Geographical variations in carbon and nitrogen stable isotope ratios of Japanese anchovy, Engraulis japonicus

Size: px
Start display at page:

Download "Geographical variations in carbon and nitrogen stable isotope ratios of Japanese anchovy, Engraulis japonicus"

Transcription

1 Title Geographical variations in carbon and nitrogen stable isotope ratios of Japanese anchovy, Engraulis japonicus Hiroshige Tanaka 1, Akinori Takasuka 2,Ichiro Aoki 1, Seiji Ohshimo 3 and Yozo Wada 4 1 Graduate School of Agricultural and Life Sciences, The University of Tokyo 2 National Research Institute of Fisheries Science, Fisheries Research Agency 3 Seikai National Research Institute of Fisheries Science, Fisheries Research Agency 4 Kyoto Institute of Oceanic and Fishery Science

2 Introduction Introduction( 1) Japanese anchovy Engraulis japonicus Small pelagic fish which inhabit the waters of east Asia. Habitat varies depending on its stock size. (small: inshore water only; large: inshore and offshore water) Some characteristics of life history (e.g., spawning, growth) also vary between inshore and offshore waters. Feeding habit: zooplanktivorous (e.g., copepods) Structure of food web and ecosystem would vary among areas. Objective To compare the trophic ecology of Japanese anchovy among sea areas using the methods of δ 13 C and δ 15 N. Tools for the trophic study

3 Study areas M&M(1)study areas 45º N 40º N 35º N 30º N 125º E 130º E 135º E 140º E 145º E 150º E Tsushima Strait Tsushima current Japan Sea Kuroshio current Oyashio current Kuroshio- Oyashio Transition Tr-1 Ex-2 Ex-1 Kuroshio Extension Pacific Ocean Tr-2 (July and Nov. 2001) and (Nov. 1999, Feb, Mar, April and June 2000) (Aug. 2001) Kuroshio Extension Ex-1 (June 1997) and juvenile Ex-2 (May 1999) and Kuroshio-Oyashio Transition Tr-1 (June 1997) Tr-2 (June 1999) and

4 Methods M&M(2)anal ysis Stable isotope analysis Adults: White dorsal muscle was used for analysis. Larvae: Whole body was used for analysis. Lipid fraction was removed from all samples. 13 C/ 12 C and 15 N/ 14 N were analyzed by FINNIGAN MAT isotope ratio mass spectrometer. δ 13 Cor δ 15 N = ( R sample / R standard 1) * 1000 (%o) ( R = 13 C/ 12 C or 15 N/ 14 N ) R standard : Carbon: PDB (Peedee Belemnite) Nitrogen: N 2 in the atmosphere

5 Results(1)S L-d13C Results (1) δ 13 C δ 13 C - PDB (%o) Ex-1 Ex-2 Tr-1 Tr SL (mm)

6 Results(2)SL-d15N Results (2) δ 15 N 16 δ 15 N- air (%o) Ex-1 Ex-2 Tr-1 Tr SL (mm)

7 16 14 Results (3) δ 13 C-δ 15 N Kuroshio extension (Ex-1) Kuroshio extension (Ex-2) Kuroshio-Oyashio transition (Tr-1) Kuroshio-Oyashio transition (Tr-2) July 01 Results(3)C-N plot δ 15 N - Air (%o) June 99, juvenile Mar. 00 June 99 Apr. 00 Nov. 99 Aug. 01 June 00 July 01 Feb. 00 June 97, juvenile Nov June 99 May 99 juvenile June δ 13 C - PDB (%o)

8 16 14 Results (3) δ 13 C-δ 15 N Kuroshio extension (Ex-1) Kuroshio extension (Ex-2) Kuroshio-Oyashio transition (Tr-1) Kuroshio-Oyashio transition (Tr-2) Results(3)C-N plot δ 15 N - Air (%o) 12 10, juvenile, juvenile 8 juvenile δ 13 C - PDB (%o)

9 16 14 Results (3) δ 13 C-δ 15 N Kuroshio extension (Ex-1) Kuroshio extension (Ex-2) Kuroshio-Oyashio transition (Tr-1) Kuroshio-Oyashio transition (Tr-2) Hiroshima Bay (Takai et al. 2002) Osaka Bay (Mishima and Hoshika 2002) May 99 May 99 Apr. 98 Aug. 94 Apr. 98 July 01 Oct. 97 Data with Reference δ 15 N - Air (%o) June 99, juvenile Mar. 00 June 99 Apr. 98 Apr. 00 Nov. 99 Aug. 01 June 00 July 01 Feb. 00 June 97, juvenile Nov June 99 May 99 juvenile June δ 13 C - PDB (%o)

10 16 14 Results (3) δ 13 C-δ 15 N Kuroshio extension (Ex-1) Kuroshio extension (Ex-2) Kuroshio-Oyashio transition (Tr-1) Kuroshio-Oyashio transition (Tr-2) Hiroshima Bay (Takai et al. 2002) Osaka Bay (Mishima and Hoshika 2002) Data with Reference δ 15 N - Air (%o) 12 10, juvenile, juvenile 8 juvenile δ 13 C - PDB (%o)

11 Discussion What caused the variation in δ 13 C? Discussion(2) Stepwise enrichment: 0~1.5 %o per one trophic level δ 13 C is used to detect the carbon source of primary production. Low High Kuroshio-Oyashio transition and Kuroshio extension Sagami bay and other inland sea offshore inshore δ 13 C of phytoplankton: ca. 22 %o (France 1995) δ 13 C of benthic algae: ca. 17 %o (France 1995) offshore inshore carbon source (possibility) offshore inshore phytoplankton phytoplankton + benthic algae Benthic carbon may affect the pelagic food web because of the shallow depth.

12 Discussion What caused the variation in δ 15 N? Stepwise enrichment: 3~4 %o per one trophic level δ 15 N is often used as an index of trophic level. Discussion(3) Low High Kuroshio-Oyashio transition and Kuroshio extension Sagami bay ( >> ) and other inland sea offshore inshore Larvae Prey size may not drastically differ. Oceanographic conditions for primary producers such as nitrogen source (nitrate, ammonia) may differ. Adults Size range of available prey is broader. Ontogenetic shift of feeding habit may occur more clearly in Sagami bay (inshore water) than in Tr-2. (offshore water). (Availability of the prey at high trophic levels may differ.)

