CONTRIBUTION TO THE KNOWLEDGE OF SOMALIAN ODONATA

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1 Monitore Zoologico Italiano. Supplemento ISSN: (Print) (Online) Journal homepage: CONTRIBUTION TO THE KNOWLEDGE OF SOMALIAN ODONATA Salvatore Carfì To cite this article: Salvatore Carfì (1974) CONTRIBUTION TO THE KNOWLEDGE OF SOMALIAN ODONATA, Monitore Zoologico Italiano. Supplemento, 5:1, To link to this article: Published online: 12 Jul Submit your article to this journal Article views: 63 View related articles Citing articles: 1 View citing articles Full Terms & Conditions of access and use can be found at

2 Monitore Zoologico Italiano ITALIAN JOURNAL OF ZOOLOGY PUBBLICATO DALLA UNIVERSITi\ DEGLI STUD! Dl FIRENZE CON IL CONTRIBUTO DEL CONSIGLIO NAZIONALE DELLE RICERCHE N. S. SUPPLEMENTO V NO. 13: CONTRIBUTION TO THE KNOWLEDGE OF SOMALIAN ODONATA (PUBBLICAZIONI DEL CENTRO Dl STUDIO PER LA FAUNISTICA ED ECOLOGIA TROPICAL! DEL C.N.R.: LXXX) SALvA TORE CARPI lstituto di Zoologia dell'universita di Firenze Received 18 September 1973 I. Introduction II. Systematics Genus Lestes Leach, Lestes pallidus Rambur, Lestes tridens McLachlan, Lestes uncifer Karsch, Genus Agriocnemis Selys, Agriocnemis exilis Selys, Genus Ceriagrion Selys, Ceriagrion glabrum (Burmeister, 1839) 2. Ceriagrion moorei Longfielp, Genus Enallagma Charpentier, Enallagma somalicum Longfield, Enallagma subfurcatum Selys, Genus Ischnura Charpentier, Ischnura senegalensis (Rambur, 1842) Genus Pseudagrion Selys, Pseudagrion acaciae Forster, Pseudagrion kersteni (Gerstaecker, 1869) 3. Pseudagrion lindicum Gruenberg, Pseudagrion massaicum Sjoestedt, Pseudagrion pseudomassaicum Pinhey, 1951 Genus Phaon Selys, Phaon iridipennis (Burnmeister, 1839) Genus Ictinogomphus Cowley, Ictinogomphus ferox (Rambur, 1842) page 149» 152» 152 >> 152 >> 152 >> 153 >> 155 >> 155 >> 155 >> 155 >> 155 >> 155 >> 155 >> 157 >> 157 >> 157 >> 158 >> 158 >> 158 >> 158 >> 159 >> 160 >> 161 >> 161 >> 161 >> Published online 12 Jul 2013

3 148 S. CARFt Genus Paragomphus Cowley, Paragomphus genei (Selys, 1841) Genus Anax Leach, Anax imperator Leach, Anax parthenope Selys, 1839 Genus Hemianax Selys, Hemianax ephippiger (Burmeister, 1839). Genus Acanthagyna Kirby, Acanthagyna cylindrata (Karsch, 1891). 2. Acanthagyna manderica (Gruenberg, 1902) Genus Macromia Rambur, Macromia contumax (Selys, 1879) Genus Tetrathemis Brauer, Tetrathemis polleni (Selys, 1869) Genus Nesciothemis Longfield, Nesciothemis farinosum (Forster, 1898) Genus Orthetrum Newman, Orthetrum abbotti Calvert, Orthetrum brachiate (Palisat de Beauvais, 1805) 3. Orthetrum chrysostigma (~urmeister, 1839) 4. Orthetrum sabina (Drury ) 5. Orthetrum taeniolatum (Schneider, 1845) 6. Orthetrum trinacria (Selys, 1841) Genus Palpopleura Rambur, Palpopleura lucia (Drury, 1773) 2. Palpopleura deceptor (Calvert, 1899) Genus Diplacodes Kirby, Diplacodes lefebvrei (Rambur, 1842) Genus Crocothemis Brauer, Crocothemis erythraea (Brulle, 1832) Genus Brachythemis Brauer, Brachythemis leucosticta (Burmeister, 1839) Genus Sympetrum Newmann, Sympetrum fonscolombei (Selys, 1840) Genus Philonomon Forster, Philonomon luminans (Karsch, 1893) Genus Trithemis Brauer, Trithemis annulata (Palisat de Beauvais, 1805) 2. Trithemis arteriosa (Burmeister, 1839) 3. Trithemis furva Karsch, Trithemis kirbyi ardens Gerstaecker, Trithemis stictica (Burmeister, 1839) Genus Olpogastra Karsch, Olpogastra lugubris (Karsch, 1895). Genus Rhyothemis Hagen, Rhyothemis semihyalina (Desjardins, 1835) Genus Tholymis Hagen, Tholymis tillarga (Fabricius, 1798). Genus Pantala Hagen, Pantala flavescens (Fabricius, 1798) page 161 )) 161» 162» 162» 162 :> 162» 162 >> 163» 163» 163» 163» 163» 163» 163» 164 >> 1<?4» 164» 164» 164» 165» 165» 166» 166» 166» 166 >> 167» 167» 167 >> 168» 168» 169» 169» 169» 169» 169» 169» 170» 170 >> 170» 171 >> 171» 171» 172» 172» 172» 172» 172» 172» 173» 173

4 SOMALIAN ODONATA Genus Tramea Hagen, 11:!61 I. Tramea basilaris (Palisot de Beauvois, 1805). 2. Tramea limbata (Desjardins, 1832). Genus Urothemis Brauer, l. Urothemis assignata (Selys, 1872). 2. Urothemis edwardsi (Selys, 1849). 3. Urothemis siji.nata aethiopica Nielsen, 1957 Genus Macrodiplax Brauer, 1868 l. Macrodiplax cora (Brauer, 1867) II I. List of the localities Summary Riassunto References 149 page 173 )) 173 )) li-t t 17-t )) I. INTRODUCTION The first data on Somalian Odonata is due to KIRBY ( 1896) who studied specimens collected in 1895 by Mr and Mrs Lort Phillips. Following this is CALVERT's (1899) work on the material collected by A. Donaldson Smith between 1894 and 1895 during his trip to Lake Rudolph, that of McLACHLAN ( 1900) on the insects collected by V.A. Peel between 1895 and 1897 and that of SHARPE (1901) on some material collected in 1899 by A. Donaldson Smith (this time by special request of «His Highness The Gaekwar >>) for the Museum of Baroda (India). During his expedition across North-East Africa, Carlo Freiherr von Erlanger had occasion to collect numerous dragonflies (though not always in Somalian territory) that were later studied by FORSTER ( 1906b ). Other Odonata collected by C. L. Collenette were determined by LoNGFIELD ( 1931 ). Based on a study of a small odonatological collection belonging to the Marquis S. Patrizi, NIELSEN ( 1935) drafted the «Catalogo degli Odonati del Corno orientale dell'africa» which re:::ords 37 species from Somalia (considering Lestes ictericus Gerstaecker, 1869 and Lestes somalicus Fi.)rster, 1906 as one species). Further data on the Somalian odonatological fauna are due to NIELSEN (1936, 1957) and PINHEY (1961). A fairly large number of Odonata have been collected during the course of several research missions in Somalia promoted first by the «Istituto di Zoologia» (Director Prof. L. Pardi) and the <<Musco Zoologico» (Director Prof. B. Lanza) of the University of Florence, later by the << Centro di Studio per Ia Faunistica ed Ecologia Tropicali del Consiglio Nazionale delle Ricerche» (Director Prof. L. Pardi). From the examination of this material and bibliographic records, I was able to ascertain that within the actual confines of the Somali Democratic Re-

5 150 S. CARFl public, the order Odonata includes seven families with 30 genera (seven new to Somalia) and 53 species of which the following 12 are new to Somalia: Lestes tridens McLachlan, Lestes uncifer Kai'sch, Agriocnemis exilis Selys, Pseudagrion acaciae Forster, Ictinogomphus ferox (Rambur, 1842). Tetrathemis polleni (Selys, 1869). Olpogastra lugubris (Karsch, 1895). Tholymis tillarga (Fabricius, 1798). Tramea basilaris (P. de Beauvais, 1805). Tramea limbata (Desjardins, 1832). Urothemis edwardsi (Selys, 1849). Macrodiplax cora (Brauer, 1867). On the contrary, the following species have to be excluded from the Somalian fauna, at least temporarily: Macromia bicornis Forster, Referring to pveceding data, PINHEY (1961, p. 117 and 1962, p. 210) gives this as present in Somalia while FoRSTER (1906b, p. 321) says: «Mbundjo am mittleren Mongo im Bakundulande, Kamerum, von Herrn Missionar Bohner rnitgeteilt». Macromia monoceros Forster, PINHEY (1961, p. 112 and 1962, p. 211 ) cites this for Somalia probably because of a misinterpretation of FoRSTER (1906b, p. 320): «Nguelo im Bergland von Ost-Usambara» (i.e., Tanzania). Macromia pallidinervis Forster, PINHEY (1961, p. 108) writes «Forster's type locality N.-E. Africa may have been Somalia» and subsequently, PINHEY (1962, p. 212) «... Somalia(? or Southern Abyssini,a)».In fact FoRSTER (1906b, p. 318) says: «Ein einziges junges 'i? aus II Hauaschgebiet, gefangen am Susukifluss am 27, November 1900 (i.e., Ethiopia). Orthetrum guineense Ris, In as much as R1s ( , p. 207 and p. 1082) doesn't cite any Somalian localities, LoNGFIELD (1955, p. 27) says «... but it has also

