POLYGALACEAE MILKWORT FAMILY

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1 POLYGALACEAE MILKWORT FAMILY Plant: herbs, shrubs and trees, rarely vines Stem: sometimes square Root: Leaves: simple, entire, mostly alternate to rarely opposite or whorled; no stipules but sometimes glands present Flowers: perfect; many irregular (zygomorphic), often with many tiny flowers; 5 (4-6-7) usually separate sepals with 2 lateral which are colored (wings); 3 or 5 petals connected into a tube, often fringed; 3-10 stamens, usually 8 (in sets of 4); ovary superior, usually 2 carpels Fruit: capsule usually, 2-chambered, seeds often hairy; others types sometimes Other: Dicotyledons Group Genera: 17+ genera; locally Polygala (milkwort) WARNING family descriptions are only a layman s guide and should not be used as definitive

2 Flower Morphology in the Sepals (5) 2 sometimes larger (not always) colored (petallike) inner sepals termed the wings. Outer 3 sepals often much smaller (colored or not). Petals (3) petals fused at base (boat-like) to almost totally fused (tube-like) in some species and surrounds the stamens and style. The lower petal (keel) often fringed at apex. An Outer Sepal 2 Wings Fringe of Keel Petal 3 Petals forming tube Racemed [Purple] Milkwort Polygala polygama Walt. In this case all sepals and petals are colored and more similar in size, the keel is fringed Pink Milkwort Polygala incarnata L.

3 POLYGALACEAE MILKWORT FAMILY Cross Milkwort [Drumheads]; Polygala cruciata L. var. Aquilonia Fern. & Schub. Pink Milkwort [Procession Flower]; Polygala incarnata L. Orange Milkwort [Red-Hot Poker]; Polygala lutea L. Candyroot; Polygala nana (Michx.) DC. Racemed Milkwort; Polygala polygama Walt. Field [Purple] Milkwort; Polygala sanguinea L. Seneca Snakeroot; Polygala senega L.

4 Cross Milkwort [Drumheads] Polygala cruciata L. var. aquilonia Fern. & Schub. Kitty Todd Nature Preserve, Lucas County, Ohio Notes: 3 petaled flower, one fringed and with other 2 forming a tube; 5 sepals, 2 larger side sepals and 3 USDA smaller ones with sharp-points and deltoid-like bases, flowers rose-purple to somewhat greenish in dense head; leaves 3-4 whorled, linear to somewhat lanceolate, thick; often in sandy soils; summer to early fall [V Max Brown, 2007]

5 Pink Milkwort [Procession Flower] Polygala incarnata L. Chaparral Prairie State Nature Preserve, Adams County, Ohio USDA Notes: 3-petaled flower, one fringed, pink, to pinkish purple to white, 5 colored sepals, in terminal clusters or racemes; leaves alternate, linear, usually gone at flowering time or reduced to scales; stem glabrous, somewhat glaucous, often branched; summer to fall [V Max Brown, 2009]

6 USDA Orange Milkwort [Red-Hot Poker] Polygala lutea L. Cedar Key Scrub State Reserve, Levy County, Florida Notes: flower petals orange, united to form a tube, sepals colored, in compact racemes; leaves alternate, basal leaves succulent and spathulate, stem leaves more lanceolate; plant rarely over 6 in high; spring to fall [V Max Brown, 2011]

7 USDA Candyroot Polygala nana (Michx.) DC. Cedar Key Scrub State Reserve, Levy County, Florida Notes: flower petals and sepals lemon yellow, petals united to form a tube, in compact racemes; leaves mostly basal, numerous, mostly sessile, succulent and spathulate; plant rarely over 6 in high; spring to early summer [V Max Brown, 2011]

8 USDA Racemed [Purple] Milkwort Polygala polygama Walt. Oak Openings Metropark, Lucas County, Ohio Notes: 3 petaled flower, one fringed and shorter than 2 side petals, sepal wings large, pinkish-purple to rarely white, on short pedicle; leaves alternate, linear to lanceolate; stems mostly unbranched; summer [V Max Brown, 2005]

9 USDA Field [Purple] Milkwort Polygala sanguinea L. Oak Openings Metropark, Lucas County, Ohio Notes: 3-petaled flower, rose-purple, greenish to white, 5 sepals, in dense oblong head; leaves alternate, numerous, and linear to lanceolate; stem often unbranched but not always; summer to fall [V Max Brown, 2005]

10 USDA Seneca Snakeroot Polygala senega L. Alley Springs, Ozark National Scenic Riverways, Shannon County, Missouri Notes: 3 petaled flower, one fringed, forming tube, 2 colored wing sepals, 3 smaller, flower white to greenish white, in racemes; leaves alternate, lanceolate and tapering at both ends; stems mostly unbranched; late spring to summer [V Max Brown, 2009]

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