13 Summary Summary Large geographical variations in δ 13 C and δ 15 N were found in Japanese anchovy, especially between inshore and offshore waters. Inshore water Carbon source: phytoplankton + benthic algae Ontogenetic shift of feeding habits may occur more clearly. Offshore water Carbon source: mainly phytoplankton Ontogenetic shift of feeding habits may not occur drastically. Differences in food web structure may be reflected in the trophic ecology of Japanese anchovy.

14 δ 15 N - Air (%o) Kuroshio extension (Ex-1) Kuroshio extension (Ex-2) Kuroshio-Oyashio transition (Tr-1) Kuroshio-Oyashio transition (Tr-2) Hiroshima Bay (Takai et al. 2002) Osaka Bay (Mishima and Hoshika 2002) Offshore water Suppl. Summary of discussion : base Suppl.For summary Inshore water Ontogenetic shift of feeding habit phytoplankton + benthic algae 8 Ontogenetic shift is not clear 6 phytoplankton δ 13 C - PDB (%o)

15 δ 15 N - Air (%o) Kuroshio extension (Ex-1) Kuroshio extension (Ex-2) Kuroshio-Oyashio transition (Tr-1) Kuroshio-Oyashio transition (Tr-2) NORPAC Ex-2 NORPAC Tr-2 NORPAC Suppl. Data of plankton Suppl.data of plankton δ 13 C - PDB (%o)

16 suppl. Samples Suppl.sampl es Area Length (mm) N Date Sampling gear Sagami bay July 1, 2001 Shirasu trawl July 5, 2001 Shirasu trawl Nov. 8, 2001 Shirasu trawl July 2, 2001 Set net Wakasa bay Nov. 29, 1999 Set net Feb. 24, 2000 Set net Mar. 14, 2000 Set net April 6, 2000 Set net June 15, 2000 Set net Tsushima strait Aug. 26, 2001 Midwater trawl Aug. 27, 2001 Midwater trawl Aug. 29, 2001 Midwater trawl Kuroshio-extention (54) June 11, 1997 (ex-1) FMT, ORI (19) May 22, 1999 (ex-2) FMT, ORI Kuroshio-Oyshio transition (29) 20 (27) June 13, 1997 (tr-1) June 1, 1999 (tr-2) FMT, ORI FMT, ORI

Changes in spatial distribution of chub mackerel under climate change: the case study using Japanese purse seine fisheries data in the East China Sea

Changes in spatial distribution of chub mackerel under climate change: the case study using Japanese purse seine fisheries data in the East China Sea Changes in spatial distribution of chub mackerel under climate change: the case study using Japanese purse seine fisheries data in the East China Sea Tohya Yasuda, Ryuji Yukami, Seiji Ohshimo Seikai National

More information

Oceanographic conditions in the JARPNII survey area from 2000 to 2013 using FRA-ROMS data

Oceanographic conditions in the JARPNII survey area from 2000 to 2013 using FRA-ROMS data Oceanographic conditions in the JARPNII survey area from 2000 to 2013 using FRA-ROMS data MAKOTO OKAZAKI 1, MASACHIKA MASUJIMA 1, HIROTO MURASE 2 AND KENJI MORINAGA 2 1 National Research Institute of Fisheries

More information

FISHERIES OCEANOGRAPHY Fish. Oceanogr. 20:2, , 2011

FISHERIES OCEANOGRAPHY Fish. Oceanogr. 20:2, , 2011 FISHERIES OCEANOGRAPHY Fish. Oceanogr. :2, 114 124, 11 Environmental variability and growth histories of larval Japanese sardine (Sardinops melanostictus) and Japanese anchovy (Engraulis japonicus) near

More information

Future of Kuroshio/Oyashio ecosystems: an outcome of the CFAME Task Team and WG20

Future of Kuroshio/Oyashio ecosystems: an outcome of the CFAME Task Team and WG20 Future of Kuroshio/Oyashio ecosystems: an outcome of the CFAME Task Team and WG20 Tsushima Current Oyashio Kuroshio/Oyashio Transition Zone (KOTZ) Kuroshio Kuroshio Extension Akihiko Yatsu, Sanae Chiba,

More information

2001 State of the Ocean: Chemical and Biological Oceanographic Conditions in the Newfoundland Region

2001 State of the Ocean: Chemical and Biological Oceanographic Conditions in the Newfoundland Region Stock Status Report G2-2 (2) 1 State of the Ocean: Chemical and Biological Oceanographic Conditions in the Background The Altantic Zone Monitoring Program (AZMP) was implemented in 1998 with the aim of

More information

Persistence of prey hot spots in southeast Alaska

Persistence of prey hot spots in southeast Alaska Persistence of prey hot spots in southeast Alaska Scott M. Gende National Park Service, Glacier Bay Field Station, 3100 National Park, Juneau, Alaska, USA; Scott_Gende Gende@nps.gov Michael Sigler National

More information

Cambridge International Examinations Cambridge International Advanced Subsidiary and Advanced Level

Cambridge International Examinations Cambridge International Advanced Subsidiary and Advanced Level Cambridge International Examinations Cambridge International Advanced Subsidiary and Advanced Level *2554656732* MARINE SCIENCE 9693/01 Paper 1 AS Structured Questions October/November 2017 1 hour 30 minutes

More information

Population dynamics model of Copepoda (Neocalanus cristatus) in the northwestern subarctic Pacific

Population dynamics model of Copepoda (Neocalanus cristatus) in the northwestern subarctic Pacific Population dynamics model of Copepoda (Neocalanus cristatus) in the northwestern subarctic Pacific Zooplankton production symposium S1-3319 3, May, 27, Hiroshima Michio J. Kishi 1)2) and Takeshi Terui

More information

Hiromichi Igarashi 1,2, Y. Ishikawa 1, T. Wakamatsu 1, Y. Tanaka 1, M. Kamachi 1,3, N. Usui 3, M. Sakai 4, S. Saitoh 2 and Y.