6 SOMALIAN ODONATA 151 been taken in... and Somaliland», without any other indication. I believe the subsequent citations of PINHEY (1961, p. 135; 1962, p. 236; 1970b, p. 295) rae due to this. Trithemis donaldsoni (Calvert, 1899). Pseudornacrornia Donaldsoni Calvert, 1899, p. 235, table X, fig. 5 (1 & n Ph.A.N.S., Stony brook(!), 17.VIII.1894). Trithernis Erlangeri Forster, 1906 b, p. 312, table A, fig. 5 (2 & & Darolifluss, 6.11!.1901; 1 & Webbe Mane). Trithernis Donaldsoni Dona/dsoni Ris, 1912, p. 782, fig The caption for fig. 446 says: «Trithemis Donaldsoni Donaldsoni cf Somalia», while on p. 784 one reads: «Die Beschreibung und Abbildung und eine Zeichnung der Genit. 2. Segment der Type, die ich Dr. CALVERT verdanke und hier reproduziere». As the figure reproduces the accessory genitalia of the CALVERT ( 1899) type coming from Stony brook, it evidently can't be Somalia. Contrary to the interpretation of R1s (1912, p. 782) the localities of FoRSTER (1906b, p. 312) are not in Somalia. I think the subsequent citations made by PINHEY (1961, p. 169; 1962, p. 270; 1970a, p. 131) are due to the above. I wish to thank Prof. E. ToRTONESE (Director of the «Museo Civico di Storia Naturale Giacomo Doria di Genova») who kindly permitted me to consult the odonatological collection of the Museum and Dr F. CAPRA (honorary curator of the same Museum); Dr C. NIELSEN of Bologna who put his rich collection of tropical Odonata at my disposition; Dr E. C. G. PINHEY (Keeper of Entomology, National Museum, Bulawayo) for the precious suggestions he kindly gave me and Mr P. WARD of the British Museum (Natural History), London, who kindly checked the determination of some species. I also wish to thank the two technicians of the «Istituto di Zoologia dell'universita» of Florence, Mr R. INNOCENTI (for the photographs) and Mr M. MANZANI (for the drawings). The following abbreviations are used in the present work: Bar.M. B.M. C.M. C.N. H.C.O. Baroda Museum (India); British Museum (Natural History), London; Coryndon Museum (now National Museum, Nairobi, Kenya); Collezione Nielsen (Bologna); Hope Collection, University Museum, Oxford; ( 1) CALVERT (1899, p. 299): << Stony brook, a tributary of the Erer river, mentioned by Dr Smith on p. 32, lies in 42"7'E- 7"35'N >>.

7 152 M.P. M.S.N.G. Ph.A.N.S. S.B.S. S. CARFl Museo Zoologico dell'universita di Firenze; Museo Civico di Storia Naturale «Giacomo Doria» di Genova; Philadelphia Academy of Natural Sciences; Spedizione Biologica fn Somalia del «Centro di Studio per la Faunistica ed Ecologia Tropicali del C.N.R.». II. SYSTEMATICS (1) 1. Lestes pallidus Rambur, ORDER ODONAT A SuBORDER ZYGOPTERA FAMILY LESTIDAE Genus LESTES Leach, 1815 Lestes ictericus: NIELSEN, 1935, p. 72 and p. 76 (Belet Amin, I d M.S.N.G., leg. S. Patrizi, 20.VI.1934). Lestes pa/lidus: PINHEY, 1961, p. 10 (Somalia, I d C.M.); PINHEY, 1962, p. 85 (the citation refers to preceding data). No specimens examined. Distribution; this species is known throughout tropical Africa and Asia as far as the Phi!llipines. Remarks: Lestes somalicus Forster, 1906, which PrNHEY, 1962, p. 95 considers as most probably a form of L. pallid us, has been described from specimens collected by C. Freiherr von Erlanger at Bucka on the Mane River. Such a locality does not fall within the confines of Somal.ia. 2. Lestes tridens McLachLan, 1895 (Fig. 1 ). Ola Uager, 2 cf cf n. 655 M.P., leg. S.B.S., 1 7.VIII.l970. Distribution: Congo, South Af11ica, Portugese East Africa. Remarks: this species is new to Somalia. ( 1) The families and genera are ordered according to the «List of Ethiopian Odonata», in PrNHEY (1962, p. 43).

8 SOMALIAN ODONATA 153 Fig Lestes tridens McLachlan, o n. 655 M.F., Ola Uager (Somalia). Left wings (A, B); anal appendages from above (C); same from right side (D). 3. Lestes uncifer Karsch, 1899 (Fig. 2). Giohar, 1 cf n. 653 M.P., leg. S.B.S., 27. VIII.l964.

9 154 S. CARFl Fig Lestes uncifer Karsch, o n. 653 M.P., Giohar (Somalia). Right wings (A, B); anal appendages from above (C); same from right side (D). Distribution: this species is known in Portugese East Africa, Tanganyika, Kenya, Uganda and Nigeria. Remarks: collected for the first time in Somalia.

10 SOMALIAN ODONATA Agriocnemis exilis Selys, FAMILY COENAGRIIDAE Genus AGRIOCNEMIS Selys, 1869 Ola Uager, 1 rf n. 654 M.F., leg. S.B.S., 28. VIII.l959; 1 ~ n. 638 M.F., leg. S.B.S., 17.VIII Distribution: this species has been found in most of the Ethiopian region, Madagascar and Mauritius (Mascaren Islands). Remarks: the Ola Uager male and female are the first to be collected in Somalia. Genus CERIAGRION Selys, 1876 l.ceriagrion glabrum (Burmeister, 1839). Ceriagrion glabrum: FoRSTER, 1906b, p. 338 (Lake Solole, 1 cl 12.VI.l901; Eleschia, a pair, leg. C. von Erlanger, 5.VII.l901); NIELSEN, 1935, p. 72 (Belet Amin, 1 d M.S.N.G., 1934; 1 d M.S.N.G., 19.VI.l934; 1 d M.S.N.G., 30.VI.l934; 1 d C.N., 20.VI.l934; leg. S. Patrizi); NIELSEN, 1935, p. 77 (the citation refers to preceding data); PINHEY, 1962, p. 119 (the citation refers to preceding data). El Bur, 1 rf n. 635 M.F., leg. S.B.S., 3-9.IX Giohar, 1 rf n. 636 M.F., leg. S.B.S., 10.VII-20.IX Distribution: Ethiopian region, Madagascar, Mauritius (Mascaren Islands), Seychelles Islands, Reunion Islands, Arabia, Australia. 2. Ceriagrion moorei Longfield, Ceriagrion moorei: PINHEY, 1961, p. 23 (British Somaliland, 1 a C.M.). Ola Uager, 4 rf rf n. 637 M.F., leg. S.B.S., 17.VIII Distribution: Nigeria, Ghana and East Africa. Genus ENALLAGMA Charpentier, Enallagma somalicum Longfield, 1931 (Fig. 3). Enallagma somalicum LoNGFIELD, 1931, p. 277, fig. p. 275 (Marojeh, 1 a Type, 2 d d Paratypes, B.M., leg. C. L. Collenette, 5.XI.l929); NIELSEN, 1935, p. 77 (the citation refers to preceding data); NIELSEN, 1957, p. 31 (Toh, 1 a M.S.N.G., leg. G. Scortecci, 29.VI.l957); PINHEY, 196i, p. 43 (British Somaliland Protectorate: Wadi Mait, 4 specimens C.M., leg. J. G. Williams;

11 156 S. CARFl Fig Enallagma somalicum Longfield, o n. 657 M.F., Run (Somalia). Right wings (A, B); anal appendages from right side (C); same from above (D); same from behind (E). Bihendullah, 4 specimens C.M., leg. P.R. 0. Bally); PINHEY, 1962, p. 135 (British Somaliland Protectorate; the citation refers to preceding data). Run, 1 cf n. 657 M.P., leg. S.B.S., 15.VIIL Remarks: as all the few known specimens of this species come from Somalia, it could probably be defined as endemic though, of course, further research might well enlarge its range; the female is still unknown. The determination of the specimen was confirmed by Mr P. Ward (British Museum) by direct comparison with the Type.

