VERTEBRATA PALASIATICA
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1 VERTEBRATA PALASIATICA pp figs ) ( ) :,,, : Q : A : (2004) , 1943 Granger Gregory ( ), ,,, 6 : ( Dianotitan) ( Arctotitan) ( Pygmaetitan) ( Acrotitan) ( Nanotitan) ( Qufutitan) (, 1959,1974 ;,1978,1982 ;,1982 ;,1983 ;Qi and Beard,1996 ;, 1997), 2 : ( ) ( Protitan cf. P. ro2 bustus) ( ) ( Protitan major sp. nov. ) Perissodactyla Owen, 1848 Brontotherioidea Marsh, 1873 Brontotheriidae Marsh,1873 Epimamteoceratinae Granger et Gregory, 1943 Protitan Granger et Gregory, 1943 ( ) Protitan major sp. nov. ( 1) Brontotheriidae gen. et sp. nov., 1978, p , 3 P4 M3 (V 13802) 1) ( : ) :
2 4 : 335 P2 (V 13803) ; DP3 DP4 (V 13804), P4, (M1 M mm) P2,,, DP3, DP4 DP3,,,, W 1 ( ), Fig. 1 Upper cheek teeth of Protitan major sp. nov.,crown view A. right P2(V 13803) ;B. left DP3 DP4 (V 13804) ; 1 2 C. right P4 M3,holotype (V 13802) ; scale bar = 40 mm P4,,,,,, 1 ( ) P2 ( V 13803) DP3 DP4 ( V 13804) Table 1 Measurements of P2 ( V 13803), DP3 and DP4 ( V 13804) of Protitan major sp. nov. (mm) (L) (W) P2 DP3 DP4 DP3 DP approximate value. ( 2),, M1, M2 M2 W
3 336 42,, M3, M2,, 2 ( ) ( V 13802) Table 2 Measurements and comparison of the upper cheek teeth ( V 13802) of Protitan major sp. nov. (mm) P4 M1 M2 M3 M1 M3 (L) P. major sp. nov. (V 13802) P. minor (Type, Am. Mus, no ) P. grangeri (Type, Am Mus. no ) P. bellus (Type, Am. Mus no ) : 1), ; 2) Granger and Gregory (1943), (Metatelmatheriinae) ( Embolotheriinae),,,, ( Granger and Gregory,1943 ;,1997 ;,1999) ( Epimanteoceras) ( E. formosus), P4, (Am Mus No ) M1 M3 157 mm ( Gnathotitan) ( G. berkeyi), (Am Mus No ) M1 M3 235mm M3,112 mm, 71 mm, 63 %, 93 % ( Dolichorhinoides) ( D. angustidens), (Am Mus No ) M1 M3 173mm,,M3 68 % ( Microtitan) ( M. mongoliensis), M1 M3 (Am Mus No ) 88mm, ( Pachytitan) ( P. ajax), (Am Mus No ) M1 M3 190mm, ( Dianotitan) ( D. lunanensis), ( Rhinotitan),, P4 3 ( R. kaiseni) (Am Mus No ) M1 M3 156mm P4 (146 %) (105 %) ( R. mongoliensis) M1 M3 (Neotype, Am Mus No ) 180mm ( R. quadri2
4 4 : 337 dens),p3 P4, P4 ( Metatitan) 3 ( M. primus) ( M. progressus) ( M. relictus),, (cement) ( Parabrontops) ( P. gobiensis), P3 P4 9, ( Protitan), P4, 6 ( P. grangeri) ( P. minor) ( P. bellus) ( P. robustus) ( P. obliquidens) ( P.? cingulatus),, m1 m3 203mm, (M1 M3 = mm) 3,, 2,,, P2,,, ( ) Protitan cf. P. robustus Granger et Gregory, 1943 ( 2) Protitan cf. P. robustus, 1978, P27 28, 3 2 ( ) (B) (A) (V 13805),, = 40mm Fig. 2 The right (A) and left (B) lower jaws (V 13805) of Protitan cf. P. robustus, crown view,scale bar = 40 mm
5 338 42, p3 m3 p2 ; p4 m3 (V 13805), p4 m3,m1 125 mm p2,, p2 40 mm p1 p2 p3, V p4,,, m3 3 3 ( ) (V 13805) Table 3 Measurements of the left lower teeth ( V 13805) of Protitan cf. P. robustus (mm) c1 p2 p3 p4 m1 m2 m3 m1 m3 (L) (W) approximate value. ( ),, Granger Gregory (1943) (Am, Mus. no ) m1 43 mm,m2 59 mm, 33 mm ; m1 m3 203 mm ( 3), ( ),, BRONTOTHERES ( PERISSODACTYLA, MAMMAL IA) FROM THE MIDDLE EOCENE OF L UNAN, YUNNAN HUANG Xue2Shi ZHENGJia2Jian ( Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing ) Key words Lunan, Yunnan, Middle Eocene, Lumeiyi Formation, Brontother ioidae Summary The brontotheres described in the present paper were collected from the Middle Eocene Lumeiyi Formation of Lunan Basin, Yunnan by a field team of IVPP in 1970s. They include two species - Protitan major sp. nov. (V ) and Protitan cf. P. robustus which represented by a left lower jaw with p3 m3 as well as broken c1 and p2, a right lower jaw with p3 m3 of same indi2 vidual (V 13805). Brontotheriidae Marsh, 1873 Protitan Granger et Gregory, 1943 Protitan major sp. nov. (Fig. 1) Brontotheriidae gen. et sp. nov. Zheng et al., 1978, P , table 3