Hiromichi Igarashi 1,2, Y. Ishikawa 1, T. Wakamatsu 1, Y. Tanaka 1, M. Kamachi 1,3, N. Usui 3, M. Sakai 4, S. Saitoh 2 and Y. PICES2015,Qingdao Oct.20,2015 Relationships between ocean conditions and interannual variability of habitat suitability index (HSI) distribution for neon flying squid in central North Pacific examined

More information

Comparing walleye pollock dynamics across the Bering Sea and adjacent areas

Comparing walleye pollock dynamics across the Bering Sea and adjacent areas Comparing walleye pollock dynamics across the Bering Sea and adjacent areas Franz J. Mueter 1, Mikhail A. Stepanenko 2 Anatoly V. Smirnov 2, and Orio Yamamura 3 1 School of Fisheries and Ocean Sciences,

More information

FISHERIES OCEANOGRAPHY Fish. Oceanogr. 18:2, , 2009

FISHERIES OCEANOGRAPHY Fish. Oceanogr. 18:2, , 2009 FISHERIES OCEANOGRAPHY Fish. Oceanogr. 18:2, 118 133, 2009 Transport and environmental temperature variability of eggs and larvae of the Japanese anchovy (Engraulis japonicus) and Japanese sardine (Sardinops

More information

over the Northern West Florida Shelf from SeaWinds and ASCAT

over the Northern West Florida Shelf from SeaWinds and ASCAT Interannual Variability of Synoptic Scale Winds over the Northern West Florida Shelf from SeaWinds and ASCAT Steve Morey Mark Bourassa Austin Todd COAPS/FSU This work is sponsored by the NASA Ocean Vector

More information

Spatial dynamics of small pelagic fish in the California Current system on the regime time-scale. Parallel processes in other species-ecosystems.

Spatial dynamics of small pelagic fish in the California Current system on the regime time-scale. Parallel processes in other species-ecosystems. PICES/GLOBEC Symposium Honolulu, Hawaii April 19-21, 2006 Spatial dynamics of small pelagic fish in the California Current system on the regime time-scale. Parallel processes in other species-ecosystems.

More information

Ocean acidification in NZ offshore waters

Ocean acidification in NZ offshore waters Ocean acidification in NZ offshore waters Cliff Law NIWA Review susceptible groups in NZ offshore waters Examples from international research of responses Identify potential impacts in the NZ EEZ Plankton

More information

Trophic position of Mediterranean bluefin tuna larvae estimated by different stable isotope analyses

Trophic position of Mediterranean bluefin tuna larvae estimated by different stable isotope analyses Trophic position of Mediterranean bluefin tuna larvae estimated by different stable isotope analyses Amaya Uriarte, R. Laiz-Carrion, J. Llopiz, J.M. Quintanilla, F. Alemany and A. García Introduction Atlantic

More information

SUPPLEMENTARY INFORMATION

SUPPLEMENTARY INFORMATION doi:10.1038/nature25504 Contents: Supplementary Text: Supplementary results and discussion: 1. Evaluations of the S-map method, 2. Ecological interpretations of the detected interspecific interactions

More information

PICES W3 [D-504], Sep 22, 2017, 11:40-12:05

PICES W3 [D-504], Sep 22, 2017, 11:40-12:05 PICES W3 [D-504], Sep 22, 2017, 11:40-12:05 Individual-based model of chub mackerel (Scomber japonicus) covering from larval to adult stages to project climate-driven changes in their spatial distribution

More information

Arctic climate change and effects on the ecosystems

Arctic climate change and effects on the ecosystems Arctic climate change and effects on the ecosystems NalânKoç Centre for Ice, Climate and Ecosystems (ICE) Norwegian Polar Institute Nalan.koc@npolar.no The Arctic Pacific Ocean Main inflow Main outflow

More information

Fukushima nuclear power plant damaged by M9 Earthquake with some focus on ocean

Fukushima nuclear power plant damaged by M9 Earthquake with some focus on ocean Fukushima nuclear power plant damaged by M9 Earthquake with some focus on ocean Moto Ikeda (Hokkaido Univ. & JAMSTEC) Oceanographic Society of Japan, Earthquake Disaster Working Group Magnitude-9 earthquake

More information

Oyashio & Kuroshio. Ocean and Climate Changes [113]

Oyashio & Kuroshio. Ocean and Climate Changes [113] Oyashio & Kuroshio Ocean and Climate Changes [113] highlights The Kuroshio and Oyashio, both of which are the western boundary currents in the North Pacific, have great influence on fisheries and climate

More information

Habitat Suitability for Forage Fishes in Chesapeake Bay

Habitat Suitability for Forage Fishes in Chesapeake Bay Habitat Suitability for Forage Fishes in Chesapeake Bay Aug 2017 Jul 2019 Mary C Fabrizio Troy D Tuckey Aaron J Bever Michael L MacWilliams 21 June 2018 Photo: Chesapeake Bay Program Motivation Production

More information

Numerical studies focusing on the early life stage of Pacific bluefin tuna (Thunnus orientalis)

Numerical studies focusing on the early life stage of Pacific bluefin tuna (Thunnus orientalis) 水研センター研報, 第 38 号,121 125, 平成 26 年 Bull. Fish. Res. Agen. No. 38, 51-55, 2014 121 Numerical studies focusing on the early life stage of Pacific bluefin tuna (Thunnus orientalis) Masachika MASUJIMA *1, Yoshiki

More information

5 Stable and radioactive isotopes

5 Stable and radioactive isotopes 5 Stable and radioactive isotopes Outline 1 Stable isotopes Measuring stable isotopic abundances Equilibrium isotope effects Kinetic isotope effects Rayleigh distillation Isotopes: a mainstay of chemical

More information

H. Igarashi 1,2, Y. Ishikawa 1, T. Wakamatsu 1, Y. Tanaka 1, M. Kamachi 1, N. Usui 3, M. Sakai 4, S. Saitoh 2, and Y. Imamura 5. 8Nov.