12 SOMALIAN ODONATA 157 Fig Ischnura senegalensis (Rambur, 1842). o n. 497 M.F., Giohar (Somalia). Right wings (A, B); anal appendages from right side (C); same from behind (D); same from above (E). 2. Enallagma subfurcatum Selys, Proischnura (Enallagma) subfurcatum: LONGFIELD, 1931, p. 277 (Medisheh, I a, I I? B.M., leg. C. L. Collenette, IX.I929). Enallagma subfurcatum: NIELSEN, 1935, p. 77 (the citation refers to preceding data); FINHEY, 1962, p. 134 (the citation refers to preceding data). No specimens examined. Distribution: Congo, Nigeri,a, South Africa, Uganda, Ethiopia, Somalia. Genus IscHNURA Charpentier, Ischnura senegalensis (Rambur, 1842) (Fig. 4 ). Jschnura senegalensis: LONGFIELD, 1931, p. 277 (Dobar, Goolis Mts, 1 I? B.M., leg. Lort Phil B.M., lips, II.I895; Medisheh, I a B.M., leg. C. L. Collenette, IX.I929; Buran, 8 a a, 11 1? 1? 13**

13 158 S. CARFi leg. C. L. Collenette, IX-X.l929); NIELSEN, 1935, p. 76 (the citation refers to preceding data); NIELSEN, 1936, p. 131 (Merca, I a M.S.N.G., leg. A. Pantano, 12.1X.I907); PINHEY, 1961, p. 46 (Somalia, I specimen C.M.); PINHEY, 1962, p. 137 (the citation refers to preceding data). Bud-Bud, 2 cf cf, 4 <.;? <.;? n. 496 M.P., leg S.B.S., 15.VIII Giohar, 5 cf cf, 4 <.;? <.;? n. 497 M.P., 16.IV-l.V.1968; 1 <.;? n. 495 M.P., ; leg. S.B.S. Mahaddei Uen (15 km to the East), 1 cf, 3 <.;? <.;? n. 494 M.P., leg. S.B.S., 17.IX Distribution: distributed throughout Africa and Asia as far as the Philippines. Genus PsEUDAGRION Selys, Pseudagrion acaciae Forster, 1906 (Fig. 5). Giohar, 1 cf n. 656 M.P., leg. S.B.S., 29. VII Distribution: Congo, East Africa from Natal to Egypt. R?marks: even though the range of this species is vast, it seems very localized which may explain the fact that the Giohar specimen is the first to be collected in Somalia. The determination of the specimen was confirmed by Mr P. Ward, British Museum (N.H.). 2. Pseudagrion kersteni (Gerstaecker, 1869). Pseudagrion kersteni: LONGFIELD, 1931, p. 277 (Medisheh, I a B.M., leg. C. L. Collenette, IX.l929); PINHEY, 1962, p. 125 (the citation refers to preceding data). Pseudagrion Kersteni: NIELSEN, 1935, p. 77 (the citation refers to preceding data). No specimens examined. Distribution: almost all the Ethiopian region. 3. Pseudagrion lindicum Gruenberg, 1902 (Fig. 6). Pseudagrion lindicum: NIELSEN, 1935, p. 37, figs 5 & 6 and p. 77 (Belet Amin, 8 a a, 7 'i1 'i1 M.S.N.G., 1934; 12 a a, I 'i1 M.S.N.G., 20.Vl.1934; 31 a a, 6 'i1 'i1 M.S.N.G., 30.Vl.1934; 2 a a, I 'i1 M.S.N.G., 20.VII.l934; 2 a a C.N., 20.VII.1934; 3 a a, 3 'i1 'i1 C.N., 30.Vl.1934, leg. S. Patrizi); NIELSEN, 1936, p. 129 (Giumbo, I a M.S.N.G., leg. A. Pan:ano, l.vl.1907); PINHEY, 1961, p. 32 (Somalia: Afgoi, I specimen C.M.); PINHEY, 1962, p. 126 (Somalia: the citation refers to preceding data). Afgoi (near Uebi Scebeli), 1 cf C.N., leg. Vatova, Ichthyological Expedition in A.O.I., 6.IX Belet Amin, 1 cf n M.P., leg. S. Patrizi, 30.VI Giohar, 1 <.;? n. 501 M.P., leg. S.B.S., 24.IX.1964; 2 <.;? <.;? n. 49& M.P., leg. S.B.S., 1970.

14 SOMALIAN ODONATA 159 Fig Pseudagrion acaciae Forster, o n. 656 M.F., Giohar (Somalia). Left wings (A, B); anal appendages from right side (C); same from above (D); same from behind (E). Mahaddei Uen ( 15 km to the East), 4 d' d', n. 499 M.F., leg. S.B.S., 17.IX Run, 3 d' d', n. 500 M.F., leg. S.B.S., 5.VIII.l964. Distribution: Congo and the East Af11ican coast. 4. Pseudagrion massaicum Sjoestedt, Pseudagrion punctum: FoRSTER, 1906b, p. 333 (Salakle, I 'i!, leg. C. von Erlanger, 7.VI.l901); NIELSEN, 1935, p. 77 (the citation refers to preceding data). No specimens examined. Distribution: South Africa, Kenya, Uganda, Congo, Angola.

15 160 S. CARFl Fig Pseudagrion lindicum Gruenberg, o n. 500 M.F., Run (Somalia). Right wings (A, B); anal appendages from above (C); same from right side (D). Remarks: in accordance with PINHEY, 1962, I ascribe the specimen examined by FoRSTER (1906b) to the species massaicum. Though very similar to this latter, P. punctum (Rambur) (PINHEY, 1962, 1964) is exclusive to the Malgascian fauna. 5. Pseudagrion pseudomassaicum Pinhey, Pseudagrion pseudomassaicum: PINHEY, 1961, p. 36 (British Somaliland Protectorate: B;hendullah, 1 specimen C.M., leg. P. Bally). No specimens examined. Distribution: Nigeria, Congo, South Africa, Mozambique, Tanzania, Uganda, Kenya and Somalia.

16 SOMALIAN ODONATA 161 FAMILY AGRIIDAE Genus PH:AON Selys, Phaon iridipennis (Burmeister, 1839). Sapho (Ph.) iridipennis iridipennis: FoRSTER, 1906b, p. 331 (Along the Giuba at Bardera, I il. leg. C. von Erlanger, 3l.V.1901). Phaon iridipennis: NIELSEN, 1935, p. 76 (the citation refers to preceding data); PINHEY, 1%2, p. 143 (the citation refers to preceding data). Eggi, 2 ~ ~ n. 481 M.P., leg. S.B.S., VIII Giohar, 1 d n. 484 M.P., 16.IV-l.V.1968; 2 ~ ~ n. 482 M.P., 3.VIII.1968; 1 ~ n. 483 M.P., 5.IX.1968; 1 d n. 485 M.P., 18.VII. 1970; leg. S.B.S. Ola Uager, 1 d, 1 ~ n. 480 M.P., leg. S.B.S., IX Distribution: all tropical and subtropical continental Africa. SuBORDER ANISOPTERA FAMILY GOMPHIDAE Genus ICTINOGOMPHUS Cowley, Ictinogomphus ferox (Rambur, 1842). Ola Uager, 1 d n. 460 M.P., 12.IX.1964; 1 d n. 461 M.P., 15.IX.1964; leg. S.B.S. Distribution: from Natal throughout the tropical belt of Africa. Remarks: as far as I know, the Ola Uager specimens are the first to be collected in Somalia though it was already known in Kenya and other nearby zones. Genus PARAGOMPHUS Cowley, Paragomphus genei (Selys, 1841). Mesogomphus bitarsatus FoRSTER, 1960b, p. 325 [Salakle on the Giuba river, 1 il (fide PINHEY, 1962, p. 185), leg. C. von Erlanger, 6.VI.1901]; NIELSEN, 1935, p. 74 (the citation refers to preceding data). Paragomphus hageni: PINHEY, 1962, p. 184 (the citation refers to the specimen studied by FORSTER, 1906b ). No specimens examined. Distribution: Africa.

17 162 S. CARFl 1. Anax imperator Leach, FAMILY AESHNIDAE Genus ANAX Leach, 1815 Anax mauricianus: LONGFIELD, 1931, p. 276 (Medisheh, 1 a B.M., leg. C. L. Collenette, IX.1929); NIELSEN, 1935, p. 73 (the citation refers to preceding data). Anax imperator: PINHEY, 1962, p. 195 (the citation refers to preceding data). Bud-Bud, 1 cf n. 467 M.P., leg. S.B.S., 15.VIII.l968. El Gambole, 1 cf, 1 9 n. 465 M.P., leg. S.B.S., 6.IX Run, n. 466 M.P., leg. S.B.S., 5.VIII.l964; 1 9 n. 469 M.P., leg. S.B.S., 16.VIII Scialambod, 1 cf n. 468 M.P., leg. S.B.S., 29.IV Distribution: Africa, Europe and Western Asia. 2. Anax parthenope Selys, Anax parthenope: LoNGFIELD, 1931, p. 277 (Buran, 5 a a B.M., leg. c. L. Collenette, IX X.1929); NIELSEN, 1935, p. 74 (the citation refers to preceding data); PINHEY, 1961, p. 96 («... extends into British Somaliland...»: the citation refers to preceding data); PINHEY, 1962, p. 196 («A north African species which infiltrates into British Somaliland...»: the citation refers to preceding data). No specimens examined. Distribution: North Africa, Southern Europe and West Asia. Genus HEMIANAX Selys, Hemianax ephippiger (Burmeister, 1839). Hemianax ephippiger: KIRBY, 1896, p. 523 (Bichen, Somaliland, 1 ~ B.M., leg. Mr & Mrs Lort Phillips, 5.II.1895); CALVERT, 1899, p. 233 (Berbera, 1 ~, leg. Donaldson Smith, 4.VII. 1894); FoRSTER, 1906b, p. 328 (on the Giuba: Maika Bebirrissa, 1 a, IO.VI.1901; Bua (Buba, Boa), 1 ~, 28.VI.1901; leg. C. von Erlanger); LONGFIELD, 1931, p. 277 (Berbera, the same specimen studied by CALVERT, 1899; Somaliland, 2 a a B.M., leg. Lort Phillips; Bichen, Somaliland, the same specimen studied by KIRBY, 1896; «Plain N. of AI Hills», 1 a B.M., leg. C. L. Collenette, XII.1929); NIELSEN, 1935, p. 68 (Belet Amin, 1 a M.S.N.G., leg. S. Patrizi, VI.1934); NIELSEN, 1935, p. 73 (the citation refers to preceding data); NIELSEN, 1957, p. 32 (Toh, I a M.S.N.G., leg. G. Scortecci, 28.VI.1953). Giohar, 1 cf n. 463 M.P., leg. S.B.S., 20.VIII.l968. Mogadiscio (surroundings), 4 cf cf n. 462 M.P., leg. S.B.S., 27.VIII Ola Uager, 1 9 n. 464 M.P., leg. S.B.S., 15.IX Distribution: from West Africa to India and the Southern palearctic region.

