THE RESPONSE TO NaCl AND IONIC CONTENT OF SELECTED SALT-TOLERANT AND NORMAL LINES OF THREE LEGUME FORAGE SPECIES IN SAND CULTURE
|
|
- Jessica Douglas
- 5 years ago
- Views:
Transcription
1 New Phytol. (1986) 104, THE RESPONSE TO NaCl AND IONIC CONTENT OF SELECTED SALT-TOLERANT AND NORMAL LINES OF THREE LEGUME FORAGE SPECIES IN SAND CULTURE BY M. A S H R A F, T. M C N E I L L Y* AND A. D. BRADSHAW Department of Botany, University of Liverpool, P.O. Box 147, Liverpool L69 3BX, UK {Accepted 15 July 1986) SUMMARY Selected NaCl-tolerant lines of Trifolium alexandrinum L., Medicago sativa L., and Trifolium pratense L. produced significantly greater dry matter than unselected control lines after 4 weeks of growth in sand culture with different NaCl concentrations. Tolerant lines of all three species generally contained less Na^ but more Cl^ in their shoots than normal lines, although these differences were not statistically significant. Selected lines also contained less K""" in their shoots than unselected lines. Ca"^ distribution was similar in T. "lexandrinum and M. sativa but not in T. pratense, in which the tolerant line had significantly higher Ca^+ in the shoot than the unselected line. Key words: Ion content, salinity, selection, NaCl tolerance, legume species. INTRODUCTION The need to produce crops with enhanced tolerance to high salt levels in the soil Has been emphasized by many workers (Dewey, 1962; Shannon, 1979; Epsteiti, 1985). The success of this approach depends upon the occurrence of appropriate genetic variability in crop species and the ability to exploit such variability using convenient selection techniques. It is important that selection at one stage of the life-cycle for increased tolerance should select those individuals also tolerant in the remaining stages. It has, however, been argued (Shannon, 1979) that the response of plants to salinity varies at different stages of the life cycle. In some species, tolerances exhibited at all growth stages are highly correlated, as for example in Medicago sativa L. (Noble, 1983). In others this may not be the case. The work described here assesses both the response to NaCl and the pattern of inorganic ion distribution in normal and salt-tolerant lines of Trifolium alexandrinum L., M. sativa L., and Trifolium pratense L. produced by selection at the seedling stage after 2 weeks of growth in saline nutrient solution. These three species are commonly used as forage crops in several countries where soil salinity is a major problem for agriculture. T. alexandrinum has been categorized as moderately tolerant, while M. sativa and T. pratetise are intermediate in salt tolerance compared with other legumes (Lauchli, 1984). Patterns of distribution of Na+, K+, Ca-+, Mg^^ and Cr within the plant after 19 days of growth in NaCl solution culture have previously been described by Winter & Lauchli (1982) for normal material of T. alexandrinum and T. * To whcun reprint requests should be addressed X/86/tlO $03.00/ The New Phytologist
2 464 M. ASHRAF, T. MCNEILLY AND A. D. BRADSHAW pratense. This paper compares the distribution of these ions in selected NaCltolerant lines and unselected control lines of these two species, and of M. sativa in response to NaCI after 28 days of growth in sand culture. MATERIALS AND METHODS Plant material Salt-tolerant lines were selected from seeds of commercial varieties of T. alexandrinum L. cv. Multifoliate, M. sativa L., cv. Euver and T. pratense L. cv. Altaswede on the basis of differences in shoot length after 2 weeks of growth in saline solution cultures (Ashraf, McNeilly & Bradshaw, unpublished data). Twenty plants were raised at the same time from unselected seeds of the same three cultivars. The plants of both the selected and unselected lines were grown to maturity in a glasshouse at 18 C and a 16 h photoperiod, and they were manually crossed at random in separate polycrosses. The resulting progeny of each line were harvested separately. Sand culture Silica sand was soaked in 2% (v/v) HCl for 5 days and then thoroughly washed with tapwater and rinsed three times with distilled deionized water. Eighteen centimetre diameter plastic pots were filled with 3-54 kg dry sand. The main object of these experiments was the comparison of lines within species and not species comparisons, so each species was considered independently. The concentrations of NaCI used were 0, 100 and 175 mol m"^. The experiment had three blocks for each species, each block having two lines and three concentrations of NaCI. One hundred seeds of each line progeny of each species were germinated on moist filter paper in Petri dishes, and five randomly selected seedlings of each line were transplanted equidistant from each other into each pot. The seedlings of both lines were of comparable size when transplanted into the pots. The experiment was conducted in a heated glasshouse at 18 C and a 16 h day length. The seedlings were grown for 2 weeks, irrigated with half-strength nutrient (Rorison) solution (Hewitt, 1966), after which time NaCI treatments were begun by adding aliquots of a 50 mol m"-' solution of NaCI in 1 1 of the half-strength nutrient solution as above, on alternate days, until the appropriate NaCI concentrations were reached. Treatments continued with addition of 1 1 of the appropriate solution on alternate days to each pot. This volume was sufficient to wash through solution already present in the sand as determined from the electrical conductivity of the effluent solution from the pots. The plants were harvested 28 days after the start of the salt treatment. Plant roots were removed carefully from the sand, shoots and roots were separated and washed with distilled deionized water. Plant material was dried at 65 C for 3 days, divided into shoots and roots and weighed. Chemical analysis Cations. Ten to twenty milligram samples of leaf, stem and root dry matter were digested in 1 ml concentrated HNO3. After digestion was complete, the sample volume was made up to 5 ml with distilled deionized water, and all the cations were determined by atomic absorption spectrometry using a Varian 1275.