H. Igarashi 1,2, Y. Ishikawa 1, T. Wakamatsu 1, Y. Tanaka 1, M. Kamachi 1, N. Usui 3, M. Sakai 4, S. Saitoh 2, and Y. Imamura 5. 8Nov. PICES2016, San Diego 8Nov.,2016 Adaptive improvement of habitat suitability index (HSI) model for neon flying squid in central North Pacific by using ocean forecasts and real-time fishery reports H. Igarashi

More information

Five years monitoring activity on radioactive cesium in seawater after the Fukushima Dai-ichi Nuclear Power Plant Accident

Five years monitoring activity on radioactive cesium in seawater after the Fukushima Dai-ichi Nuclear Power Plant Accident Five years monitoring activity on radioactive cesium in seawater after the Fukushima Dai-ichi Nuclear Power Plant Accident H. Kaeriyama 1*, D. Ambe 1, Y. Shigenobu 1, S. Miki 1, T. Morita 1, H. Sugisaki

More information

Food and Feeding ------------------------------------------------------------------------------------ 4.1. Introduction Fish like any other organisms depends on the energy received from its food to perform

More information

Environmental forcing on forage fish and apex predators in the California Current: Results from a fully coupled ecosystem model

Environmental forcing on forage fish and apex predators in the California Current: Results from a fully coupled ecosystem model Environmental forcing on forage fish and apex predators in the California Current: Results from a fully coupled ecosystem model Jerome Fiechter Institute of Marine Sciences, UC Santa Cruz Co-authors: K.

More information

Vancouver Lake Biotic Assessment

Vancouver Lake Biotic Assessment Vancouver Lake Biotic Assessment Washington State University Vancouver Aquatic Ecology Laboratory Dr. Stephen M. Bollens Dr. Gretchen Rollwagen-Bollens Co-Directors Problem: Noxious cyanobacteria blooms

More information

The abundance of juvenile yellowtail (Seriola quinqueradiata) near the Kuroshio: the roles of drifting seaweed and regional hydrography

The abundance of juvenile yellowtail (Seriola quinqueradiata) near the Kuroshio: the roles of drifting seaweed and regional hydrography FISHERIES OCEANOGRAPHY Fish. Oceanogr. 15:5, 351 362, 2006 The abundance of juvenile yellowtail (Seriola quinqueradiata) near the Kuroshio: the roles of drifting seaweed and regional hydrography SHINJI

More information

GAMINGRE 8/1/ of 7

GAMINGRE 8/1/ of 7 FYE 09/30/92 JULY 92 0.00 254,550.00 0.00 0 0 0 0 0 0 0 0 0 254,550.00 0.00 0.00 0.00 0.00 254,550.00 AUG 10,616,710.31 5,299.95 845,656.83 84,565.68 61,084.86 23,480.82 339,734.73 135,893.89 67,946.95

More information

6th Grade Science Scope & Sequence Darul Arqam Houston, Texas 1st Quarter (43 Days) Resources:

6th Grade Science Scope & Sequence Darul Arqam Houston, Texas 1st Quarter (43 Days) Resources: 1st Quarter (43 Days) 1st: Aug 9-11 (3 days) What is Science? Science in Action Models in Science 6. Define science and identify questions that science cannot answer. Compare and contrast theories and

More information

Investigating the contribution of allochthonous subsidies to kelp forests in central California

Investigating the contribution of allochthonous subsidies to kelp forests in central California Investigating the contribution of allochthonous subsidies to kelp forests in central California melissa m foley UCSC Institute of Marine Science and Center for Ocean Solutions system connectivity rivers

More information

Ice matters. Arctic and Antarctic under-ice communities linking sea ice with the pelagic food web

Ice matters. Arctic and Antarctic under-ice communities linking sea ice with the pelagic food web Ice matters Arctic and Antarctic under-ice communities linking sea ice with the pelagic food web Hauke Flores, J. A. van Franeker, C. David, B. Lange, V. Siegel, E. A. Pakhomov, U. Bathmann... Hauke.flores@awi.de

More information

Name Period Part I: INVESTIGATING OCEAN CURRENTS: PLOTTING BUOY DATA

Name Period Part I: INVESTIGATING OCEAN CURRENTS: PLOTTING BUOY DATA Name Period Part I: INVESTIGATING OCEAN CURRENTS: PLOTTING BUOY DATA INTRODUCTION: Ocean currents are like huge rivers in the sea. They carry drifting organisms, vital dissolved chemical nutrients and

More information

Advice September 2012

Advice September 2012 9.4.23 Advice September 2012 ECOREGION STOCK Widely distributed and migratory stocks European seabass in the Northeast Atlantic Advice for 2013 ICES advises on the basis of the approach to data-limited

More information

Arctic marine mammals and climate change. SIO 133 Marine Mammal Biology April 26, 2017

Arctic marine mammals and climate change. SIO 133 Marine Mammal Biology April 26, 2017 Arctic marine mammals and climate change SIO 133 Marine Mammal Biology April 26, 2017 The Arctic Ocean: Extreme seasonal cycle of light and sea ice Arctic Ocean Food Webs Short growing season Intense

More information

Buoyancy and vertical distribution of Pacific mackerel eggs and larvae and its implication to the recruitment variability.

Buoyancy and vertical distribution of Pacific mackerel eggs and larvae and its implication to the recruitment variability. 2015 Oct. 20 BIO-P 10586 Buoyancy and vertical distribution of Pacific mackerel eggs and larvae and its implication to the recruitment variability. Hwa Hyun Lee 1, Sukyung Kang 2, Kyungmi Jung 2, Suam

More information

Mapping and Valuing Pelagic Ecosystem Services in the Lesser Sunda Ecoregion Preliminary Results on the Manta Rays Pelagic Fisheries for Tourism

Mapping and Valuing Pelagic Ecosystem Services in the Lesser Sunda Ecoregion Preliminary Results on the Manta Rays Pelagic Fisheries for Tourism Mapping and Valuing Pelagic Ecosystem Services in the Lesser Sunda Ecoregion Preliminary Results on the Manta Rays Pelagic Fisheries for Tourism Luky Adrianto. Yudi Wahyudin, Alan F Koropitan and Benny

More information

to climatic regime shifts?

to climatic regime shifts? Can we explain and predict stock fluctuations of Japanese common squid, Todarodes pacificus related to climatic regime shifts? Yasunori SAKURAI 1, Jun YAMAMOTO 2, Ken MORI 3 Tsuneo GOTO 4 and Hideaki KIDOKORO

More information

Offshore entrainment of anchovy larvae and its implication for their survival in a frontal region of the Kuroshio