18 SOMALIAN ODONATA 163 Genus AcANTHAGYNA Kirby, Acanthagyna cylindrata (Karsch, 1891 ). Gynacantha cylindrata: NIELSEN, 1935, p. 68 and 74 (Belet Amin, I \? M.S.N.G., leg. S. Patrizi, VI.1934; Villaggio Duca degli Abruzzi). Acanthagyna cylindrata: PINHEY, 1961, p. 98 («Known from... Somalia... : the citation refers to preceding data); PINHEY, 1962, p. 198 («... Somalia... : the citation refers to preceding data). No specimens examined. Distribution: equatorial Africa. 2. Acanthagyna manderica (Gruenberg, 1902). Gynacantha manderica: NIELSEN, 1935, p. 67 and 74 (Belet Amin, I ~ C.N., 20.VI.l934; I ~ M.S.N.G., 30.VI.1934; I \? M.S.N.G., 15.VI.1934; leg. S. Patrizi). Acanthagyna manderica: PINHEY, 1962, p. 199 («... Somalia...»: the citation refers to preceding data). Giohar, 1 S? n. 489 M.F. 27.VII.1964; 1 S? n. 502 M.F., 6.VIII.l968; leg. S.B.S. Distribution: almost all equatorial Africa. FAMILY CORDULIIDAE Genus MACROMIA Rambur, Macromia contumax (Selys, 1879). Macromia (Phyllomacromia) biflava: NIELSEN, 1935, p. 68 and 74, fig. 3 (Ola Uager, I ~, I \? M.S.N.G., I \? C.N., leg. S. Patrizi, 15.VIII.l934). Macromia contuma.x: PINHEY, 1962, p. 211 (the citation refers to the specimens studied by NIELSEN, 1935). No specimens examined. Distribution: the collecting sites thus far known are all located m the equatorial belt of Africa. FAMILY LIBELLULIDAE Genus TETRATHEMIS Brauer, T etrathemis polleni (Selys, 1869). Ola Uager, 1 S? n. 491 M.F., 12.IX.1964; 1 cf n. 490 M.F., 15.IX. 1964; 1 S? n. 492 M.F., 17.VIII.1970; leg. S.B.S. Uadi Manas, 1 S? n. 644 M.F., leg. S.B.S., VII.1962.

19 164 S. CARFl Distribution: Nigeria, Congo, Uganda; East Africa from Natal to Kenya; Comore Islands, Madagascar. Remarks: the specimens cited are the first of this species to be collected in Somalia. Genus NESCIOTHEMIS Longfield, Nesciothemis farinosum (Forster, 1898). Orthetrum farinosum: Ris, 1909, p. 222 (Somaliland, 2 55, I!i' B.M., leg. G. W. Bury, 1906); LONGFIELD, 1931, p. 276 (Somaliland, the same specimens studied by Ris, 1909; Medisheh, 2 55 B.M., leg. C. L. Collenette, IX.l929; Marojeh, 2 55 B.M., leg. C. L. Collenette, IX.I929); NIELSEN, 1935, p. 74 (the citation refers to preceding data); NIELSEN, 1957, p. 32 (Toh, I 5 M.S.N.G., leg. G. Scortecci, 28.VI.l952). Nesciothemis farinosum: PINHEY, 1962, p. 240 («... Somalia... : the citation refers to preceding data). Baidoa, 1 cf n. 631 M.P., leg. S.B.S., 27.VIII.l968. Giohar, 1 cf n. 632 M.P., leg. S.B.S., l.viii Distribution: almost throughout continental Africa. Genus 0RTHETRUM Newman, Orthetrum abbotti Calvert, Orthetrum phi/lipsi KIRBY, 1896, p. 522 (Dobar, Goolis Mts, I 5 B.M., leg. Mr & Mrs Lort Phillips, ). Orthetrum chrysostigma chrysostigma (nee Burmeister, 1839): RIS, 1909, p. 204 (the citation refers to the Type of 0. phi/lipsi Kirby, 1896); LoNGFIELD, 1931, p. 276 (the citation refers to the Type of 0. phillipsi and the following unedited material: Einad, I 5 B.M., leg. C. L. Colienette, XI.1929 and Schimba Beris, 2 d 5 B.M., leg. C. L. Collenette, XII.1929). Orthetrum chrysostigma (nee Burmeister, 1839): NIELSEN, 1935, p. 74 (the citation refers to preceding data). Orthetrum abbotti: LoNGFIELD, 1955, p. 48 (the citation refers to the specimen studied by KIRBY, 1896, Type of 0. phillipsi); PINHEY, 1962, p. 232 (the citation refers to preceding data). No specimens examined. Distribution: almost all the Ethiopian region. 2. Othetrum brachiate (Palisot de Beauvais, 1805). Orthetrum brachiale: CALVERT, 1899, p. 237 (Berbera, I 5, leg. Donaldson Smith, 4.VII.C9~); LONGFIELD, 1931, p. 276 (Berbera, the same specimen studied by CALVERT, 1899; Medisheh, I 5 B.M., leg. C. L. Collenette, IX.1929; Buran, 2 55 B.M., leg. C. L. Collenette, IX.I929); NIELSEN, 1935, p. 74 (the citation refers to preceding data); LONGFIELD, 1955, p. 18 («... Somaliland... : the citation refers to preceding data); PINHEY, 1962, p. 233 (the citation refers to preceding data).

20 SOMALIAN ODONATA 165 Orthetrum stemmale capense: NIELSEN, 1935, p. 70 and 74 (Belet Amin, 2 ~ ~, I I' M.S.N.G., 20.VI.I934; 2 ~ ~ M.S.N.G., I ~ C.N., I I' C.N., 30.VI.I934; 1 ~ C.N., 1 I' M.S.N.G., IO.VII.I934; I ~ M.S.N.G., no date: leg. S. Patrizi; Ola Uager, I ~ juvenile C.N., 1 I' M.S.N.G., 15.VIII.I934, leg. S. Patrizi). Eggi, 3 d' d' n. 620 M.F., VII.1962; 1 < n. 714 M.F., VIII.1962; leg. S.B.S. Giohar, 1 d' n M.F., leg. Vatova, Ichthyological Expedition in A.O.I., 2.IX.1937; 1 d' n. 623 M.F., 27.VII.1964; 1 <.i n. 711 M.F., IV.1968; 1 d' n. 624, 3l.VI.1968; 1 < n. 712 M.F., 30.VII.1968; 3 d' d' n. 621 M.F., l.viii.1968; 1 d' n. 777 M.F., 1 d' n. 779 M.F., 3 < < n. 710 M.F., 1 < n. 715 M.F., 2.VIII.1968; 1 d' n. 766 M.F., 1 < n. 778 M.F., 2l.VIII.1968; 3 d' d' n. 619 M.F., 22.VIII.1968; 1 < n. 626 M.F., 18.VII.l970; 1 d' n. 630 M.F., 19.VII.1970; 1 d' n. 629 M.F., 1 d' n. 780 M.F., 29.VII.1970; 1 d' n. 628 M.F., l.viii.1970; 1 d', 1 < n. 625 M.F., 21.VIII.l970; 1 d' n. 627 M.F., VIII.1970; leg. S.B.S. Giuba, 1 < n. 713 M.F., leg. Paoli, VII Mahaddei, 1 d' n. 622 M.F., leg. S.B.S., 30.VIII Ola Uager, 1 d' n. 781 M.F., 12.VIII.1970; 2 d' d', 2 < < n. 670 M.F., 17.VIII.l970; leg. S.B.S. Distribution: almost all Africa. Remarks: of the specimens examined, n. 713 had already been determined as 0. stemmale capense; n. 1214, donated by Nielsen, had already been determined by him as 0. brachiate; n. 630, 779, 780 and 781 were kindly checked by Dr E. Pinhey of Bulawayo; n. 619 was captured while devouring a Diplacodes lefebvrei. 3. Orthetrum chrysostigma (Burmeister, 1839). Orthetrum chrysostigma: PINHEY, 1961, p. 136 (British Somaliland Protectorate, 35 specimens C.M.); PINIIEY, 1970b, p. 279 (Somalia, the citation refers to preceding data). Baidoa, 1 d' n. 616 M.F., 2.Il Uadi Manas, 3 d' d' n. 615 M.F., leg. S.B.S VII Distribution: all Africa, Southern Spain, Syria, Arabia. 4. Orthetrum sabina (Drury, 1770). Orthetrum Sabina: CALVERT, 1899, p. 238 (Berbera, I ~, I I', leg. Donaldson Smith, 4.VII.l894). Orthetrum sabina: Rrs, 1909, p. 225 (Somaliland, I I' B.M., leg. G. W. Bury, 19~6); LONG FIELD, 1931, p. 276 (Berbera, the same specimens studied by CALVERT, 1899; Dobar, Goolis Mts, 1 ~ B.M., leg. Lort Phillips, ; Berbera, the same specimen studied by Ris, 1909; Somaliland, l()o15'n- 45 IO'E, I 'i' B.M., leg. M. Portal Hyatt, XI.I928; Berbera, I ~ B.M., leg. C. L. Collenette, VIII.l928); NIELSEN, 1935, p. 74 (the citation refers to preceding data);

