Chemical components of wood

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1 Chemical components of wood Cellulose beta 1-4 glucan Extractives - various organics Lignin - Phenol propanoid monomers

2 Straight from Wikipedia The three types of reaction catalyzed by cellulases:1. Breakage of the non-covalent interactions present in the crystalline structure of cellulose (endo-cellulase) 2. Hydrolysis of the individual cellulose fibers to break it into smaller sugars (exo-cellulase) 3. Hydrolysis of disaccharides and tetrasaccharides into glucose (beta-glucosidase).

3 Chemical components of wood Cellulose beta 1-4 glucan Extractives - various organics Lignin - Phenol propanoid monomers

4 A fuzzy image of lignin from Wikipedia

5 Wood X-section (hardwood) Wood X-section (conifer)

6

7

8 Distribution of water and nitrogen in wood

9 Chemical components of wood Cellulose beta 1-4 glucan Extractives - various organics Lignin - Phenol propanoid monomers

10 Why the CODIT theory is all wet thanks to Rayner and Boddy

11

12 Echinodontium tinctorium Indian paint fungus a fungus that enters branch stubs and waits for years to cause heartrot

13 Hypoxylon an Ascomycete that decays wood and colonizes endophytically

14 Diagram of Hypoxylon spore germination in response to bark exudates - work by Ignatio Chapella

15 Access from one individual tree to another Hymenochaete corrugata - the glue or the bondage fungus

16 Brown rots Fomitopsis pinicola conifers Oligoporus amarus (incense cedar only) Oligoporus sequoiae (coastal redwood only) O. balsameus (Cupressus spp.) Laetiporus sulphureus (wide host range, but esp. eucalyptus and oak)? Phaeolus schweinitzii resinous conifers Heartrots on living trees White rots Phellinus pini (pines, Douglas-fir, & others) Echinodontium tinctorius (true fir and hemlock) Ganoderma applanatum (primarily hardwoods, Oak etc.) Cryptoporus volvatus ( conifers) Sterum hirsutum (hardwoods) Trichaptum abietinum (conifers) Trametes versicolor (hardwoods) Armillaria mellea gr. Heterobasidion annosum Phellinus weirii Dead trees Dead trees

17 Phellinus pini gr. red ring rot See this on the street side of Tolman Hall

18 The genus Phellinus (& Inonotus) has setae, and a brown hymenium

19 Ganoderma a common heartrot, white rot

20 Brown rots Fomitopsis pinicola conifers Oligoporus amarus (incense cedar only) Oligoporus sequoiae (coastal redwood only) O. balsameus (Cupressus spp.) Laetiporus sulphureus (wide host range, but esp. eucalyptus and oak)? Phaeolus schweinitzii resinous conifers Saprobes on living trees White rots Phellinus pini (pines, Douglas-fir, & others) Echinodontium tinctorius (true fir and hemlock) Ganoderma applanatum (primarily hardwoods, Oak etc.) Cryptoporus volvatus ( conifers) Sterum hirsutum (hardwoods) Trichaptum abietinum (conifers) Trametes versicolor (hardwoods) Armillaria mellea gr. Heterobasidion annosum Phellinus weirii Dead trees Dead trees

21 Zone lines in wood cause by vegetative interactions between different genotypes of decay fungi

22 Wood endophytes like Hypoxylon get first dibs

23 Trichaptum

24

25 Saprobes & pathogens that can persist as sabrobes Brown rots Oligoporus amarus (incense cedar only) Oligoporus sequoiae (coastal redwood only) O. balsameus (Cupressus spp.) on living trees White rots Phellinus pini (pines, Douglas-fir, & others) Echinodontium tinctorius (true fir and hemlock) Fomitopsis pinicola conifers Laetiporus sulphureus (wide host range, but esp. eucalyptus and oak)? Phaeolus schweinitzii resinous conifers Ganoderma applanatum (primarily hardwoods, Oak etc.) Cryptoporus volvatus ( conifers) Sterum hirsutum (hardwoods) Trichaptum abietinum (conifers) Trametes versicolor (hardwoods) Armillaria mellea gr. Heterobasidion annosum Phellinus weirii Dead trees Dead trees

26 edge of root disease center note progressively thinner crowns and shorter heights

27 Lion s tailing A crown symptom caused by lack of expansion of shoot and lower needle retention

28 Tree failure, a symptom of root decay Heterobasidion annosum P-strain in action at Yosemite village

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