Offshore entrainment of anchovy larvae and its implication for their survival in a frontal region of the Kuroshio MARINE ECOLOGY PROGRESS SERIES Vol. 248: 237 244, 2003 Published February 20 Mar Ecol Prog Ser Offshore entrainment of anchovy larvae and its implication for their survival in a frontal region of the Kuroshio

More information

ESP Process Flow. 2/27/2012 Environmental Studies Program 1

ESP Process Flow. 2/27/2012 Environmental Studies Program 1 ESP Process Flow 2/27/2012 Environmental Studies Program 1 Studies Management Process Timeline Oct Nov Dec Jan Feb Mar Apr May Jun Jul Aug SDP Guidance Stakeholder Input Due SDP Preparation Draft SDPs

More information

Oct.23,2014. PICES2014,Yeosu

Oct.23,2014. PICES2014,Yeosu PICES2014,Yeosu Oct.23,2014 A multi-model ensemble prediction of habitat suitability index (HSI) models for neon flying squid in central North Pacific by using 3-D ocean data assimilation product Hiromichi

More information

Relationship between temperature and fluctuations in sandfish catch (Arctoscopus japonicus) in the coastal waters off Akita Prefecture

Relationship between temperature and fluctuations in sandfish catch (Arctoscopus japonicus) in the coastal waters off Akita Prefecture ICES Journal of Marine Science, 54: 1 12. 1997 Relationship between temperature and fluctuations in sandfish catch (Arctoscopus japonicus) in the coastal waters off Akita Prefecture Kazumi Sakuramoto,

More information

Trophic Ecology of Summer Flounder in Lower Chesapeake Bay Inferred from Stomach Content and Stable Isotope Analyses

Trophic Ecology of Summer Flounder in Lower Chesapeake Bay Inferred from Stomach Content and Stable Isotope Analyses Transactions of the American Fisheries Society 140:1240 1254, 2011 C American Fisheries Society 2011 ISSN: 0002-8487 print / 1548-8659 online DOI: 10.1080/00028487.2011.618364 ARTICLE Trophic Ecology of

More information

by co-chairs Lisa Hendrickson, Hassan Moustahfid and Alexander Arkhipkin

by co-chairs Lisa Hendrickson, Hassan Moustahfid and Alexander Arkhipkin Summary of an FAO workshop regarding the effects of climate variability and change on short-lived species and their forecasting with a focus on squid stocks and Western Boundary Currents by co-chairs Lisa

More information

POPULATION DYNAMICS AND PREDATION IMPACT OF THE INTRODUCED CTENOPHORE MNEMIOPSIS LEIDYI IN THE GULLMARS FJORD, WEST COAST OF SWEDEN

POPULATION DYNAMICS AND PREDATION IMPACT OF THE INTRODUCED CTENOPHORE MNEMIOPSIS LEIDYI IN THE GULLMARS FJORD, WEST COAST OF SWEDEN POPULATION DYNAMICS AND PREDATION IMPACT OF THE INTRODUCED CTENOPHORE MNEMIOPSIS LEIDYI IN THE GULLMARS FJORD, WEST COAST OF SWEDEN Lene Friis Møller & Peter Tiselius Dep. Of Marine Ecology Kristineberg

More information

BIOLOGICAL OCEANOGRAPHY

BIOLOGICAL OCEANOGRAPHY BIOLOGICAL OCEANOGRAPHY AN INTRODUCTION 0 ^ J ty - y\ 2 S CAROL M. LALLI and TIMOTHY R. PARSONS University of British Columbia, Vancouver, Canada PERGAMON PRESS OXFORD NEW YORK SEOUL TOKYO ABOUT THIS VOLUME

More information

Zooplankton of the Okhotsk Sea: A Review of Russian Studies 19

Zooplankton of the Okhotsk Sea: A Review of Russian Studies 19 Zooplankton of the Okhotsk Sea: A Review of Russian Studies 19 Figure 18. Distribution of zooplankton biomass (mg m 3 ) near the western coast of Kamchatka in the 0-50 m layer: 1 = 100-500, 2 = 500-1,000,

More information

Understanding the role of the YS Bottom Cold Water ( 10 C) on the survival strategy of Euphausia pacifica throughout the hot summer

Understanding the role of the YS Bottom Cold Water ( 10 C) on the survival strategy of Euphausia pacifica throughout the hot summer Understanding the role of the YS Bottom Cold Water ( 10 C) on the survival strategy of Euphausia pacifica throughout the hot summer Euphausia pacifica Se-J. Ju, H.S. Kim, W.S. Kim, D.H. Kang and A.R. Ko

More information

Stable isotope analysis in fisheries food webs

Stable isotope analysis in fisheries food webs Stable isotope analysis in fisheries food webs Rod M. Connolly Australian Rivers Institute, Griffith University, PMB, Gold Coast Mail Centre, Queensland 9726, Australia Email: r.connolly@griffith.edu.au

More information

Climate change, ocean acidification and individual-based models: Why the little things matter

Climate change, ocean acidification and individual-based models: Why the little things matter Climate change, ocean acidification and individual-based models: Why the little things matter Author Richards, Russell, Meynecke, Olaf, Chaloupka, M, Tomlinson, Rodger Benson Published 2012 Conference

More information

Chapter 10. Marine Ecology

Chapter 10. Marine Ecology Chapter 10 Marine Ecology Copyright 2016 McGraw-Hill Education. All rights reserved. No reproduction or distribution without the prior written consent of McGraw-Hill Education. Marine Ecology Ecology is

More information

Jacqueline M. Grebmeier Chesapeake Biological Laboratory University of Maryland Center for Environmental Science, Solomons, MD, USA

Jacqueline M. Grebmeier Chesapeake Biological Laboratory University of Maryland Center for Environmental Science, Solomons, MD, USA Update on the Pacific Arctic Region Synthesis Activity as part of the ICES/PICES/PAME Working Group on Integrated Ecosystem Assessment of the Central Arctic Ocean (WGICA) Jacqueline M. Grebmeier Chesapeake

More information

HOMEWORK PACKET UNIT 2A. Part I: Introduction to Ecology

HOMEWORK PACKET UNIT 2A. Part I: Introduction to Ecology CP Biology Name Date Period HOMEWORK PACKET UNIT 2A Part I: Introduction to Ecology Name Class Date 3.1 What Is Ecology? Studying Our Living Planet 1. What is ecology? 2. What does the biosphere contain?