21 166 S. CARFl LoNGFIELD, 1955, p. 53 («... Somaliland..., the citation refers to preceding data); PINHEY, 196!, p. 141 (Berbera, I specimen C.M.); PINHEY, 1962, p. 238 («... Somalia... British Somaliland...», the citation refers to preceding data); PINHEY, 1970b, p. 272 («... to Somalia...», the citation refers to preceding data). No specimens examined. Distribution: North Africa from Algiers to Somalia; Asia along the tropical belt as far as Australia. 5. Orthetrum taeniolatum (Schneider, 1845). Orthetrurn brevistylurn KIRBY, 1896, p. 521 (Dobar, Goolis Mts., I c5 B.M., leg. Lort Phillips, 4.II.l895). Orthetrurn taeniolatum: Rrs, 1909, p. 192 (Dobar, the same specimen studied by KIRBY, 1896, Type of 0. brevistylurn); LoNGFIELD, 1931, p. 276 (Dobar, the same specimen studied by KIRBY, 1896, Tipe of 0. brevistylurn; Saugamore, 4 c5 c5 B.M., leg. Lort Phillips, II.I897; Buran, I c5 B.M., leg. C. L. Collenette, IX.I929); NIELSEN, 1935, p. 74 (the citation refers to preceding data); LoNGFIELD, 1955, p. 49 (<... Somaliland...»: the citation refers to preceding data); PIN HEY, 1962, p. 239 («... Somalia... British Somaliland...»: the citat:on refers to preceding data); PINHEY, 1970b, p. 282 («... Somalia...»: the citation refers to preceding data). No specimens examined. Distribution: Nigeria, Somalia, Ethiopia, Sudan, Western Asia. 6. Orthetrum trinacria (Selys, 1841). Orthetrurn trinacria: FoRSTER, 1906 b, p. 316 (along the Giuba at Maika Guele-Gedud, I ~, 2.VI.l901; Salakle, 2 ~ ~, 6.VI.l901; lake Geile, I c5, IS.VI.l901; Umfudu, 2 ~ ~, 18.VI.l901; leg. C. von Erlanger); NIELSEN, 1935, p. 74 (the citation refers to preceding data). Run, 1 d n. 612 M.F., 5.VIII.l964; 2 d d, 1 <;? n. 614 M.F., 11.VIII.l969; 3 d d n. 613 M.F., 11-VIII.l969; 1 <;? n. 611 M.P., 16.VIII.l969; leg. S.B.S. Distribution: all continental Africa, Asia Minor, Sicily. Genus PALPOPLEURA Rambur, Palpopleura lucia (Drury, ). Palpopleura marginal a: FoRSTER, 1906b, p. 303 [Arbarone, 3 c5 c5, V.I901; Haro Bussa, I c5, 2l.V.I901; Ganale, near lake Geile (perhaps within the confines of Somalia or at any rate in the nearby vicinity), 3 c5 c5, I ~, IS.V.l901; Ganale, near Jonte, I ~, IS.VII, I c5, 16.VII.I901; leg. C. von Erlanger]. Palpopleura lucia: NIELSEN, 1935, p. 71 (Oia Uager, I c5, 4 ~ ~, M.S.N.G., leg. S. Patrizi, VIII.l934); NIELSEN, 1935, p. 75 (the citation refers to preceding data); PINHEY, 1962, p. 243 (the citation refers to preceding data). Ola Uager, 1 <;? n. 470 M.P., 13.IX.1964; 1 d n. 471 M.P., 15.IX.

22 SOMALIAN ODONATA ; 1 cf n. 474 M.F., 12.VIII.l970; 1 cf n. 473 M.F., 17.VIII.l970; 1 cf n. 475 M.F., 18.VIII.1970; leg. S.B.S. Run, 1 cf n. 476 M.F., leg. S.B.S., 10.VIII Distribution: almost all continental Africa. Remarks: of the specimens examined, only the Ola Uager 9 M.F. n belongs to the form lucia. All the other refer to the form portia. 2. Palpopleura deceptor (Calvert, 1899). deceptor: FoRSTER,!906b, p. 304 (Haru Bu"a. along the Giuba ri\'cr, 2 cl cl Hemisti~moides one of which is n Ph.A.~.S., leg. C. Von Erlanger, V.1901 ). Pa/popleura deceptor: Rts, 1911, p. 326 (the citation refers to preceding data); NIELSEN, 1935, p. 75 (the citation refers to preceding data); PtNIIEY, 1961, p. 144 (the citation refer> to preceding data); Pt)';IIEY, 1962, p. 242 (the citation refers to preceding data). No specimens examined. Distribution: tropical Africa. Remarks: FORSTER (1906b) cited two among the other specimens as surely coming from Somalia (see above). The latter citations report these species as Somalian, not because of Forster's data but because Rrs ( 1910) interpreted the locality Sheikh Husein as Somalian for the Type of CALVERT (1899). Genus DIPLACODES Kirby, Displacodes lefebvrei (Rambur, 1842). Diplacodes tetra: FoRSTER, 1906b, p. 308 (Haru Bussa. I 0, 22.V.1901; along the Giuba: between Maika Bebirrissa and Lake Solule, I o, 3 I' I'. IO.VI.1901; Lake Solole, 2 I' I', II. VI. 1901; Umfudu, I 0, 18.VI.I901; Mansundu, I I', 3.VII.1901; leg. C. von Erlanger). Dip/acodes Lcfeb1 rei: NtELSE", 1935, p. 71 (Belet Amin, I 9 M.S.N.G., 19.V1.1934; I o, 2 I' I' M.S.N.G., 20.V1.1934; I o, 2 I' I' M.S.N.G., 3G.VI.l934; I I' C.N., 20.VI.1934; I o C.N., 30.VI.1934; I I' C.N., Vll.1934; leg. S. Patrizi. Ola Uager, I o M.S.N.G., leg. S. Patrizi, 15.VII1.1934); NtEL~EN, 1935, p. 75 (the citation refers to preceding data); NtELSE", 1936, p. 129 (Benadir. Merca Uagadi, I o M.S.N.G., leg. A. Pantano, 14.IX.1907). Diplacodes /efebvrei: Pt,.HH, 1962, p. 255 («Ita!. East Africa.. : the citation refers to preceding data). Afmadu, 1 cf n. 591 M.F., 1 9 n. 599 M.F., leg. S.B.S., Balad, 1 cf n. 586 M.F., leg. S.B.S., 13.VIII Gelib, 1 9 n. 588 M.F., leg. S.B.S., 4.IX Ghersale, 1 cf juvenile n. 584 M.F., leg. S.B.S., 22.IV Giohar, 2 cfcf n. 585 M.F., IV.1968; 1 cf n. 597 M.F., IV.1968; 6 cfcf, n. 595 M.F., 16.IV-l.V.1968; 2 cfcf, 1 9 n. 592 M.F., 30.VII.1968; 2 cfcf, 1 9 n. 596 M.F., 22-VIII.1968; 1 9 n. 590 M.F., 13.IX.1968; leg. S.B.S.

23 168 S. CARFl Mahaddei, 2 dd, 2 ~ ~ n. 594 M.F., 17.IX.1968; 1 ~ n. 593 M.F., 23.IX.1968; leg. S.B.S. Modun, 1 d n. 518 M.F., leg. S.B.S., VIII Mogadiscio (surrounding), 1 ~ n. 598 M.F., leg. S.B.S., 27.VIII Ola Uager, 1 ~ n. 514 M.F., 16.VIII.l970; 3 d d, 2 ~ ~ n. 513 M.F., 17.VIII.1970; 1 d n. 515 M.F., 18.VIII.l970; leg. S.B.S. Run, 1 d, 1 ~ n. 589 M.F., 5.VIII.1964; 2 ~ ~ n. 520 M.F., 7. VIII.1969; 1 ~ n. 522 M.F., 10.VIII.l969; 1 d n. 517 M.F., 11.VIII. 1969; 1 ~ n. 521 M.F., 11.VIII.l969; 1 ~ n. 523 M.F., 12.VIII.1969; 1 ~ n. 524 M.F., 19.VIII.l969; leg. S.B.S. Scialambod, 1 d n. 587 M.F., leg. S.B.S., 29-IV Distribution: almost all Africa, Madagascar, Southern Spain and Asia Minor. Genus CROCOTHEMIS Brauer, Crocothemis erythraea (Brulle, 1832). Orthetrum lorti KIRBY, 1896, p.522 (Dobar, Goolis Mts, I I? B.M., leg. Mr & Mrs Lort Phillips, 4.II.1895). Crocothemis erythraea: CALVERT, 1899, p. 234 (Berbera, 3 ~ ~, 4 I? I?, leg. Donaldson Smith, 4-5.VII.1894; Lafarok, 1 ~, leg. Donaldson Smith, 13.VII.1894); SHARPE. 1901, p. 8 (Bulhar, 2 specimens Bar. M., leg. Donaldson Smith, ); FoRSTER, 1906b, p. 311 (along the Giuba, ~ ~ & I? I? from the following localities: Salakle, 7.VI.1901; Dogge, 3.VI.1901; Umfudu, 18.Vi.1901; Bua, 28.VI.1901; Mansundu, 3-4.VII.1901; leg. C. von Erlanger); RIS, 1911, p. 536 (Somalia; the citation refers to the same specimen studied by KIRBY, 18%, Type of 0. lorti); LONGFIELD, 1931, p. 276 (Berbera, the same specimens studied by CALVERT, 1899; Lafarok, the same specimen studied by CALVERT, 1899; Dobar, Goolis Mts, the same specimen studied by KIRBY, 1896, Type of 0. lorti; Bulhar, the same specimens studied by SHARPE, 1901; Buran, 4 ~ ~, 1 I? B.M., leg. C. L. Collenette, IX-X.1929; Marojeh, 1 I? B.M., leg. C. L. Collenette, X1.1929); NIELSEN, 1935, p. 71 (Belet Amin, 3 ~ ~ M.S.N.G V.1934; I I? M.S.N.G., VII. 1934; leg. S. Patrizi); NIELSEN, 1935, p. 75 (the citation refers to preceding data); PINHEY, 1961, p. 158 [«Somalia, British Somaliland Protectorate (at Baidoa)», 28 specimens, C.M.]; PINHEY, 1962, p. 258 (the citation refers to preceding data). Giohar, 1 d n. 650 M.F., 10.VII-20.IX.1962; 1 d n. 535 M.F., 16. IV-l.V.1968; 4 d d n. 533 M.F., 22.VIII.1968; 1 d n. 530 M.F., VIII.l970; leg. S.B.S. Run, 2 d d, 1 ~ n. 536 M.F., 5.VIII.1964; 1 d n. 537 M.F., 19. V.1969; 1 d n M.F., 6.VIII.l969; 1 ~ n. 531 M.F., 11.VIII. 1969; 3 d d, 2 ~ ~ n. 532 M.F., 11.VIII.1969; 1 d n. 529 M.F., 12.VIII.l969; 1 ~ n. 528 M.F., 19.VIII.l969; leg. S.B.S. Scialambod, 1 ~ n. 538 M.F., leg. S.B.S., 29-IV Uadi Manas, 1 d n. 534 M.F., VII.1962; 1 d n. 649 M.P., VII.1962; leg. S.B.S. Distribution: all Africa and a large part of the palearctic region.