3 Ion uptake and salt tolerance 465 s Z c o in * Z i I S O c.2 " O 2 6 ta t; _ Z S 7 1 ro ca b do i'^ 00 ^ CO S 3 c U c.-i to <u o o V ft, ^ t3 QJ U O C O - O > -S o o o a, Iv oi. Po» 6 ^.^ Z*
4 466 M. ASHRAF, T. MCNEILLY AND A. D. BRADSHAW Chloride. Ten milligram subsamples were extracted by boiling in distilled deionized water for 3 h, and Cl^ content of the extracts were determined with a CMT 10 chloride titrator (Radiometer). Analyses of variance were carried out separately for the ion content of each plant part within each species O NaCl concentrations in sand culture (mol m"^) Fig. 1. Mean dry weights of plants from selected and unselected lines of three legume species [Trifolium alexandrinum (a), Medicago sativa (b), Trifolium pratense (e)] after 4 weeks of growth in sand culture at different NaCl concentrations. %, selected line; O, unselected line. RESULTS Mean dry weight per plant The results for mean dry weight per plant (Fig. 1) and their analyses of variance (Table 1) show that dry matter yield of both lines of all three species declined linearly with increasing NaCl concentration {P < 0-001). However, the selected line of each species produced significantly greater dry matter than the unselected line (T. alexandrinum, P<0-05; M. sativa, P < 0-001; T. pratense, P < 0-01). The selected lines of T. alexandrinum, M. sativa and T. pratense yielded about 25 %, 30 % and 45 % respectively more dry matter than the unselected lines at 175 mol m^'. The lines x concentration interaction was only significant in M. sativa (P < 0-05). This may have been due to the large differences in dry weight between the lines at 0 and 100 mol m ' NaCl. Shoot/root ratio Dry weight was also expressed as shoot/root ratios (Table 2). There were no significant differences in shoot/root ratio in T. alexandrinum and T. pratense, but a significant difference between the lines (P < 0-01) was found in M. sativa. The unselected line of M. sativa had a higher shoot/root ratio than the selected line, suggesting a more adverse effect in root dry weight in the former. The shoot/root ratio of the T. pratense unselected line was also greater than that of the selected line at 175 mol m^'', although the difference was not statistically significatit. Ion content Sodium. The Na+ content (Fig. 2) of the roots of the selected tolerant line of T. alexandrinum was significantly greater than in the unselected line (P < 0-01), and the leaves of the selected line contained less Na"*" than those of the unselected
5 Ion uptake and salt tolerance 467 line, but this difference was not statistically significant. The Na+ content in the stems of the two T. alexandrinum lines did not differ. By contrast, the different plant parts of selected and unselected lines of M. sativa had similar Na"*" contents. In T. pratense, the Na+ content was lower in both the leaves and the stems of the selected line than in the unselected line, although this difference was again not statistically significant. The Na"*" contents in the three species were in the order T. pratense > M. sativa > T. alexandrinum.. Chloride. The Cl" contents (Fig. 3) of the leaves and stems were generally higher in the selected than in the unselected lines of all three species, although the o S NaCI concentrations in sand culture [mol m~^) Fig. 2. Na^ contents in different parts of plants from selected and unselected lines of three legume species [Trifolium alexandrinum (a), Medieago sativa (b), Trifotium pratense (c)] after 4 weeks of growth in sand culture at different NaCI concentrations. Selected (#) and unselected (O) leaves; selected ( ) and unselected (D) stems; selected (A) and unselected (A) roots O NGCI concentrations in sand culture (mol m -) Fig. 3. Cl~ contents in different parts of plants from selected and unselected lines of three legume species [Trifolium alexandrinum (a), Medieago sativa (b), Trifolium pratense (c)]; details as in Figure 2.
6 468 M. AsHRAF, T. MCNEILLY AND A. D. BRADSHAW differences were not statistically significant. For the roots, the selected line of T. alexandrinum absorbed more CI" at 100 mol m~-* than the unselected line and the lines X concentration interaction was significant (P < 0-05). There were diflferences between species in patterns of CI" content within the plant. T. alexandrinum leaves contained less Cr than the stem, while in other species the reverse situation was found. Overall, Cl~ contents in the leaves of T NoCI concent rolions m sond culture (mol Fig. 4. K+ contents in different parts of plants from selected and unselected lines of three legume species [Trifolium alexandrinum (a), Medicago sativa (b), Trifolium pratense (c)]; details as in Figure 2. m"^) alexandrinum were lower than those in M. sativa, which in turn were lower than those in T. pratense. Roots of all three species had lower Cl~ contents than the stems or leaves. T. alexandrinum and M. sativa retained almost the same amount of CI, i.e. lower than that found in T. pratense. Potassium. The leaves and stems of the selected lines of T. alexandrinum and M- sativa contained less K+ overall (Fig. 4) than those of unselected lines {P < 0-01). In T. pratense, all plant parts in the selected line again retained less K"*", but this difference was not statistically significant. The three species showed different patterns of K+ accumulation. In M. sativa, roots retained more K^ than the other parts, while in T. alexandrinufjj and T. pratense, shoots were the K"''-accumulating organs. The order of accumulation of K"'' was T. pratense > M. sativa > T. alexandrinum. Little difference in the accumulation of K+ between plant parts was found in T. alexandrinum and M. sativa, but the differences in accumulation by different plant parts were highly significant in T. pratense. Calcium. The T. alexandrinum selected line had less (not statistically significant) Ca^+ in its leaves at 175 mol m"'' than the unselected line but did not differ at 0 and 100 mol m"' NaCI (Fig. 5). In M. sativa, the selected line contained significantly less Ca''^^ than the unselected line in both leaves and stem {P < 0-01). In T. pratense, the leaves and stems of the selected line contained significantly greater amounts of Ca^* (leaves, P < 0-05; stem, P < 0-001) than the unselected
7 Ion uptake and salt tolerance _ NaCi concentrations in sand culture (mol Fig. 5. Ca" contents in different parts of plants from selected and unselected lines of three legume species [Trifolium alexandrinum (a), Medicago sativa (b), Trifolium pratense (c)]; details as in Figure (a) A \ m. '~ '~ ~O S ^ _j ^ O ^ 0 NaCI concentrations in sand culture Imol m Fig 6 Mg" contents in different parts of plants from selected and unselected lines of three legume species [Trifolium alexandrinum (a), Medicago sativa (b), Trifolium pratense (c)]; details as in Figure 2. ') line. A reverse situation was observed in the roots, the unselected line containing more Ca^^ than the selected line (P < 0-01). Comparing the species, the Ca2+ contents were in the order T. pratense > M. sativa > T. alexandrinum. Magnesium. The Mg contents (Fig. 6) of the leaves and roots of the two T. alexandrinum lines did not differ significantly, but the stems of the unselected line contained more Mg'^+ than the selected line (P < 0-001). Similarly, the selected line of M. sativa contained less Mg2+ in the leaves and stem than the unselected