More information

The ecological importance of Antarctic Silverfish

The ecological importance of Antarctic Silverfish PCAS 18 (2015/2016) Critical Literature Review (ANTA602) The ecological importance of Antarctic Silverfish Rata Pryor Rodgers Student ID: 41102419 Word count: 3113 Abstract The Antarctic marine ecosystem

More information

PICES XIV (2005) in Vladivostok

PICES XIV (2005) in Vladivostok PICES XIV (2005) in Vladivostok Present and future of the North Pacific simulated by a high resolution coupled atmosphere-ocean model Hiroyasu HASUMI (CCSR) Tatsuo SUZUKI (FRCGC) Takashi SAKAMOTO (FRCGC)

More information

Chaetognatha of the Namibian Upwelling Region: Taxonomy, Distribution and Trophic Position

Chaetognatha of the Namibian Upwelling Region: Taxonomy, Distribution and Trophic Position Chaetognatha of the Namibian Upwelling Region: Taxonomy, Distribution and Trophic Position Karolina Bohata*, Rolf Koppelmann Institute for Hydrobiology and Fisheries Science, Center for Earth Systems Research

More information

Catastrophic reduction of seaice in the Arctic Ocean - its impact on the marine ecosystems in the polar region-

Catastrophic reduction of seaice in the Arctic Ocean - its impact on the marine ecosystems in the polar region- 1/12 Catastrophic reduction of seaice in the Arctic Ocean - its impact on the marine ecosystems in the polar region- KAKENHI No.22221003 Naomi Harada (JAMSTEC) J. Onodera, E. Watanabe, K. Matsuno, K. Kimoto,

More information

Outflow of Okhotsk Sea Water and the oceanic condition of the sea east of Hokkaido

Outflow of Okhotsk Sea Water and the oceanic condition of the sea east of Hokkaido Sea ice, water mass and freshwater processes/coastal lagoons Outflow of Okhotsk Sea Water and the oceanic condition of the sea east of okkaido Yutaka Nagata Marine Information Research Center, Japan ydrographic

More information

Distribution and transport variations of source waters for North Pacific Intermediate Water formation revealed by multiple tracer analysis

Distribution and transport variations of source waters for North Pacific Intermediate Water formation revealed by multiple tracer analysis Distribution and transport variations of source waters for North Pacific Intermediate Water formation revealed by multiple tracer analysis Yugo Shimizu 1, Lynne D. Talley 2, Shin-ichi Ito 1, and Miyuki

More information

9.0 Consistency with the Magnuson-Stevens Fishery Conservation and Management Act (MSFCMA)

9.0 Consistency with the Magnuson-Stevens Fishery Conservation and Management Act (MSFCMA) 9.0 Consistency with the Magnuson-Stevens Fishery Conservation and Management Act (MSFCMA) This analysis was prepared in accordance with the requirements of the Magnuson- Stevens Fishery Conservation and

More information

Archimer

Archimer Please note that this is an author-produced PDF of an article accepted for publication following peer review. The definitive publisher-authenticated version is available on the publisher Web site Journal

More information

Identifying risks of geoduck aquaculture: the role of larval transport

Identifying risks of geoduck aquaculture: the role of larval transport Material intended for presentation only. Please do not cite, copy or distribute Bivalve life cycle: a simplified view however: High fecundities (female geoduck 50 million eggs/year) High larval mortalities

More information

SARSIM Model Output for the Distribution of Sardine in Canadian, US and Mexican Waters. Richard Parrish October 13, 2015

SARSIM Model Output for the Distribution of Sardine in Canadian, US and Mexican Waters. Richard Parrish October 13, 2015 SARSIM Model Output for the Distribution of Sardine in Canadian, US and Mexican Waters. Richard Parrish October 13, 2015 Agenda Item H.1.c The information presented below was taken from a model that I

More information

Tiphaine Chouvelon, Laurie Violamer, Aurélie Dessier, Paco Bustamante, Françoise Mornet, Cécilia Pignon-Mussaud, Dupuy Christine

Tiphaine Chouvelon, Laurie Violamer, Aurélie Dessier, Paco Bustamante, Françoise Mornet, Cécilia Pignon-Mussaud, Dupuy Christine Small pelagic fish feeding patterns in relation to food resource variability: an isotopic investigation for Sardina pilchardus and Engraulis encrasicolus from the Bay of Biscay (north-east Atlantic) Tiphaine

More information

tentacular club

tentacular club click for previous page - 130 - Moroteuthis ingens (Smith, 1881) ONYCHO Moro 1 Onychoteuthis ingens Smith, 1881, Proc.Zool.Soc.Lond., 1881:25. Synonymy : Onychoteuthis ingens Smith, 1881. En - Greater

More information

Changes in carbon and nitrogen stable isotopes of chironomid larvae during growth, starvation and metamorphosis

Changes in carbon and nitrogen stable isotopes of chironomid larvae during growth, starvation and metamorphosis RAPID COMMUNICATIONS IN MASS SPECTROMETRY Rapid Commun. Mass Spectrom. 2007; 21: 997 1002 Published online in Wiley InterScience (www.interscience.wiley.com).2925 Changes in carbon and nitrogen stable

More information

Tiphaine Chouvelon, Alexis Chappuis, Paco Bustamante, Sébastien Lefebvre, Françoise Mornet, Gaël Guillou, Laurie Violamer, Dupuy Christine

Tiphaine Chouvelon, Alexis Chappuis, Paco Bustamante, Sébastien Lefebvre, Françoise Mornet, Gaël Guillou, Laurie Violamer, Dupuy Christine Trophic ecology of European sardine Sardina pilchardus and European anchovy Engraulis encrasicolus in the Bay of Biscay (north-east Atlantic) inferred from δ13c and δ15n values of fish and identified mesozooplanktonic

More information

Aggregation Hotspots

Aggregation Hotspots Photo: Mike Brittain Aggregation Hotspots George L. Hunt, Jr. School of Aquatic and Fishery Sciences University of Washington Aggregation Hotspots Predictable in Time and Space Places with high rates of

More information

Biology (Biology_Hilliard)

Biology (Biology_Hilliard) Name: Date: 1. There are two types of modern whales: toothed whales and baleen whales. Baleen whales filter plankton from the water using baleen, plates made of fibrous proteins that grow from the roof

More information

Large pelagic squids, mid-level trophic gateways?