24 SOMALIAN ODONATA 169 Genus BRACHYTHEMIS Brauer, Brachythemis leucosticta (Burmeister, 1839). Cacergate leucosticta: McLACHLAN, 1900, p. 34 (Sinnadohgo, Haweea Country, I 11 H.C.O., leg. A. Peel, 8.IX.I897). Brachythemis teucosticta: NIELSEN, 1935, p. 71 (Belet Amin, 2 d d M.S.N.G., leg. S. Patrizi, 30.VI.l934); NIELSEN, 1935, p. 75 (the citation refers to preceding data); NIELSEN, 1936, p. 128 (Giumbo, I d M.S.N.G., leg. G. Ferrari, 13.VII.1907; Merca Plain, I d M.S.N.G., leg. A. Pantano, IO.X.1907); PINHEY, 1962, p. 261 (the citation refers to preceding data). Balad, 2 rf rf n. 567 M.P., leg. S.B.S., 30.VIII.l964. Eggi, 4 rf rf n. 563 M.P., leg. S.B.S., VII Giohar, 1 rf n. 565 M.P., 27.VIII.l964; 1 rf, 2 S' S' n. 571 M.P., IV.1968; 2 rfrf, 1 S' n. 568 M.P., 16.IV-l.V.1968; 8 rfrf, 2 S' S' n. 510 M.P., 7.VIII.l970; 3 rf rf, 1 S' n. 511 M.P., VIII.1970, leg. S.B.S. Ola Uager, 1 rf n. 566 M.P., 15.IX.1964; 2 rf rf, 1 S' n. 512 M.P., 17.VIII.l970; leg. S.B.S. Distribution: almost all Africa, Asia Minor, central Portugal, province of Malaga in Spain. Genus SYMPETRUM Newmann, Sympetrum fonscolombei (Selys, 1840). Sympetrum Fonscolombii (sic): FORSTER, 1906b, p. 307 (Ganale, near Dogge, I d, leg. C. von Erlanger, IO.VI.1901). Sympetrum Fonscolombei: NIELSEN, 1935, p. 75 (the citation refers to preceding data). Sympetrum fonscolombei: PINIIEY, 1962, p. 263 (the citation refers to the specimen cited by FoRsTER, 1906b ). No specimens examined. Distribution: almost all continental Africa, Europe and Asia. Genus PHILONOMON Forster, Philonomon luminans (Karsch, 1893 ). Philonomon Erlangeri FoRSTER, 1906a, p. 10 and 18 (Somaliland). FoRsTER, 1906b, p. 310, table A, fig. 4, along the Giuba at Gedud, I d, 2.VI.1901; Dogge, 2 d d, 9-10.VI 1901; Umfudu, 1 \1, 18.VI.1901; leg. C. von Erlanger). Philonomon luminans: NIELSEN, 1935, p. 71 (Belet Amin, I d M.S.N.G., leg. S. Patrizi, 30. VI.1934). NIELSEN, 1935, p. 75 (the citation refers to preceding data); PINHEY, 1961, p. 155 (British Somaliland, 26 specimens C.M.); PINHEY, p. 264 («... Somalia...»: the citation refers to preceding data).

25 170 S. CARFI Giohar, 1 d' n. 486 M.P., leg. S.B.S., 16.IV-l.V Ola Uager, 2 d'd' n. 488 M.P., 17.VIII.l970; 1 d' n. 487 M.P., 17.VIII.1970; leg. S.B.S. Distribution: almost all of continental Africa. Genus TRITHEMIS Brauer, Trithemis annulata (Palisot de Beauvois, 1805). Trithemis annulata: NIELSEN, 1935, p. 71 and 75 (Belet Amin, 2 & & M.S.N.G., 1934; 1 & M.S.N.G., 30.VI.l934; I 'i' M.S.N.G., VII.1934; 1 & C.N., 20.VI.l934; leg. S. Patrizi). PINHEY, 1961, p. 162 (Somalia, specimens C.M.); PINHEY, 1962, p. 268 (the citation rekrs to preceding data). Baidoa, 2 d' d' n. 556 M.P., leg. S.B.S. 27.VIII El Bur, 1 d' n. 648 M.P., leg. S.B.S., 3-9.IX Giohar, 1 d' C.N., leg. U. Piechter, 1936; 1 <? n. 647 M.P., 10.VII- 20.1X.1962; 1 d', 1 <? n. 558 M.P., 16.IV-l.V.1968; 2 d' d', 1 <? n. 552 M.F., 30.VII.1968; 2 d' d' n. 554 M.F., l.viii.1968; 1 <? n. 503 M.P., 18.VII.1970; 1 d' n. 506 M.P., 19.VII.1970; 4 d' d', 1 <? n. 508 M.F., 29.VII.1970; 2 <? <? n. 507 M.F., l.viii.1970; 1 d' n. 509 M.F., 6.VIII.1970; 1 d', 2 <? <? n. 505 M.P., VIII.1970; leg. S.B.S. Modun, 1 <? n. 504 M.P., leg. S.B.S., VIII Ola Uager, 1 <? n. 555 M.P., leg. S.B.S., 15.1X Run, 1 d' n. 553 M.P., 8.VIII.1964; 1 d' n. 560 M.F., 10.VIII.1969; 1 d' n. 559 M.P., 16.VIII.1969; leg. S.B.S. Uadi Manas, 1 d' n. 557 M.P., 1 d', 1 <? n. 646 M.F., leg. S.B.S., VII Distribution: throughout continental Africa, Madagascar, part of Asia Minor, Southern Italy, Sicily and Sardinia. 2. Trithemis arteriosa (Burmeister, 1839). Trithemis arteriosa: KIRBY, 1896, p. 521 (Dobar, Goolis Mts, 2 & & B.M., leg. Mrs Lort Phillips, ; Bichen, Somaliland, 1 & B.M., leg. Mr & Mrs Lort Phillips, ); LoNGFIELD, 1931, p. 275 (Dobar, Goolis Mts, the same specimens studied by KIRBY, 1896; Bichen, the same specimen studied by KIRBY, 1896; Bihen Andola, I & B.M., leg. Mrs Lort Phillips, ; Saugamore, 7 & & B.M., leg. Mrs Lort Phillips, 1897; Medisheh, I & B.M., leg. C. L. Collenette, IX.I929; Buran, 10 & &, 2 'i' 'i' B.M., leg. C. L. Collenette, IX-X.I929); NIELSEN, 1935, p. 75 (the citation refers to preceding data); NIELSEN, 1957, p. 33 (Toh, 2 & & M.S.N.G., leg. G. Scortecci, VI.l953); PINHEY, 1961, p, 164 (Somalia, specimens C.M.); PINHEY, 1962, p. 269 (the citation refers to preceding data). Trithemis Stuhlmanni: FoRSTER, 1906b, p. 311 (Lake Geile near the Giuba, I &, leg. C. von Erlanger, 15.VI.l901).