8 47 M. ASHRAF, T. MCNEILLY AND A. D. BRADSHAW lines (leaves, P < 0-001; stem, P < 0-01), but accumulation in the roots of the two lines did not differ significantly. In T. pratense, the selected line contained less Mg^"*" in its leaves than the unselected line, and an increase of NaCI in the external medium had a very pronounced effect in reducing the internal Mg^"'' content by approximately 50% and 60% in the selected and unselected lines respectively. DISCUSSION The selected lines had been produced from the progeny of individuals showing superior lengths of shoot at the seedling stage after two weeks of growth in nutrient solution containing NaCI. These produced significantly greater yields of dry matter than the unselected lines in all three species. It has been argued that, in some species, tolerance at the seedling stage may not reflect tolerance at the adult stage (Akbar & Yabuno, 1974; Shannon, 1979). On the other hand, the seedling stage has been considered more sensitive than the adult stage (Pasternak, Twersky & De Malach, 1979; Noble, 1983), and it has been argued that to improve the salt tolerance of a species, tolerance at the most sensitive stage must be increased (Noble, 1983). Whether selection is carried out at the adult or at early stages, the important factor is the intensity of selection which can be applied for isolation of tolerant individuals. The NaCl-tolerant lines produced for use in this study were the product of selection of a very few individuals out of a total of at least seedlings of normal cultivars of the three species. The individuals selected were 0-32%, 0-76% and 0-80% of the total population in T. alexandritium, M. sativa and T. pratense, respectively. Differences in shoot length were used as the selection criterion in these dicotyledonous species following the proposal of Rozema & Visser (1981). Lines of M. sativa with enhanced NaCI tolerance produced by selection have recently been described by Noble (1983). These were the product of seleetion of adult plants from samples of only 500 individuals, but nevertheless they gave correlated NaCI tolerance at all other growth stages. The procedure outlined in the present work is based upon the screening of seedlings. Clearly, this method can also produce individuals whose progeny show enhanced tolerance in T. alexandrinum, T. pratense, as well as in M. sativa. Mesophytes respond to saline environments either by excluding salt and synthesizing organic solutes, or by accumulating high concentrations of electrolytes to maintain their turgor (Elzam & Epstein, 1969; Greenway & Munns, 1980). Although the selected line of T. alexandrinum had higher Na"*" and Cl" contents in the roots compared with those of the unselected line, it excluded Na^, but had higher Cl~ in the leaves. Similar results have been observed for the salt-tolerant soybean variety 'Lee' (Lauchli & Wieneke, 1979), although the leaves of tolerant T. alexandrinum line had a higher Cl~ content. The selected line also had lower K^, Ca^^\ and Mg^+ eontents in its tissues, in spite of the important role of the former two ions in maintaining the turgor of plants under saline stress. In M. sativa, the selected and unselected lines did not differ in Na"*" and K"*^ content, although the selected line had higher Cl~ in the leaves at high NaCI concentration. This line also had less Ca"'^^ in its shoot compared with the unselected line. The results for Cl" content agree with those of Croughan, Stavarek & Rains (1978) who found that, as in a typical halophyte, a selected NaCl-tolerant line of M. sativa from tissue culture accumulated Cl~. The tolerant line of T. pratense had less Na"*" in both leaves and stems, but took
9 Ion uptake and salt toleranee up less K+ and Mg''^+ and had higher Ca^+ contents in its shoot. This species is more sensitive to salt than T. alexandrinum (data presented here, and that of Winter & Lauchli, 1982) and M. sativa. The lower Na+ content in the shoot of the tolerant line is again similar to the pattern in salt-tolerant soybean (Lauchli & Wieneke, 1979). If a forage species accumulates high quantities of salt in its tissue, it may not be suitable for fodder (Downton, 1984; Epstein, 1985). However, the tolerant lines of all the three species examined here showed a considerable variability in ion distribution compared with non-tolerant lines. Further investigation of two basic points of great importance is clearly needed. First, if further cycles of selection are carried out in the same way, lines of these species that have further increases in NaCI tolerance may show a greater tendency to exclude Na"*" from their tissues. Second, it is crucial to investigate plant-to-plant variability in Cl~ or Na"^ and to select CI" or Na+ excluders from tolerant lines for use in further breeding programmes. ACKNOWLEDGEMENTS The authors gratefully acknowledge financial support to M. Ashraf through an Overseas Research Studentship Award and a University of Liverpool Research Studentship. REFERENCES AKBAR, M. & YABUNO, T. (1974). Breeding for saline-resistant varieties of rice. U. Comparative performanee of some riee varieties to salinity during early development stages. Japan Journal of Breeding, 24, CROUGHAN.T. P., STAVAREK, S. J. & RAINS, D. W. (1978). Selectionof a NaCI tolerant line of cultured alfalfa eells. Crop Scienee, 18, DEWISY, D. R. (1962). Breeding crested vvheatgrass for salt tolerance. Crop Science, 2, DOWNTON, W. J. S. (1984). Salt tolerance of food crops: perspectives for improvements. CRC Critical Review in Plant Sciences, 1, ELZAM, O. E. & EPSTEIN, E. (1969). Salt relations of two grass species differing in salt tolerance. I. Growth and salt content at different salt concentrations. Agroctiimica, 13, EPSTEIN, E. (1985). Salt-tolerant crops: origins, development, and prospects of the concept. Plant and Soil, 89, GREENAWAY, H. & MuNNS, R. (1980). Mechanisms of salt tolerance in nonhalophytes. Annual Reviezv of Plant Physiology, 31, HEWITT, E. J. (1966). Sand and Water Culture Methods used in ttie Study of Plant Nutrition, 2nd edn. Commonwealth Agricultural Bureau. Technical Communication No. 22. LAUCHLI, A. (1984). Salt exclusion: an adaptation of legumes for pastures and saline eonditions. In: Salinity Tolerance in Plants: Strategies for Crop Improvement. (Ed. by R. C. Staples & G. H. Toeniessen), pp John Wiley & Sons, New York. LAUCHM, A. & WIENEKE, ]. (1979). Studies on growth and distribution of Na+, K+ and CI" in soybean varieties differing in salt tolerance. Zeitschrift fiir Pflanzenernahrung und Bodenkunile, 142, NOBLE, C. L. {\983). Genetical and ptiysiological aspects of.uilt-tolerance in lucerne {Medicafro sativa L.). Ph.D. thesis. University of Melbourne, Australia. PASTERNAK, D., TWERSKY, M. & DE MAI.ACH, Y. (1979). Salt resistance in agricultural crops. In: Stress Phvsiology in Crop Plants (Ed. by H. Mussel & R. C. Staples), pp John Wiley & Sons, New YoVk. RozEMA, J. & Vissia!, M. (1981). The applicability of the rooting technique measuring salt resistance in populations of Festuca rubra and Juncus species. Plant and Soil, 62, SHANNON, M. C. (1979). In quest of rapid screening techniques for plant salt tolerance. HortScience, 14, WINTER, E. & LAUCHLI, A. (1982). Salt tolerance of Trifolium alexandrinum L. 1. Comparison of salt response of T. alexandrinum and T. pratense. Australian Journal of Plant Physiology, 9,
10
Heterotic Effects in Spring Wheat (Triticum aestivum L.) Under Saline Conditions
Pakistan Journal of Biological Sciences, 1 (3): 159-162, 1998 Research Article Heterotic Effects in Spring Wheat (Triticum aestivum L.) Under Saline Conditions M. Ahsan* and D. Wright School of Agriculture
More informationEffect of saline culture on the concentration of Na +, K + and Cl in Agrostis tolonifera
Current Research Journal of Biological Sciences 2(1): 76-82, 2010 ISSN: 2041-0778 Maxwell Scientific Organization, 2009 Submitted Date: October 30, 2009 Accepted Date: November 21, 2009 Published Date:
More informationDIFFERENTIAL RESPONSE OF THE EDAPHIC ECOTYPES IN CYNODON DACTYLON (L)
DIFFERENTIAL RESPONSE OF THE EDAPHIC ECOTYPES IN CYNODON DACTYLON (L) PERS. TO SOIL CALCIUM BY P. S. RAMAKRISHNAN* AND VIJAY K. SINGH Department of Botany, Panjab University, -^, India {Received 24 April
More informationGenetic Divergence Studies for the Quantitative Traits of Paddy under Coastal Saline Ecosystem
J. Indian Soc. Coastal Agric. Res. 34(): 50-54 (016) Genetic Divergence Studies for the Quantitative Traits of Paddy under Coastal Saline Ecosystem T. ANURADHA* Agricultural Research Station, Machilipatnam
More informationSALT TOLERANCE IN THE HALOPHYTE SUAEDA MARITIMA (L.) DUM. THE EFFECT OF SALINITY ON THE CONCENTRATION OF SODIUM IN THE XYLEM
New Phytol. (1987) 105, 359-366 359 SALT TOLERANCE IN THE HALOPHYTE SUAEDA MARITIMA (L.) DUM. THE EFFECT OF SALINITY ON THE CONCENTRATION OF SODIUM IN THE XYLEM BY N. J.W. CLIPSON* AND T. J. FLOWERSf School
More informationIrrigation water salinity limits faba bean (Vicia faba L.) photosynthesis
5 th CASEE Conference Healthy Food Production and Environmental Preservation The Role of Agriculture, Forestry and Applied Biology Irrigation water salinity limits faba bean (Vicia faba L.) photosynthesis
More informationWATER RELATIONS OF WHEAT ALTERNATED BETWEEN TWO ROOT TEMPERATURES
Nezo Phytol. (1979) 82, 89-96 89 WATER RELATIONS OF WHEAT ALTERNATED BETWEEN TWO ROOT TEMPERATURES BY M. B. KIRKHAM Department of Agrofiomy, Oklahoma State University^ StiHwater, Oklahoma 74074, U,S.A.
More informationS. F. AFZALI 1*, M. A. HAJABBASI 1, H. SHARIATMADARI 1, K. RAZMJOO 2, AND A. H. KHOSHGOFTARMANESH 1. Abstract
Pak. J. Bot., 38(5): 1709-1714, 2006. COMPARATIVE ADVERSE EFFECTS OF PEG- OR NaCl- INDUCED OSMOTIC STRESS ON GERMINATION AND EARLY SEEDLING GROWTH OF A POTENTIAL MEDICINAL PLANT MATRICARIA CHAMOMILLA S.
More informationAMMONIUM UPTAKE FROM DILUTE SOLUTIONS BY PINUS RADIATA SEEDLINGS
10 Vol. 9 AMMONIUM UPTAKE FROM DILUTE SOLUTIONS BY PINUS RADIATA SEEDLINGS JAMES W. FLEWELLING School of Forest Resources, University of Georgia, Athens, Georgia, U.S.A. (First received for publication
More informationEFFECT OF SOIL ACIDITY NEUTRALIZATION AND COPPER TOXICITY ON MAIZE PRODUCTIVITY, COPPER UPTAKE AND BIOMASS CATION CONTENT
56 BULG. J. PLANT PHYSIOL., 1996, 22(3 4), 56 65 EFFECT OF SOIL ACIDITY NEUTRALIZATION AND COPPER TOXICITY ON MAIZE PRODUCTIVITY, COPPER UPTAKE AND BIOMASS CATION CONTENT Antoaneta Arsova N. Poushkarov
More informationEFFECTS OF SEED SIZE AND EMERGENCE TIME ON SUBSEQUENT GROWTH OF PERENNIAL RYEGRASS
Phytol (980) 84, 33-38 EFFECTS OF SEED SIZE AND EMERGENCE TIME ON SUBSEQUENT GROWTH OF PERENNIAL RYEGRASS BY ROBERT E. L. NAYLOR School of Agriculture, The University, Aberdeen {Accepted 2 January 979)
More informationTitle Allantoin by Inosine in Nutrient So. Author(s) Toshihiro; Yokoi, Daisuke; Osaki, M
Title Rice Root Growth with Increasing in Allantoin by Inosine in Nutrient So Author(s) Tokuhisa, Dai; Okazaki, Keiki; Shin Toshihiro; Yokoi, Daisuke; Osaki, M Citation The Proceedings of the Internationa
More informationNa+ recirculation in alfalfa and phloem transfer cell structure. N?ziha Boughanmi 1, Philippe Michonneau 2 and Pierrette Fleurat-Lessard 2
Na+ recirculation in alfalfa and phloem transfer cell structure N?ziha Boughanmi 1, Philippe Michonneau 2 and Pierrette Fleurat-Lessard 2 1 D?partement des Sciences de la Vie. Facult? des Sciences de Bizerte.
More informationGFP GAL bp 3964 bp
Supplemental Data. Møller et al. (2009) Shoot Na + exclusion and increased salinity tolerance engineered by cell type-specific alteration of Na + transport in Arabidopsis Supplemental Figure 1. Salt-sensitive
More informationCommon Effects of Abiotic Stress Factors on Plants
Common Effects of Abiotic Stress Factors on Plants Plants are living organisms which lack ability of locomotion. Animals can move easily from one location to other. Immovable property of plants makes it
More informationCOMPARATIVE PERFORMANCE OF BRASSICA NAPUS AND ERUCA SATIVA UNDER WATER DEFICIT CONDITIONS: AN ASSESSMENT OF SELECTION CRITERIA
Journal of Research (Science), Bahauddin Zakariya University, Multan, Pakistan. Vol.15, No.4, December 24, pp. 439-446 ISSN 121-112 COMPARATIVE PERFORMANCE OF BRASSICA NAPUS AND ERUCA SATIVA UNDER WATER
More informationEvaluating SYDlbiotic Potential of Rhizobia
SECTION III Evaluating SYDlbiotic Potential of Rhizobia SIGNIFICANCE OF SYMBIOTIC NITROGEN FIXATION TO AGRICULTURE The value of legumes in improving and sustaining soil fertility was well known to agriculturalists,
More informationSHORT COMMUNICATION PREDICTION OF LEAF AREA IN PHASEOLUS VULGARIS BY NON-DESTRUCTIVE METHOD BULG. J. PLANT PHYSIOL., 2003, 29(1 2),
96 BULG. J. PLANT PHYSIOL., 2003, 29(1 2), 96 100 SHORT COMMUNICATION PREDICTION OF LEAF AREA IN PHASEOLUS VULGARIS BY NON-DESTRUCTIVE METHOD Bhatt M. and Chanda S.V.* Department of Biosciences, Saurashtra
More informationScreening methods for salinity tolerance: a case study with tetraploid wheat
Plant and Soil 253: 201 218, 2003. 2003 Kluwer Academic Publishers. Printed in the Netherlands. 201 Screening methods for salinity tolerance: a case study with tetraploid wheat Rana Munns 1 &RichardA.James
More informationModel of Dry Matter and Plant Nitrogen Partitioning between Leaf and Stem for Coastal Bermudagrass. I. Dependence on Harvest Interval
JOURNAL OF PLANT NUTRITION Vol. 27, No. 9, pp. 1585 1592, 2004 Model of Dry Matter and Plant Nitrogen Partitioning between Leaf and Stem for Coastal Bermudagrass. I. Dependence on Harvest Interval A. R.
More informationTropical Agricultural Research & Extension 16(4): 2014
Tropical Agricultural Research & Extension 16(4): 2014 EFFECTS OF MYCORRHIZAE AS A SUBSTITUTE FOR INORGANIC FERTILIZER ON GROWTH AND YIELD OF TOMATO (LYCOPERSICON ESCULENTUM L.) AND SOY- BEAN (GLYCINE
More informationWhere we re at! The Kikuyu program at the University of Sydney
Where we re at! The Kikuyu program at the University of Sydney Selection 946 tidal mark Black Ada Swamp, Tarthra FACULTY OF AGRICULTURE & ENVIRONMENT Mr Duncan Fraser M.Agr Plant Breeding Institute Meeting;
More informationUnderstanding how vines deal with heat and water deficit
Understanding how vines deal with heat and water deficit Everard Edwards CSIRO AGRICULTURE & FOOD How hot is too hot? Cell death will occur in any vine tissue beyond a threshold (lethal) temperature cell
More informationSymbiotic Fungal Endophytes that Confer Tolerance for Plant Growth in Saline and Dry Soils Zakia Boubakir, Elizabeth Cronin, Susan Kaminskyj
Symbiotic Fungal Endophytes that Confer Tolerance for Plant Growth in Saline and Dry Soils Zakia Boubakir, Elizabeth Cronin, Susan Kaminskyj Department of Biology University of Saskatchewan 1 Outline Background
More informationUC Davis The Proceedings of the International Plant Nutrition Colloquium XVI
UC Davis The Proceedings of the International Plant Nutrition Colloquium XVI Title Varietal differences in salinity tolerance and mineral nutrition in tomatoes (Solanum lycopersicum) Permalink https://escholarship.org/uc/item/65bn6h9
More informationTHE EFFECTS OF FOLIAR APPLICATION OF SALICYLIC ACID ON QUALITATIVE AND QUALITATIVE YIELD OF WHEAT UNDER SALINE CONDITIONS
International Research Journal of Applied and Basic Sciences. Vol., 2 (9), 366-370, 2011 Available online at http://www. irjabs.com 2011 THE EFFECTS OF FOLIAR APPLICATION OF SALICYLIC ACID ON QUALITATIVE
More informationName: B5 PLANT HORMONES. Class: Practice questions. Date: 53 minutes. Time: 53 marks. Marks: Biology Only. Comments: Page 1 of 25
B5 PLANT HORMONES Practice questions Name: Class: Date: Time: 53 minutes Marks: 53 marks Comments: Biology Only Page of 25 Hormones called auxins control plant growth. A student investigated plant growth
More informationWEATHERING ACCORDING TO THE CATIONIC BONDING ENERGIES OF COLLOIDS I ABSTRACT
WEATHERING ACCORDING TO THE CATIONIC BONDING ENERGIES OF COLLOIDS I By E. R. GRAHAM University of Missouri ABSTRACT A study was made of the energy changes of several colloidal systems in relation to weathering.
More informationAPPLICATION OF NEAR INFRARED REFLECTANCE SPECTROSCOPY (NIRS) FOR MACRONUTRIENTS ANALYSIS IN ALFALFA. (Medicago sativa L.) A. Morón and D. Cozzolino.
ID # 04-18 APPLICATION OF NEAR INFRARED REFLECTANCE SPECTROSCOPY (NIRS) FOR MACRONUTRIENTS ANALYSIS IN ALFALFA (Medicago sativa L.) A. Morón and D. Cozzolino. Instituto Nacional de Investigación Agropecuaria.
More informationDeterminations by Atomic Absorption Spectroscopy and Inductively Coupled Plasma-Atomic Emission
0 chapter Sodium and Potassium Determinations by Atomic Absorption Spectroscopy and Inductively Coupled Plasma-Atomic Emission Spectroscopy 67 S. S. Nielsen (ed.), Food Analysis Laboratory Manual Springer
More informationBiology and management of dodder a new threat to the canola industry
Biology and management of dodder a new threat to the canola industry Abul Hashem Department of Agriculture and Food Western Australia Centre for Cropping Systems P.O. Box 483, Northam WA 641, Australia
More informationIn vitro digestibility and neutral detergent fibre and lignin contents of plant parts of nine forage species
Journal of Agricultural Science, Cambridge (1998), 131, 51 58. 1998 Cambridge University Press Printed in the United Kingdom 51 In vitro digestibility and neutral detergent fibre and lignin contents of
More informationGENETIC ANALYSES OF ROOT SYSTEM DEVELOPMENT IN THE TOMATO CROP MODEL
GENETIC ANALYSES OF ROOT SYSTEM DEVELOPMENT IN THE TOMATO CROP MODEL Kelsey Hoth 1 Dr. Maria Ivanchenko 2 Bioresourse Research 1, Department of Botany and Plant Physiology 2, Oregon State University, Corvallis,
More informationSimplified Analysis for Soil Samples with Compact Water Quality Meter <LAQUAtwin>
Feature Article Simplified Analysis for Soil Samples with Compact Water Quality Meter Measurement of exchangeable calcium ion and potassium ion in soil Keiko KUWAMOTO Exchangeable calcium ion
More informationCRITICAL PETIOLE POTASSIUM LEVELS AS RELATED TO PHYSIOLOGICAL RESPONSES OF CHAMBER- GROWN COTTON TO POTASSIUM DEFICIENCY
Summaries of Arkansas Cotton Research 23 CRITICAL PETIOLE POTASSIUM LEVELS AS RELATED TO PHYSIOLOGICAL RESPONSES OF CHAMBER- GROWN COTTON TO POTASSIUM DEFICIENCY D.L. Coker, D.M. Oosterhuis, M. Arevalo,
More informationModel Analysis for Growth Response of Soybean
COMMUNICATIONS IN SOIL SCIENCE AND PLANT ANALYSIS Vol. 34, Nos. 17 & 18, pp. 2619 2632, 2003 Model Analysis for Growth Response of Soybean A. R. Overman * and R. V. Scholtz III Agricultural and Biological
More informationMariana Cruz Campos. School of Plant Biology Faculty of Natural and Agricultural Sciences
Mariana Cruz Campos School of Plant Biology Faculty of Natural and Agricultural Sciences Mariana holds a Bachelor degree with Honours in Biological Sciences from the University of São Paulo, Brazil, where
More informationEVOLUTION IN CLOSELY ADJACENT PLANT POPULATIONS
EVOLUTION IN CLOSELY ADJACENT PLANT POPULATIONS III. AGROSTIS TENUIS ON A SMALL COPPER MINE T. McNEILLY * Deportment of Agricultural Botany, University College of North Wales, Bangor Received 24.iv.6 1.
More informationEFFECT OF DROUGHT STRESS ON PHYSIOLOGICAL GROWTH PARAMETERS OF TOMATO INBRED LINES AT GERMINATION STAGE
EFFECT OF DROUGHT STRESS ON PHYSIOLOGICAL GROWTH PARAMETERS OF TOMATO INBRED LINES AT GERMINATION STAGE Kibreab M. Ghebremariam, MA Liang Yan, Prof. Zhengcai Zhang, MA Qiaoli Wang, MA Northwest A&F University,
More informationROLE OF THE ALLELOPATHY IN MIXED VEGETABLE CROPS IN THE ORGANIC FARMING
Abstract Scientific Papers. Series A. Agronomy, Vol. LVI, 2013 ISSN 2285-5785; ISSN CD-ROM 2285-5793; ISSN Online 2285-5807; ISSN-L 2285-5785 ROLE OF THE ALLELOPATHY IN MIXED VEGETABLE CROPS IN THE ORGANIC
More informationMAIZE-GAMAGRASS INTERSPECIFIC HYBRID, ZEA MAYS
International Journal of Latest Research in Science and Technology Volume 4, Issue 1: Page No.128-133 January-February 2015 http://www.mnkjournals.com/ijlrst.htm ISSN (Online):2278-5299 MAIZE-GAMAGRASS
More informationLEAF APPEARANCE RATE IN Brachiaria decumbens GROWN IN NITROGEN AND POTASSIUM RATES. Abstract
ID # 01-30 LEAF APPEARANCE RATE IN Brachiaria decumbens GROWN IN NITROGEN AND POTASSIUM RATES M.D.C. Ferragine 1, F.A Monteiro 2 and S. C. da Silva 3 1,2 Departamento de Solos e Prod. Vegetal, Universidade
More informationTHE ROLE OF CELL WALL PEROXIDASE IN THE INHIBITION OF LEAF AND FRUIT GROWTH
264 BULG. J. PLANT PHYSIOL., SPECIAL ISSUE 2003, 264 272 THE ROLE OF CELL WALL PEROXIDASE IN THE INHIBITION OF LEAF AND FRUIT GROWTH T. Djaković 1, Z. Jovanović 2 1 Maize Research Institute, Slobodana
More informationGenetic and physiological approach to elucidation of Cd absorption mechanism by rice plants
Genetic and physiological approach to elucidation of Cd absorption mechanism by rice plants Satoru Ishikawa National Institute for Agro-Environmental Sciences, 3-1-3, Kannondai, Tsukuba, Ibaraki, 305-8604,
More informationNumerical Data. Effect of copper on nutrients content (mg g 1 dry wt.) of radish (45th day) N P K Na Ca Mg
Numerical Data Effect of copper on nutrients (mg g 1 dry wt.) of radish (45th day) Copper added in the soil N P K Na Ca Mg (mg kg 1 ) Control 32.66 6.31 43.52 1.70 13.56 3.99 50 37.35(+14.36) 6.97(+10.45)
More informationGeneral Medicine 2016/17
ÚSTAV LÉKAŘSKÉ BIOCHEMIE A LABORATORNÍ DIAGNOSTIKY 1. LF UK Buffers, buffer capacity. Oxidoreduction, electrode processes Practical lesson on medical biochemistry General Medicine Martin Vejražka, Tomáš
More informationInteractive Effects of Salinity and Growth Substances on Grermination of Pea (Pisum sativum,) Seeds
Phyton (Austria) Vol. 25 Faso. 1 87-91 28. 2/1985 Interactive Effects of Salinity and Growth Substances on Grermination of Pea (Pisum sativum,) Seeds Ramesh Chander SETIA and Shelly NARANG *) Received
More informationEffects of pre-sowing seed treatments on seed germination and salinity tolerance of Rice (Oryza sativa L.) seedlings
International Journal of Agronomy and Agricultural Research (IJAAR) ISSN: 2223-7054 (Print) 2225-3610 (Online) http://www.innspub.net Vol. 9, No. 6, p. 112-117, 2016 RESEARCH PAPER OPEN ACCESS Effects
More informationTitle: Salt tolerance in the halophyte Salicornia dolichostachya Moss: growth, morphology and physiology
Title: Salt tolerance in the halophyte Salicornia dolichostachya Moss: growth, morphology and physiology Authors: Diana Katschnig, Rob Broekman, Jelte Rozema PII: S0098-8472(12)00089-5 DOI: doi:10.1016/j.envexpbot.2012.04.002
More informationUse of Western Pozzolan ATS soil amendment for golf course fairways.
Use of Western Pozzolan ATS soil amendment for golf course fairways. Study was undertaken to examine the properties of Western Pozzolan material for use in golf course fairways to improve water relations
More informationKevin Foster. School of Plant Biology Faculty of Natural and Agricultural Sciences
Kevin Foster School of Plant Biology Faculty of Natural and Agricultural Sciences Kevin holds a Bachelor of Science degree from Curtin University and a Diploma in Agricultural Technology. He is currently
More informationVitiNotes. Grapevine nutrition 3: Petiole analysis
VitiNotes www.crcv.com.au 2006 Grapevine nutrition 3: Petiole analysis Current titles in this Grapevine nutrition VitiNote series include: 1. Nitrogen fertilisation 2. Phosphorus fertilisation 3. Petiole
More informationCONTRIBUTION TO THE STUDY OF THE IMPACT OF A FERTILIZER COMMONLY USED IN ALGERIA (NPK) ON GERMINATION AND RESPIRATORY METABOLISM OF TRITICUM DURUM
Proceedings of the 13 th International Conference of Environmental Science and Technology Athens, Greece, 5-7 September 2013 CONTRIBUTION TO THE STUDY OF THE IMPACT OF A FERTILIZER COMMONLY USED IN ALGERIA
More informationPOTASSIUM IN PLANT GROWTH AND YIELD. by Ismail Cakmak Sabanci University Istanbul, Turkey
POTASSIUM IN PLANT GROWTH AND YIELD by Ismail Cakmak Sabanci University Istanbul, Turkey Low K High K High K Low K Low K High K Low K High K Control K Deficiency Cakmak et al., 1994, J. Experimental Bot.
More informationBy the end of this lesson, you should be able to
Allelopathy 1 Allelopathy By the end of this lesson, you should be able to define allelopathy explain the difference between allelopathy and competition identify the key interactions in allelopathy provide
More informationTREATMENT OF LANDFILL LEACHATE USING ZEOLITE
TREATMENT OF LANDFILL LEACHATE USING ZEOLITE Grant Redvers and Renate Timm (City Design Ltd), Garry Peters (Auckland City Council) and Craig Mowatt (NZ Natural Zeolite Ltd) Introduction City Design Ltd,
More informationAvoidance of sodium accumulation by the stomatal guard cells of the halophyte Aster tripolium
Journal of Experimental Botany, Vol. 48, No. 38, pp. 77-711, March 1997 Journal of Experimental Botany Avoidance of sodium accumulation by the stomatal guard cells of the halophyte Aster tripolium L..R.R.
More informationSalt Tolerance of Forage Kochia, Gardner's Saltbush, and Halogeton: Studies in Hydroponic Culture
Utah State University DigitalCommons@USU All Graduate Theses and Dissertations Graduate Studies 2016 Salt Tolerance of Forage Kochia, Gardner's Saltbush, and Halogeton: Studies in Hydroponic Culture Joseph
More informationTREES. Functions, structure, physiology
TREES Functions, structure, physiology Trees in Agroecosystems - 1 Microclimate effects lower soil temperature alter soil moisture reduce temperature fluctuations Maintain or increase soil fertility biological
More informationEFFECTS OF WATER DEFICIT ON PHYSIOLOGY AND MORPHOLOGY OF THREE VARIETIES OF NERICA RAINFED RICE (Oryza sativa L.)
EFFECTS OF WATER DEFICIT ON PHYSIOLOGY AND MORPHOLOGY OF THREE VARIETIES OF NERICA RAINFED RICE (Oryza sativa L.) Sikuku P. A., Netondo G. W., Onyango J. C. and Musyimi D. M. Department of Botany and Horticulture,
More informationResponse of Hybrid Lowland Rice (Oryza sativa L.) Cultivar FARO44 to Salinity in Coastal Plain Sands of the Niger Delta, Nigeria
Journal of Plant Sciences 2015; 3(2): 106-110 Published online April 16, 2015 (http://www.sciencepublishinggroup.com/j/jps) doi: 10.11648/j.jps.20150302.20 ISSN: 2331-0723 (Print); ISSN: 2331-0731 (Online)
More information15 Acids, Bases, and Salts. Lemons and limes are examples of foods that contain acidic solutions.
15 Acids, Bases, and Salts Lemons and limes are examples of foods that contain acidic solutions. Chapter Outline 15.1 Acids and Bases 15.2 Reactions of Acids and Bases 15.3 Salts 15.4 Electrolytes and
More informationFineMapping and Further Characterisation of a chromosomal Region of 3H with QTLs for Salinity Tolerance in Barley (Hordeum vulgare L.
FineMapping and Further Characterisation of a chromosomal Region of 3H with QTLs for Salinity Tolerance in Barley (Hordeum vulgare L.) Han Hoang Nguyen FineMapping and Further Characterisation of a chromosomal
More informationESTIMATION OF LEAF AREA IN WHEAT USING LINEAR MEASUREMENTS
P L A N T B R E E D I N G A N D S E E D S C I E N C E Volume 46 no. 2 2002 S.V. Chanda, Y.D. Singh Department of Biosciences, Saurashtra University,Rajkot 360 005, India ESTIMATION OF LEAF AREA IN WHEAT
More informationDuring photosynthesis, plants convert carbon dioxide and water into glucose (C 6 H 12 O 6 ) according to the reaction:
Example 4.1 Stoichiometry During photosynthesis, plants convert carbon dioxide and water into glucose (C 6 H 12 O 6 ) according to the reaction: Suppose that a particular plant consumes 37.8 g of CO 2
More informationNaCl (aq) + HOH (l) + (aq) + Cl (l) Eg. HCl (aq) + NH 3(aq) In both cases the acid and base react and neutralize each other.
Acid Base Reactions An acid base reaction is called a neutralization reaction. Eg. HCl (aq) + NaOH (aq) NaCl (aq) + HOH (l) In grade 10 you learned that acid base reactions produce salt and water, however,
More informationEelgrass biomass and production
Eelgrass biomass and production Objectives To introduce methods for assessing basic parameters fundamental to many seagrass studies such as shoot size and stand structure expressed as biomass and shoot
More informationC.v. Dr. Mohammed Ali Hussein
C.v. Dr. Mohammed Ali Hussein - Dr Mohammed Ali Hussien Al- Falahe Email: dr.mohammed1953@yahoo.com Tele : 07507718671. - Was born in Baghdad Iraq 1953. - Graduated from Al-Nasar primary school in 1966.
More informationDrought Tolerant Criterion of Wheat Genotypes Using Carbon Isotopes Discrimination Technique
Journal of Earth Science and Engineering 5 (2015) 256-261 doi: 10.17265/2159-581X/2015. 01. 005 D DAVID PUBLISHING Drought Tolerant Criterion of Wheat Genotypes Using Carbon Isotopes Discrimination Technique
More information25-3 Plant Adaptations Slide 2 of 29
2 of 29 Aquatic Plants How are plants adapted to different environments? 3 of 29 Aquatic Plants Aquatic Plants To take in sufficient oxygen, many aquatic plants have tissues with large air-filled spaces
More informationAdel Sepehry, Davoud Akhzari, Mohammad Pessarakli and Hossein Barani. School of Plant Sciences, University of Arizona, Tucson, AZ 85721, USA
American-Eurasian J. Agric. & Environ. Sci., 12 (10): 1411-1419, 2012 ISSN 1818-6769 IDOSI Publications, 2012 DOI: 10.5829/idosi.aejaes.2012.12.10.1765 Effect of Salinity, Aridity and Grazing Stress on
More informationTropical Grasslands (1999) Volume 33,
Tropical Grasslands (1999) Volume 33, 122 126 122 Effects of leachates from swards of Bothriochloa pertusa and Urochloa mosambicensis on the growth of four test species, B. pertusa, U. mosambicensis, Stylosanthes
More informationPHARMACY CALCULATIONS
THE OBJECTIVES INCLUDE THE FOLLOWING: PHARMACY CALCULATIONS Discuss importance of calculations in a pharmacy practice. Outline steps to avoid calculation errors Explain importance of reading the prescription
More informationEFFECTS OF DIFFERENT DOSES OF GLYCINE BETAINE AND TIME OF SPRAY APPLICATION ON YIELD OF COTTON (GOSSYPIUM HIRSUTUM L.)
Journal of Research (Science), Bahauddin Zakariya University, Multan, Pakistan. Vol.17, No.4, October 2006, pp. 241-245 ISSN 1021-1012 EFFECTS OF DIFFERENT DOSES OF GLYCINE BETAINE AND TIME OF SPRAY APPLICATION
More informationMEASUREMENT OF THE TRANS-ROOT ELECTRICAL POTENTIAL OF PLANTS GROWN IN SOIL
New Phytol. (1972) 71, 111-117. MEASUREMENT OF THE TRANS-ROOT ELECTRICAL POTENTIAL OF PLANTS GROWN IN SOIL BY A. Q. ANSARI* AND D. J. F. BOWLING Department of Botany, University of Aberdeen {Received %
More informationModel of Dry Matter and Plant Nitrogen Partitioning between Leaf and Stem for Coastal Bermudagrass. II. Dependence on Growth Interval
JOURNAL OF PLANT NUTRITION Vol. 27, No. 9, pp. 1593 1600, 2004 Model of Dry Matter and Plant Nitrogen Partitioning between Leaf and Stem for Coastal Bermudagrass. II. Dependence on Growth Interval A. R.
More informationChapter 4 Solution Stoichiometry. Dr. Sapna Gupta
Chapter 4 Solution Stoichiometry Dr. Sapna Gupta Concentrations of Solutions A solution is solute dissolved in a solvent. To quantify and know exactly how much of a solute is present in a certain amount
More informationTHE EFFECT OF CATIONS ON THE ABSORPTION OF PHOSPHATE BY BEECH MYCORRHIZAL ROOTS
THE EFFECT OF CATIONS ON THE ABSORPTION OF PHOSPHATE BY BEECH MYCORRHIZAL ROOTS BY D. H. JENNINGS Botany Department, Leeds University {Received 30 April 1964) SUMMARY Pretreatment of beech mycorrhizal
More informationFlushing Out the Moles in Lab: The Reaction of Calcium Chloride with Carbonate Salts
Flushing Out the Moles in Lab: The Reaction of Calcium Chloride with Carbonate Salts Pre-lab Assignment: Reading: 1. Chapter sections 3.3, 3.4, 3.7 and 4.2 in your course text. 2. This lab handout. Questions:
More informationInstitute of Natural Resources and Environmental Sciences, Pakistan Agricultural Research Council, Islamabad-45500, Pakistan 2
GEN. APPL. PLANT PHYSIOLOGY, 2005, 31(1-2), 89-104 89 EFFECT OF PRE-GERMINATION AND POST-GERMINA- TION TREATMENT WITH GROWTH HORMONES (KINETINE AND ABSCISIC ACID) ON ION CONCENTRATION AND BIOCHEMICAL CONTENTS
More informationCHAPTER 4 TYPES OF CHEMICAL REACTIONS & SOLUTION STOICHIOMETRY
Advanced Chemistry Name Hour Advanced Chemistry Approximate Timeline Students are expected to keep up with class work when absent. CHAPTER 4 TYPES OF CHEMICAL REACTIONS & SOLUTION STOICHIOMETRY Day Plans
More informationSELECTING NEW Brachiaria FOR BRAZILIAN PASTURES. 2 CNPq fellow. Abstract
ID # 13 14 SELECTING NEW Brachiaria FOR BRAZILIAN PASTURES C.B. do Valle 1,2, V.P.B. Euclides 1,2, M.C.M. Macedo 1,2, J R. Valério 1,2 and S. Calixto 1 1 Embrapa Gado de Corte, Caixa Postal 154, 79002-970
More informationIdentification of Rhizosphere Abiotic Stress Tolerance and Related Root Traits in Soybean [ Glycine max ( L. ) Merr. ]
31 9 2005 9 1132 1137 ACTA AGRONOMICA SINICA Vol131, No19 pp1 1132-1137 Sept1, 2005 1,2 1, 3 1 Ξ ( 1,, 210095 2, 056038) : 301 62,,, P 1 4 6 2, 7 3 3, : : S565 Identification of Rhizosphere Abiotic Stress
More informationJournal of Water and Soil Vol. 25, No. 6, Jan-Feb 2012, p
Journal of Water and Soil Vol. 25, No. 6, Jan-Feb 2012, p. 1310-1320 ( ) 1310-1320. 1390-6 25 4 3 2 1 - - - 89/9/9: 90/5/1:.. - ( ) 900 100.. ( ) ( ) 5. ( ) 150... 200 100. 700. 700 500 : 190 118 105 75
More informationEffect of PGPR on growth promotion of rice (Oryza sativa L.) under salt stress
Available online at www.pelagiaresearchlibrary.com Asian Journal of Plant Science and Research, 2014, 4(5):62-67 ISSN : 2249-7412 CODEN (USA): AJPSKY Effect of PGPR on growth promotion of rice (Oryza sativa
More informationList of Equipment, Tools, Supplies, and Facilities:
Unit D: ph of Soil Lesson 2: Identifying ph Connection With Plant Growth Student Learning Objectives: Instruction in this lesson should result in the students achieving the following objectives: 1. Explain
More informationEffect of diazotrophs on the mineralization of organic nitrogen in the rhizosphere soils of rice (Oryza sativa)
Journal of Crop and Weed 3(1) : 47-51 (7) Effect of diazotrophs on the mineralization of organic nitrogen in the rhizosphere soils of rice (Oryza sativa) A. C. DAS AND D. SAHA Department of Agricultural
More informationAcid-Base Equilibria and Solubility Equilibria
Acid-Base Equilibria and Solubility Equilibria Acid-Base Equilibria and Solubility Equilibria Homogeneous versus Heterogeneous Solution Equilibria (17.1) Buffer Solutions (17.2) A Closer Look at Acid-Base
More informationModel Analysis for Partitioning of Dry Matter and Plant Nitrogen for Stem and Leaf in Alfalfa
Communications in Soil Science and Plant Analysis, 36: 1163 1175, 2005 Copyright # Taylor & Francis, Inc. ISSN 0010-3624 print/1532-2416 online DOI: 10.1081/CSS-200056889 Model Analysis for Partitioning
More informationDistribution and mobility of aluminium in an Al-accumulating plant, Fagopyrum esculentum Moench.
Journal of Experimental Botany, Vol. 52, No. 361, pp. 1683±1687, August 2001 Distribution and mobility of aluminium in an Al-accumulating plant, Fagopyrum esculentum Moench. Renfang Shen and Jian Feng
More informationINFLUENCE OF FOLIAR APPLICATION OF POTASSIUM ON WHEAT (TRITICUM AESTIVUM L) UNDER SALINE CONDITIONS
Sci., Tech. and Dev., 32 (4): 285-289, 2013 INFLUENCE OF FOLIAR APPLICATION OF POTASSIUM ON WHEAT (TRITICUM AESTIVUM L) UNDER SALINE CONDITIONS AMEER KHAN* AND MUNAWER AZIZ Department of Biological Sciences,
More informationGENETIC VARIABILITY WITHIN TWO ADAPTED POPULATIONS OF TALL WHEATGRASS (THYNOPYRUM PONTICUM) IN ARGENTINA.
ID # 12-03 GENETIC VARIABILITY WITHIN TWO ADAPTED POPULATIONS OF TALL WHEATGRASS (THYNOPYRUM PONTICUM) IN ARGENTINA A. Andrés 1 and R. Guillen 2 1 INTA EEA Pergamino. CC 31 (2700) Pergamino, Buenos Aires.,
More informatione Crop Management in Sugarcane... easi g Cane, Sugar and Jaggery Yields Souvenir Proceedings
T ational Seminar on e Crop Management in Sugarcane easi g Cane, Sugar and Jaggery Yields Souvenir cum Proceedings Venue Andhra University Campus, Visakhapatnam 5th & 6th December, 2014, ', Organised by
More informationSTUDIES IN THE PHYSIOLOGY OF LICHENS
STUDIES IN THE PHYSIOLOGY OF LICHENS V. TRANSLOCATION FROM THE ALGAL LAYER TO THE MEDULLA IN PELTIGERA POLYDACTYLA BY D. C. SMITH AND E. A. DREW Department of Agriculture, University of Oxford {Received
More informationSalinity effects on the stomatal behaviour of grapevine
New Phytol. (1990), 116, 499-503 Salinity effects on the stomatal behaviour of grapevine BY W. J. S. DOWNTON, B. R. LOVEYS AND W. J. R. GRANT CSIRO Division of Horticulture, GPO Box 350, Adelaide, 5001,
More informationCurrent State of Extraction Don t Be Deceived! Sharon F. Webb, Ph.D. Director of Quality Program
Current State of Extraction Don t Be Deceived! Sharon F. Webb, Ph.D. Director of Quality Program Overview Factors Purpose of Dissolution Quality Objectives of Program Effectiveness of Dissolution Technique
More informationRecent advances in genetics of salt tolerance in tomato
Plant Cell, Tissue and Organ Culture 76: 101 119, 2004. 2004 Kluwer Academic Publishers. Printed in the Netherlands. 101 Review of Plant Biotechnology and Applied Genetics Recent advances in genetics of
More informationAP Chapter 4: Reactions in Aqueous Solution Name
AP Chapter 4: Reactions in Aqueous Solution Name WarmUps (Show your work for credit) Date 1. Date 2. Date 3. Date 4. Date 5. Date 6. Date 7. Date 8. AP Chapter 4: Reactions in Aqueous Solution 2 WarmUps
More informationPlant Growth & Development. Growth Processes Photosynthesis. Plant Growth & Development
Plant Growth & Development Growth Processes Growth Requirements Types of Growth & Development Factors Growth Processes Photosynthesis Creating carbohydrates (stored energy) from CO 2 + water + sunlight
More informationThe Chemistry of Salinity in Soils. Naomi Waissman Assadian Texas A&M Research and Extension Center at El Paso September 4, 2002
The Chemistry of Salinity in Soils Naomi Waissman Assadian Texas A&M Research and Extension Center at El Paso September 4, 2002 Introduction Salinity concentrations in soils have profound effects on soil
More information