcv as vc ul as as cul ar cul ar cul ar ul pl ar ar apl ants pla nts pla nts a fnts fo ufo un fo un d ou nd iu nd i nd in d ti n th in et hn te hth ehe e isl isl isl a an isl an ds, an ds, an ds, nc ds, co co mc ocm pl oom pl et mpl et mpl eet pl ed ete d etb ed eb y d d by by mb y mor y mor e mor me tor or e th e at he ta n hth an h4 an a4 0n n f4 0 full- 0 full- 0 f fc ull- col col oc ol co ur ol ol ur our po ur ph ur ph otp hot po hot gr ot ot gr ao gr oa pgr gr ap hs. ap hs. ap hs. p Nhs. Not Not Not on ot onl ot onl y onl oy tnl nl y th y et hy te hth ehe e m most most most mpr ost pr epr ev pr al pr ev eal v ev al ent val ent als ent es pnt nt es pci se psci pe es pe ci es eci ci bes but es but bals ut but als ut als o o rals o ar ro ar ro e rar e ara ar e an e an d e an d ua nd u nd un us d un us u nus nal us us uual plu al pl al pl ants pla nts pla nts aof nts of st of of st artli of st st st nartli ng ng bng eb ng ae bg aut be be aut y ea ut ay ut ar ut y ar y ear y ear ar eee ill ill ill ustr ill atustr eat ed ate d at aed an ed an d an d ad n es n d es d cri des dcri es es cri b eb cri ed, bbe d, be sd, es od, so ms oe sm oe om of me of me te of th of of et hte hm th em hf eem for m for m tf or ft hor or e th te hfirst th e he e first ti ti mti e. mti mme. me. e. M Mu cm umc h umu ch eu ch ec h me h mp eem ph masis ph mp hph h asis gis is vgi ev gie gi n vve n tve n to en to n eto en to en do e nd en dnmi de mi de c emi c smi c mi sp c es pc sci ep sci pe es pe ci es eci tes cies th es at th tat h hat gr hat gr at ogr ogr w gr ow on w ono w nw own own ho wh ower hwe her e her els ere e else e iels e in e in te in th in et hn te hth ehe e w worl worl worl d w d orl ad an d d an an d apl nd npl d ad pl ant pla nt pla int ai nt nt in via nv in ad vnv ers ad va ers dad ers dt ers th at th tat h chat co at co nstit co co o nstit ut ut e ut e ut a e ut a e se a eri sa eri sa seri os ous eri ous tous thr us thr et hr tat ehr at hr ete at et at o at to nt o tat no o at nur nat ur nat al at ur al ur ur eal ce al cos eec os ec y- os cy- os os y- y- y- st st est est ms est eems of ms of ms tof th of of e th te hta he hz e Ae zor AAz or Aes. zor zes. or or es. A es. A us A us A ef us A ef us us eful ef glul ul gloss ar ar oss y ary iar ar y in y in y Ei n ie n n E glis En En n h, glis h, Ph, Port h, Port Port up ug ort gu ues g ug es uge ues aes e an e an d e an d an d Gnd er Gd GGer mg er am er an mma nma nan n i intr intr intr oi od ntr od uoc du od es cu des uc uc les ces al es ay lla y rl ay er ay ae ry ar de re ers ad ea ers dad ers dt ers to o tto ht o e th o te hs th e ci sh e ci se sci entifi sci e cie e c ntifi c tc er tc er tc ter mi ter mi er nmi nol mi mi nol on ol no gol olo gy. og y. og y. gy. y. A An An ia in An i vn ial nv in al vnu val au val al bl uua bl e ua bl e abl aie bl ai e d ai e d fai ai d for d for d for all for all or all pl all pl apl ant pla nt pll ant al ont nt lo vlers ov lo ers vov ers iv iers n tin th in e th n te hta he hz Ae e zor AAz or Aes: zor zes: or or es: St es: Stu Stu dst St eu due nts, dud e de e tnts, et ae tca te tc eh aea ers, ch ac hch h fers, arm ers, m mm m rers, ar an rra ng ra ers, ng an gng g sers, ci s ci ses ci entists, sci e cie e ntists, t to to urists. to to o urists. f ffar- f ar- T This This This T shis es ce scs oe se co nec on d, co nd, on d, en d, enl d, enl ar enl ar enl gar nl eg ar ar ed gge d g ed e ditid d editi e eo diti on on io i on cl i n icl niu ncl un dcl cl es ud ues dud tdes th es e th te hfirst th e he e first d et d et der det er det er et mi er mi er nmi ati nmi mi nnati on on ati on ok n oe kn n ey kke y tk ey to ey to y all to tall o o all all g eg en gger nge er nea ner a ner of er a of a ta of tof hof e th te hth Ae he za e zor AAz or Ae zor za eor aor n eea n fl ea n fl an or fl n or fl a. fl or a. or or a. a. a M Ma amr ag M gar gaf r gaf gr af Pr af Pu af Pu bli Pu Publi su bli sh bli sh esh re shs re hsr eer srs s C Co- Co- pc o- Cp uo- o- pu bli pu bli pus usbli hbli esh ed she d hed wit ed wit d wit h wit h wit Bh h Ba h Bc abk ca Bak ch ac kh uck hy ukh s yu hus yuy Ps ys Pu s Pu bli Pu bli Pus usbli hbli sh ers her s her hs er er s ss c cmcmc mc mm
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