26 SOMALIAN ODONATA 171 No specimens examined: Distribution: all continental Africa, Syria. Remarks: though this Trithemis is considered to be the most widespread of the species in Africa, in Somalia - or rather, in the Somalian localities where the various «S.B.S.» have had occasion to collect - T. annulata seems to prevail. 3. Trithemis furva Karsch, Trithemis distanti distanti: LoNGFIELD, 1931, p. 276 (Medisheh, 3 d d B.M., leg. C. L. Collenette, IX-X.1929). Trithemis Distanti: NIELSEN, 1935, p. 75 (the citation refers in part to the specimens studied by LoNGFIELD, 1931). Medisheh, 2 d d B.M , leg. C. L. Collenette, 2 8.XII Distribution: perhaps throughout continental Ethiopian Africa. Remarks: two of the three specimens cited by LoNGFIELD (1931) located among the paratypes of T. risi Longfield, were examined. One of these bears a label reading: «On small stream descending rocky valley». I am deeply indebted to Sandra L. Bedford for their loan. 4. Trithemis kirbyi ardens Gerstaecker, Trithemis kirbyi ardens: LoNGFIELD, 1931, p. 276 (Saugamore, 3 d d B.M., leg. Mrs Lort Phillips, ; Medisheh, I d B.M., leg. C. L. Collenette, IX.1929); PINHEY, 1961, p. 163 (Somalia, 22 specimens C.M.); PINHEY, 1962, p. 272 (the citation refers to preceding data). Trithemis Kirbyi ardens: NIELSEN, 1935, p. 76 (the citation refers to preceding data). Baidoa, 1 d n. 652 M.P., leg. G. Stefanini & N. Puccioni, Uadi Manas, 1 d n. 651 M.P., leg. S.B.S., VIII Distribution: a large part of continental Ethiopian Africa, Madagascar, Com oro Islands. 5. Trithemis stictica (Burmeister, 1839). Trithemis stictica: LoNGFIELD, 1931, p. 276 (Medisheh, 3 d d B.M.. leg. C. L. Collenette, IX-XI.1929); NIELSEN, 1935, p. 76 (the citation refers to preceding data); PINHEY, 1962, p. 273 (... British Somaliland...», the citation refers to preceding data); PINHEY, 1970a, p. 125 (... Somalia...», the citation refers to preceding data). No specimens examined. Distribution: Madagascar. widely distributed in continental Ethiopian Afrioa,

27 172 S. CARFl Genus 0LPOGASTRA Karsch, Olpogastra lugubris (Karsch, 1895). Giohar, 1 <.;? n. 519 M.F., leg. S.B.S., VIII.l970. Distribution: Southwest Africa, Rhodesia, Mozambique, Tanzania, Congo, Uganda, Kenya, Nigeria, Cameroon, Chad, Sudan, Somalia. Remarks: according to my records, the specimen from Giohar is the first to be collected in Somalia. Genus RHYOTHEMIS Hagen, Rhyothemis semihyalina (Desjardins, 1835). Rhyothemis hemihyalina: FoRSTER, 1906, p. 303 (Ganale near Dogge, 1 a, leg. C. von Erlanger, 10.VI.1901). Rhyothemis semihyalina: NIELSEN, 1935, p. 72 and 76 (Belet Amin, I a M.S.N.G., I a C.N., ieg. S. Patrizi. IO.VII.I934); PINHEY, 1962, p. 281 (the citation refers to preceding data). specimens examined: Giohar, 1 d, 1 <.;? n. 547 M.F., IV.1968; 1 d n. 548 M.F., 3.VIII.l968; 1 d n. 549 M.F., VIII.1969; leg. S.B.S. Ola Uager, 1 d, 1 <.;? n. 551 M.F., leg. S.B.S., 17.VIII Distribution: almost all continental Africa; Madagascar. Genus THOLYMIS Hagen, Tholymis tillarga (Fabricius, 1798). Giamana (surroundings), 4 d d n. 516 M.F., leg. S.B.S., 24.VIII Giohar, 1 <.;? n. 540 M.F., leg. S.B.S., 2.VIII Distribution: tropical and subtropical belt of the world. Remarks: though this species is amply distributed geographically, it is the first time that it has been collected in Somalia, probably because of its crepuscular habits.

28 SOMALIAN ODONATA 173 Genus PANT ALA Hagen, Pan tala flavescens (Fabricius, 1798 ). Panta/a {lavescens: KIRBY, 1896, p. 251 (Dobar, Goolis Mts, 3 & d B.M., leg. Mr & Mrs Lort Phillips, ); SHARPE, 1901, p. 8 (Bulhar, 6 specimens Bar. M., leg. A. Donaldson Smith, ); LoNGFIELD, 1931, p. 275 (Dobar, Goolis Mts, the same specimens studied by KIRBY, 1896, 2 'i' 'i' B.M., leg. Mrs Lort Phillips, ; Bihen, I 'i' B.M., leg. Mrs Lort Phillips; Bulhar, the same specimens studied by SHARI'E, 1901; Somali land, I(Jui5'N- 45oJO'E, I d B.M., leg. M. Portal Hyatt, Xl.1928; Wadamago, I 'i' B.M., leg. C. L. Collenette, VIII.l929; Buran, 5 d &, 3 'i' 'i' B.M., leg. C. L. Collenette, IX-X.I929); NIELSEN, 1935, p. 72 (Belet Am in. 3 d d, I 'i' M.S.N.G., 1934; I &, I 'i' M.S.N.G., 20.VI.1934; leg. S. Patrizi. I & C.N., leg. S. Patrizi, Vl1.1934; Ola Uager, 2 & & M.S.N.G., leg. S. Patrizi, IS.VII1.1934); NIELSEN, 1935, p. 76 (the citation refers to preceding data); NIELSEN, 1936, p. 129 (Giumbo, I & M.S.N.G., leg. A. Pantano, I.VI.1907; specimen already determined by L. Navas); PISHEY, 1961, p. 178 (Somalia, British Somaliland Protectorate, various specimens C.M.); PINHEY, p. 285 (the citation refers to preceding data). Bur Duldir - Bugda Acable, 1 d, 1 '( n. 543 M.F., leg. S.B.S., 4.IX Giamama (surroundings), 1 d, 1 '( n M.F., leg. S.B.S., 24. VIII Giohar, 1 '( n. 539 M.F., IV.1968; 3 d d, 1 '( n. 544 M.F., 16.IV-l.V.1968; 1 d, 1 '( n. 541 M.F., 22.VIII.1968; 1 '( n. 546 M.F., 18.VII.1970; leg. S.B.S. Jesomma, 1 d, 1 '( n. 542 M.F., leg. S.B.S., VIII Ola Uager, 1 '( n M.F., leg. S.B.S., 17.VIII Distribution: this circum-tropical species is abundantly distributed throughout the world. Genus TRAMEA e) Hagen, Tramea basilaris (Palisot de Beauvais, 1805). Ola Uager, 2 '( '( n. 576 M.F., 16.VIII.1970; 1 d, 2 '( '( n. 575 M.F., 17.VIII.1970; leg. S.B.S. Distribution: this species is known in numerous localities of the Ethiopian region as well as in India, Ceylon, Birmania, Tibet and Malaysia. Remarks: the specimens examined are the first collected in Somalia even though this species was already known from nearby localities, such as Dahele (FORSTER, 1906b, p. 303 ). This locality, considered as «Djehle (l) GLOYD, K. L. (1972).

29 174 S. CARFl lower Juba» by NIELSEN, 1935, p. 76 and then «Somalia» by PrNHEY, 1962, p. 288, is not in Somalian territory. 2. Tramea limbata (Desjardins, 1832). Bud Bud, 1 cf n. 572 M.F., leg. S.B.S., 15.VIII Mogadiscio and Chismaio (between), 1 cf n. 574 M.F., leg. S.B.S., 8.VIII Distribution: this species, known from numerous localities in the Ethiopian region, is distributed across Asia as far as the Pacific Ocean. Remarks: the specimens examined are the first collected in Somalia. Genus UROTHEMIS Brauer, Urothemis assignata (Selys, 1872). Urothemis sanguinea designata: FORSTER, 1906b, p. 316 (Ghaghabli, along the Giuba, 2 & &, 4.VI.1901; Dogge, 1 &, 1 <;!, 9.VI.l901; Geile, 1 il, 1S.VI.l901; leg. C. von Erlanger). Urothemis assignata: NIELSEN, 1935, p. 76 (the citation refers to preceding data); NrELSnN, 1936, p. 129 (MERCA, 1 & M.S.N.G., leg. A. Pantano, 12.IX.1907); PINHEY, 1962, p. 291 (the citation refers to preceding data). Chisimaio (surroundings), 1 cf n. 604 M.F., leg. S.B.S., 19.VIII.l970. Giohar, 1 cf n. 608 M.F., 27.VIII.1964; 1 cf n. 609 M.F., IV.1968; 1 cf, 1 9 n. 607 M.F., 30.VIII.l968; n. 606 M.F., 8.IX.1968; 1 9 n M.F., 7.VIII.1970; leg. S.B.S. Giuba, 1 9 n. 610 M.F., leg. p,aoli, VII Modun (surroundings), 1 9 n. 603 M.F., leg. S.B.S., VIII Ola Uager, 1 cf n. 605 M.F., leg. S.B.S., 16.VIII.l970. Distribution: from Natal to East and West Africa. Remarks: among specimens in old collections I found the 9 n. 610, already determined by Dr C. Nielsen but never published. The left posterior wing of the cf n. 607 is deformed, perhaps due to an accident during emergence (Fig. 7). 2. Urothemis edwardsi (Selys, 1849). Giohar, 1 cf n. 602 M.F., 30.Vll.1968; 2 cf cf n. 600 M.F., 7.VIII.1970; leg. S.B.S. Distribution: tropical and subtropical Africa. Remarks: the specimens examined are the first collected in Somalia.