6 4 : 339 Type A fragmentary right upper jaw with P4 M3 (V 13802). Referred specimen An isolated right P2 (V 13803) and broken left DP3 DP4 (V 13804). Locality and horizon Nearby Lumeiyi Village. Lunan County, Yunnan Province ;Lower part of Middle Eocene Lumeiyi Formation. Diagnosis A species of Protitan larger than other Known species of the genus, length of M1 M3 being mm. Etymology The species name shows it is the largest species of the genus so far Known. Remarks The described upper molars, especially the last two ones, are longer than wide, the size of which is larger than those of Metatelmatheriinae and smaller than Embolotheriinae in general, indicating that the material may belong to Epimanteoceratinae ( Granger and Gregory, 1943). There are at least ten genera within the last subfamily. Of them the Lumeiyi upper cheek teeth are close to those of Protitan mainly in having no hypocone (tetartocone) or hypocone ridge on P4. The size, however, is larger than any species else. So the Lumeiyi specimen should represent a new species2 Protitan major, the largest species of the genus. References Chow M C ( ), Chang Y P ( ), Ting S Y ( ), Some Early Tertiary Perissodactyla fromlunan Basin, E. Yunnan. Vert PalAsiat ( ), 12(4) : (in Chinese with English summary) Chow M C ( ), Hu C C ( ), A new species of Parabrontops from the Oligocene of Lunan, Yunnan. Acta Palaeont Sin ( ), 7(2) : (in Chinese with English summary) Granger W, Gregory W K, A revision of the Mongolian titanotheres. Bull Am Mus Nat Hist, 80(10) : Miao D S ( ), Early Tertiary fossil mammals from the Shinao basin, Panxian County, Guizhou Province. Acta Palaeont Sin ( ), 21(5) : (in Chinese with English abstract) Qi T, Beard K C, Nanotitan shanghuangensis, gen. et sp. nov. : The smallest known brontothere (Mammalia : Perissodacty2 la). J Vert Paleont, 16 (3) : Wang B Y ( ), Perissodactyla from the Late Eocene of Lantian, Shensi. Prof Pap Stratigr Palaeont ( ), (7) : (in Chinese with English summary) Wang B Y ( ),1982. Osteology and phylogenetic relationship of Rhinotitan mongoliensis. Mem Inst Vert Paleont Paleoan2 throp, Acad Sin, (16) : 1 75 (in Chinese with English abstract) Wang Y ( ), Guo J W ( ), Wang J W ( ), A review of Chinese Brontotheres. In :Wang YQ, Deng T eds. Proc Seventh Ann Meeting Chinese Soc Vert Paleont ( ). Beijing : China Ocean Press (in Chinese with English summary) Wang Y ( ),Wang J W ( ), A new brontothere from late Middle Eocene of Qufu, Shandong. Vert PalAsiat ( ), 35(1) : (in Chinese with English summary) Ye J ( ), Mammalian fauna from the Late Eocene of Ulan Shireh area, Inner Mongolia. Vert PalAsiat ( ), 21 (2) : (in Chinese with English summary) Zheng J J ( ), Tang YJ ( ), Zhai R J ( ) et al., Early Tertiary strata of Lunan Basin, Yunnan. Prof Pap Stratigr Palaeont ( ), (7) : (in Chinese)
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