Large pelagic squids, mid-level trophic gateways? Large pelagic squids, mid-level trophic gateways? Trophic ecology of two pelagic squids, revisited Matthew Parry, Ph.D. Pacific Islands Regional Office, NOAA Why study the trophic ecology of these squids?

More information

Species specific geographical distribution patterns in a warm Barents Sea: haddock vs. cod

Species specific geographical distribution patterns in a warm Barents Sea: haddock vs. cod Species specific geographical distribution patterns in a warm Barents Sea: haddock vs. cod Nordic Climate-Fish 2nd Conference: Latitudinal changes in marine resources, exploitation and society within the

More information

Synchronicity of climate driven regime shifts among the East Asian Marginal Sea waters and major fish species

Synchronicity of climate driven regime shifts among the East Asian Marginal Sea waters and major fish species Synchronicity of climate driven regime shifts among the East Asian Marginal Sea waters and major fish species SM Mustafizur Rahman 1, Chung Il Lee 2 1 Research Institute for Dok-do and Ulleung-do, Kyungpook

More information

The biological importance of the major ocean currents

The biological importance of the major ocean currents The biological importance of the major ocean currents Squid and the western boundary currents Illex illecebrosus, the short-finned squid Squid use the Gulf Stream to facilitate their migration. The center

More information

Variability and trend of the heat balance in the southeast Indian Ocean

Variability and trend of the heat balance in the southeast Indian Ocean Variability and trend of the heat balance in the southeast Indian Ocean Ming Feng, CSIRO Marine & Atmospheric Research Arne Biastoch, Claus Böning, Leibniz-Institut für Meeresforschung Nick Caputi, Department

More information

Atmospheric circulation analysis for seasonal forecasting

Atmospheric circulation analysis for seasonal forecasting Training Seminar on Application of Seasonal Forecast GPV Data to Seasonal Forecast Products 18 21 January 2011 Tokyo, Japan Atmospheric circulation analysis for seasonal forecasting Shotaro Tanaka Climate

More information

Ecology of krill in Icelandic waters. Teresa Silva

Ecology of krill in Icelandic waters. Teresa Silva Ecology of krill in Icelandic waters Teresa Silva What are krill? Video used with permission from Steinunn H. Ólafsdóttir (MFRI) Introduction Life-Cycle Larval krill: plankton, i.e., drift with the currents

More information

JAPAN GLOBEC JAPAN GLOBEC JAPAN GLOBEC JAPAN GLOBEC

JAPAN GLOBEC JAPAN GLOBEC JAPAN GLOBEC JAPAN GLOBEC Progress of the Japan-GLOBEC project Coordinator: Yasunori Sakurai (sakurai@fish.hokudai.ac.jp) Graduate School of Fisheries Sciences, Hokkaido University, Hokkaido, Japan Japan is located near the confluence

More information

Critical Issues in Assessment of Offshore Wind Farm Development on Dispersion and Settling of Scallop Larvae in the Northeast U.S.

Critical Issues in Assessment of Offshore Wind Farm Development on Dispersion and Settling of Scallop Larvae in the Northeast U.S. Critical Issues in Assessment of Offshore Wind Farm Development on Dispersion and Settling of Scallop Larvae in the Northeast U.S. Coastal Ocean Changsheng Chen School for Marine Science and Technology

More information

Two of the main currents in the Arctic region are the North Atlantic Current (in red) and the Transport Current (in blue).

Two of the main currents in the Arctic region are the North Atlantic Current (in red) and the Transport Current (in blue). Have you ever enjoyed playing in the snow or making snowmen in the wintertime? The winter season is our coldest season. However, some of the coldest days we have here in Indiana have the same temperature

More information

Biodiversity Classwork Classwork #1

Biodiversity Classwork Classwork #1 Biodiversity Classwork Classwork #1 1. What is biodiversity? 2. In the boxes below, create two ecosystems: one with low biodiversity and one with high biodiversity. Explain the difference. Biodiversity

More information

The mechanisms influencing the timing, success and failure of spawning in natural populations of the sea urchin Strongylocentrotus intermedius

The mechanisms influencing the timing, success and failure of spawning in natural populations of the sea urchin Strongylocentrotus intermedius The mechanisms influencing the timing, success and failure of spawning in natural populations of the sea urchin Strongylocentrotus intermedius in the northwestern Sea of Japan P. M. Zhadan 1, M. A. Vaschenko

More information

CHAPTER 5 WARM UPS. Mrs. Hilliard

CHAPTER 5 WARM UPS. Mrs. Hilliard CHAPTER 5 WARM UPS Mrs. Hilliard CHAPTER 5 VOCABULARY 1. Photosynthesis 2. Cellular respiration 3. Producer 4. Consumer 5. Decomposer 6. Food chain 7. Food web 8. Trophic level 9. Carbon cycle 10. Nitrogen-fixing

More information

Courtesy of John Mitchell

Courtesy of John Mitchell Shrimp trawl fisheries Marine fish stocks are under increasing threat from a range of spatially based problems that might be natural or involve socio-economic factors. To best manage spatial problems,

More information

GODAE Ocean View Activities in JMA (and Japan)

GODAE Ocean View Activities in JMA (and Japan) GOVST VIII, Nov. 6 th, 2017, Bergen, Norway GODAE Ocean View Activities in JMA (and Japan) Yosuke Fujii 1, Norihisa Usui 1, Takahiro Toyoda 1, Nariaki Hirose 1, Hiromichi Igarashi 2, and Japan GODAE group

More information

Temporal and Spatial Patterns of th.

Temporal and Spatial Patterns of th. Temporal and Spatial Patterns of th Titleconchopheria on the Shell of the In coronatus coreensis Author(s) Xing, Yongmei; Wada, Keiji Citation PUBLICATIONS OF THE SETO MARINE BIO LABORATORY (2001), 39(2-3):

More information

Algae and Dissolved Oxygen Dynamics of Landa Lake and the Upper Spring Run

Algae and Dissolved Oxygen Dynamics of Landa Lake and the Upper Spring Run Algae and Dissolved Oxygen Dynamics of Landa Lake and the Upper Spring Run Why study algae and dissolved oxygen dynamics of Landa Lake and the Upper Spring Run? During low-flow conditions, extensive algal

More information

Time-series observations in the Northern Indian Ocean V.V.S.S. Sarma National Institute of Oceanography Visakhapatnam, India

Time-series observations in the Northern Indian Ocean V.V.S.S. Sarma National Institute of Oceanography Visakhapatnam, India The Second GEOSS Asia-Pacific Symposium, Tokyo, 14-16 th April 28 Time-series observations in the Northern Indian Ocean V.V.S.S. Sarma National Institute of Oceanography Visakhapatnam, India Seasonal variations

More information

Marine biologists have identified over 250,000 marine species. This number is constantly increasing as new organisms are discovered.

Marine biologists have identified over 250,000 marine species. This number is constantly increasing as new organisms are discovered. A wide variety of organisms inhabit the marine environment. These organisms range in size from microscopic bacteria and algae to the largest organisms alive today blue whales, which are as long as three

More information

Report R.V. HEINCKE cruise No. 161

Report R.V. HEINCKE cruise No. 161 Report R.V. HEINCKE cruise No. 161 Institute: Institute for Marine Research Kiel, Department of Fisheries Biology, Kiel University Date: from 14.11.2001 to 26.11.2001 Area: German Bight, Fisher, Skagerrak

More information

PRINCIPLE OF OCEANOGRAPHY PBBT101 UNIT-1 INTRODUCTION OF OCEANIC ENVIRONMENT. PART-A (2 Marks)

PRINCIPLE OF OCEANOGRAPHY PBBT101 UNIT-1 INTRODUCTION OF OCEANIC ENVIRONMENT. PART-A (2 Marks) PRINCIPLE OF OCEANOGRAPHY PBBT101 UNIT-1 INTRODUCTION OF OCEANIC ENVIRONMENT 1. Define marine ecosystem. 2. What is geography? 3. Give two Oceanic zones 4. What is sea? 5. Define oceanography? 6. Enlist

More information

Modelling the Channel and the Bay of Biscay using the MARS model. Contributions: Dyneco Physed / Pelagos / Benthos EMH

Modelling the Channel and the Bay of Biscay using the MARS model. Contributions: Dyneco Physed / Pelagos / Benthos EMH Modelling the Channel and the Bay of Biscay using the MARS model Contributions: Dyneco Physed / Pelagos / Benthos EMH Overview 1. The MANGA Configuration 2. Model validation: Sea Surface Temperature 3.

More information

2004 State of the Ocean: Physical Oceanographic Conditions in the Newfoundland and Labrador Region

2004 State of the Ocean: Physical Oceanographic Conditions in the Newfoundland and Labrador Region Canadian Science Advisory Secretariat Science Advisory Report 25/18 Research vessel CCGS Teleost 24 State of the Ocean: Physical Oceanographic Conditions in the Newfoundland and Labrador Region Background

More information

SHIFT IN SIZE-AT-AGE OF THE STRAIT OF GEORGIA POPULATION OF PACIFIC HAKE (MERLUCCIUS PRODUCTUS)

SHIFT IN SIZE-AT-AGE OF THE STRAIT OF GEORGIA POPULATION OF PACIFIC HAKE (MERLUCCIUS PRODUCTUS) SHIFT IN SIZE-AT-AGE OF THE STRAIT OF GEORGIA POPULATION OF PACIFIC HAKE (MERLUCCIUS PRODUCTUS) JACQUELYNNE R. KING Pacific Biological Station Fisheries and Oceans Canada Nanaimo, British Columbia V9S

More information

BIOS 569: Practicum in Field Biology. Impact of DOC in the Zooplankton Community Composition. Amarilis Silva Rodriguez. Advisor: Patrick Kelly

BIOS 569: Practicum in Field Biology. Impact of DOC in the Zooplankton Community Composition. Amarilis Silva Rodriguez. Advisor: Patrick Kelly BIOS 569: Practicum in Field Biology Impact of DOC in the Zooplankton Community Composition Amarilis Silva Rodriguez Advisor: Patrick Kelly 2013 Abstract: Dissolved organic carbon (DOC) plays an important

More information

Primary Productivity in a Cold Water Mass and the Neighborhood Area Occurring off Enshu-Nada in the Late Summer of 1989

Primary Productivity in a Cold Water Mass and the Neighborhood Area Occurring off Enshu-Nada in the Late Summer of 1989 Journal of Oceanography Vol. 48, pp. 105 to 115. 1992 Primary Productivity in a Cold Water Mass and the Neighborhood Area Occurring off Enshu-Nada in the Late Summer of 1989 AKIHIRO SHIOMOTO and SATSUKI

More information

Eddy and Chlorophyll-a Structure in the Kuroshio Extension Detected from Altimeter and SeaWiFS

Eddy and Chlorophyll-a Structure in the Kuroshio Extension Detected from Altimeter and SeaWiFS 14th Symposium on Integrated Observing and Assimilation Systems for the Atmosphere, Oceans, and Land Surface (IOAS-AOLS), AMS Atlanta, January 17-21, 21 Eddy and Chlorophyll-a Structure in the Kuroshio

More information

History and meaning of the word Ecology A. Definition 1. Oikos, ology - the study of the house - the place we live

History and meaning of the word Ecology A. Definition 1. Oikos, ology - the study of the house - the place we live History and meaning of the word Ecology. Definition 1. Oikos, ology - the study of the house - the place we live. Etymology - origin and development of the the word 1. Earliest - Haeckel (1869) - comprehensive

More information

Title NOTES ON SOME ANMAL PLANKTONS COLL FROM THE SEA OF JAPAN OFF SAN'N-D Author(s) Furuhashi, Kenzo Citation PUBLCATONS OF THE SETO MARNE BO LABORATORY (1952), 2(2): 257-260 ssue Date 1952-10-05 URL

More information

Primary Producers. Key Ideas

Primary Producers. Key Ideas Primary Producers Kelp forests are one of the ocean s most productive habitats. 1 Key Ideas Energy flows through living systems, but matter is recycled. Primary producers (autotrophs) synthesize glucose

More information