30 SOMALIAN ODONATA Urothemis signata aethiopica Nielsen Urothemis signata subsp. aethiopica NIELSEN, 1957, p. 33, fig. p. 34 (Toh, I is Type, 2 'i! 'i! Cotypes M.S.N.G., I 'i! Cotype n M.F., leg. G. Scortecci, 28.VI.1953). Urothemis bisignata aethiopica: PINHEY, 1962, p. 292 (the citation refers to the specimens studied by NIELSEN, 1957). No specimens examined. Distribution: the typical subspecies is known in India, Ceylon, Burma and Indochina. The subspecies aethiopica is so far known only from the type locality. Fig Urothemis assignata (Selys, 1872). o n. 607 M.P., Giohar (Somalia). Specimen with deformed left hindwing. Genus MACRODIPLAX Brauer, Macrodiplax cora Brauer, 1867). Bud-Bud, 1 cf n. 582 M.F., leg. S.B.S., 15.VIII Giohar, 1 cf n. 583 M.F., leg. S.B.S., 7.VIII Distribution: Socotra, Mauritius, Southern India, Ceylon, Southeast Asia, Samoan Islands, New Caledonia.

31 176 S. CARFI Remarks: the specimens examined are the first collected in Somalia. This is of particular interest because it widens the range of a typically oriental species towards the west. III. LIST OF THE LOCALITIES Afgoi ( = Afkoi ), Benadir region Afmadu ( = Afmedu), lower Giuba, about 110 km NNW of Chisimaio Ararchi Eggi (see Eggi) Arbarone ( = Habrona = Aproni ), about midway along the trail between El Uach and Bardera Baidoa ( = Iscia Baidoa), upper Giuba Balad (along the Uebi Scebeli river, about 40 km N of Mogadiscio) Bardera (upper Giuba) Belet Am in ( orographically, to the left of the Giuba river) Berbera (Migiurtinia, ex British Somaliland) Ber Birisso (see Maika Bebirrissa) Bichen (unidentified locality in Migiurtinia, ex British Somaliland, not far from Dobar) Bihen (unidentified locality in Migiurtinia, ex British Somaliland, as it is one of many homonymous localities) Bihen Andola (unidentified locality m Migiurtinia, ex British Somaliland. It may correspond to Bihendullah) Bihendullah (in Migiurtinia, ex British Somali land) Bua ( = Buba = Boa), lower Giuba Bud-Bud Bugda Acable Bulhar (in Migiurtinia, ex British Somaliland) 2 8'30"N '35"E 0 31' 3 "N '59"E 2 28' N ' E 3o 7' N ' ~ 2 2l'40"N '30"E 2 20' N ' E 10 10' N ' E 0 36' N ' E 4 12' N ' E 4 3'18"N '47"E

32 SOMALIAN ODONATA Buran (in Migiurtinia, ex British Somaliland) Bur Duldir (hills N of Bugda Acable, not far from Bulo Burti) Bu Sar (see Haro Bussa) Chisimaio ( = Kismayu), lower Giuba Dogge Dobar ( = Dubur = Dubar ), the maps I consulted show a Dubur = Dubar about 13 km S of Berbera and a Plane of Dubur about 70 km further S, at the foot of the Goolis Mountains Eggi ( = Ararchi Eggi) Einad (in Migiurtinia, ex British Somaliland) El Bur (Mudugh region) El Gambole (Southern Somalia) Eleschia (the map tracing Erlanger's itinerary - SPRIGADE, shows an Eleschid about 10 km N of Jonte and an Helischid about 13 km WNW of the former. Neither of these should be confused with Hele Scid, situated much farther N and well out of the indicated itinerary) Ganale (often used to indicate the upper course of the Giuba river) Garoe (Nogal Valley, Migiurtinia) Gedud (the map tracing Erlanger's Itin erary - SPRIGADE, shows a Maika Gedud and, nearby, a Maika Guele Gedud, both slightly WNW of Bardera) Geile, Lake (along the Giuba river, slightly S of the Solole hollow) Gelib ( = Jilib ), along the Giuba river, lower Giuba Ghaghabli '40"S '50"E 1 41' N ' E 11 2' N ' E 4 40' N ' E 3 4'20"N ' E 0 29' N '30" 2 10' N ' E

33 178 S. CARFI Ghersale (along the road between Mogadiscio and Giohar, about 25 km S of Giohar) Giamama ( = Margherita), along the Giuba river, lower Giuba Giohar ( = Giauher = Villaggio Duca degli Abruzzi = Villabruzzi ), along the Uebi Scebeli river. Benadir Giuba (river, Southern Somalia) Giumbo Habrona (see Arbarone) Haro Bussa ( = Haro Bussar Hun (see Run) Iscia Baidoa (see Baidoa) Jilib (see (Gelib) Bu Sar) Jesomma J onte (village on the trail between Chisimaio and Gelib, along the Giuba river) Kismayu (see Chisimaio) Lafarok ( = Lafarug) Mahaddei Uen ( = Mahaddei ), along the shores of the Uebi Scebeli river, Benadir Maika Bebirrissa ( = Ber Birisso ), along the Giuba river Maika Guele Gedud (see Gedud) Mansundu ( = Mans undo) Margherita (see Giamama) Marojeh (in Migiurtinia, ex British Somaliland) Medisheh (in Migiurtinia, ex British Somaliland) Merca (along the coast, Benadir region) Merca Uagadi (Uagadi is a village about 15 km NW of M(!rca) Modun (at the fork for Brava, along the road from Mogadiscio to Gelib) Mogadiscio 4 3' N ' E 0 7' s ' E 1003' N ' E 2 58'14"N ' E 1 40' N 42 26' E 0 6' N ' E 10 47' N 4r35' E 1 42'44"N '25" 1" 9' N 43 59' E 2 1 '58"N - 45"20'56"E

34 SOMALIAN ODONATA 179 Mufudu (see Umfudu) Ola Uager «Plain N of Al Hills» (in Migiurtinia, ex British Somaliland) Run ( = Hon = Hun), ghelta on the Uadi Run, at the base of the hills on the N side of the Nogal Valley, about 50 km NNE of Garoe Salakle Saugamore (unidentified locality in Migiurtinia, ex British Somaliland) Scialambod ( = Vittorio d'africa), on the road between Afgoi and Brava, not far from Merca Schimba Beris Sinnadohgo (Haweea Country) Solole, Lake (in the Solole hollow, near Gebbi) Toh (springs situated on the slopes of the Ahl Mascat mountains, about 15 km SW of Candala in Migiurtinia, 750 m a.s.l.) Uadi Manas (a water hole on the trail between Baidoa and Bardera, about 32 km S of Baidoa) Umfudu ( = Mufudu), on the left bank of the Giuba river Villaggio Duca degli Abruzzi ( = Villabruzzi, see Giohar) Vittorio d'africa (see Scialambod) Wadamago Wadi Mait (unidentified locality in Migiurtinia, ex British Somaliland) 1 40' 10 45' 5 23' N ' N ' N ' N ' N ' E E E E E E E E E E E E SUMMARY The author has studied the Odonata collected in the Somali Democratic Republic during scientific expeditions organized first by the Institute of Zoology and by the Museum of Zoology of the University of Florence

35 180 S. CARFl and, presently, by the «Centro di Studio per la Faunistica ed Ecologia Tropicali del C.N.R.». He has also included a list as complete as possible of all the quotations on Somalian Odonata. According to the literature and specimens studied, the Somalian Dragonflies pertain to seven families, 30 genera (seven of which are new for Somalia) and 53 species ( 15 Zygoptera and 38 Anisoptera), 12 of which are new for Somalia: Lestes tridens McLachlan, Lestes uncifer Karsch, Agriocnemis exilis Selys, Pseudagrion acaciae Forster, Ictinogomphus ferox (Rambur), Tetrathemis polleni (Selys), Olpogastra lugubris (Karsch), Tholymis tillarga (Fabricius), Tramea basilaris (P. de Beauvois), Tramea limbata (Desjardins), Urothemis edwardsi (Selys), Macrodiplax cora (Brauer). RIASSUNTO L' Au tore ha studiato gli Odonati raccolti nella Repubblica Democratica Somala durante le spedizioni biologiche organizzate prima dall'istituto di Zoologia e dal Museo Zoologico dell'universita di Firenze e quindi dal «Centro di Studio per la Faunistica ed Ecologia Tropicali del C.N.R.». Inoltre ha cercato di riunire, in un quadro il piu completo possibile, tutte le citazioni sugli Odonati della regione. Dalla letteratura e dal materiale esaminati risulta che l'ordine degli Odonati comprende, nella regione considerata, 7 famiglie, 30 generi (di cui 7 nuovi per la Somalia) e 53 specie ( 15 Zigotteri e 38 Anisotteri), delle quali le 12 seguenti risultano essere nuove per la Somalia: Lestes tridens McLachlan, Lestes uncifer Karsch, Agriocnemis exilis Selys, Pseudagrion acaciae Forster, Ictinogomphus ferox (Rambur), Tetrathemis polleni (Selys), Olpogastra lugubris (Karsch), Tholymis tillarga (Fabricius), Tramea basilaris (P. de Beauvois), Tramea limbata (Desjardins), Urothemis edwardsi (Selys), Macrodiplax cora (Brauer). REFERENCES CALVERT, PH. P Neuropterous insects collected by Dr. A. Donaldson Smith in Northeastern Africa. Proc. Acad. nat. Sci. Philad. 1: , table X. ERLANGER, C. FRHR. VON Bericht iiber meine Expedition in Nordost-Afrika in den Jahren Z. Ges. Erdk. Berl.: , tables 2-5. FoRSTER, F a. Die Libellulidengattungen von Afrika und Madagascar. Jber. Ver. Naturk. Mannheim 71-72: FoRSTER, F b. Forschungereise durch Siidschoa, Galla und die Somalilander von Carlo Freiherr von Erlanger. Libellen. ]b. nassau Ver. Naturk. 59: , table A. GLOYD, K. L Tramea, Trapezostigma and time (Anisoptera: Libellulidae) a nomenclatural problem. Odonatologica: 1: