Artiodactyla Owen,1848 Anthracotheriidae Gill,1872. rma nicus Pilgrim et Cotter,1916 , C P1 M3, I2 C P3 M3 ( IVPP V 12718) 4 3 P4,

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1 V ERTEBRATA PALASIATICA pp pl. I 1) 1,2 3 ( ) ( ) ( ) ( A nthracokeryx birmanicus),,,,,-, Q ,,, 7, 1990,, Artiodactyla Owen,1848 Anthracotheriidae Gill,1872 Ant hracokeryx Pilgrim et Cotter,1916 Ant hracokeryx bi rma nicus Pilgrim et Cotter,1916 ( I,12), C P1M3,I2C P3M3 ( IVPP V 12718),, 4 3 P4,,,,,, CM3, I2C P3M3 (1) 1) ( :013116) :

2 292 39,I2 I3,5mm ; I3, 22mm 1 Anthracokeryx birmanicus Table 1 Measurements of Anthracokeryx birmanicus and comparison with other specimens (mm) IVPP V IVPP V 4948 GSI B621 AMNH I I P P P P M M M P1P4 51 M1M3 51 (diastema) I2I I3C CP1 5 P1P P2P : IVPP, Insititute of Vertebrate Paleontology and Paleoanthropolopy,Chinese Academy of Seiences,Beijing,China ; GSI, Geological Survey of India,Calcutta,India ;AMNH,American Museum of Natural History,New York,USA. 2 Anthracokeryx birmanicus Table 2 Measurements of canine of Anthracokeryx birmanicus (mm) (canine) (height of crown) (length of root) left right ,,, (2),,,, P1P3 P1, P2,, P2 P1 P3,P3 P3,,,, P4,,,, P1P4,M1 M3,

3 4 : 293,,,,M2 M1 M3 ;,,M3 M2 M1,,,, A nthracokeryx A nthracokeryx,,a nthracokeryx kw angsiensis ;, A nthracokeryx cf. A. bambusae ; A nthracokeryx gungkangensis ;,M1 M3, A nthracokeryx bi rm anicus A nthracokeryx bi rm anicus Pilgrim Cotter (1916), A nthracokeryx (1957) (1977),,A nthracokeryx, P4 A nthracokeryx bi rm anicus P4,,,,,, A. kw angisensis (,1977) A. gungkangensis P4, (,1977) A. dawsoni P4, (,1985) A. bi rm anicus,,,,,,,, (Aitchison,1946 ;,1992), ( ) (,1992), A. bi rm anicus,,,,,,, ( 1986) ( L ist riodon ), L ist riodon ;,,,,,,,, A. brimanicus,,

4 A nthracokeryx bri m anicus (,1979 ;,1989) , 9 13, (,1989), A. bi rm anicus, A. morit urus, A nthracothem a rubricae, Indomeryx cotteri, Metatel m atherium? brow ni Deperetella (,1984) A nthracokeryx A nthracothem a (,1977) Ducrocq (1999) Krabi A nthracothem a A nthracokeryx, Krabi (,1984 ;,1989) (1995),, 1500m, ; 500m,, ( Tsubamoto et al.,2000), Tabyin, N um m ulites acut us, Yaw, N um m ulites yawensis, Discocyclina sella, O perculina cf. O. canalif era V elates perversus, NP216 NP220 ( Takai et al,2000 ;Cavelier and Pomerol,1986),,, Montpellier Ducrocq,

5 4 : 295 NEW MATERIAL OF AN THRACO KER YX B RIMANICUS FROM GUANGXI,CHINA L I Qian 1,2 CHEN Geng2Jiao 3 (1 Insitit ute of Vertebrate Paleontology and Paleoanthropolopy, Chi nese Academy of Sciences Beijing ) (2 L aboratory of Palaeobiology & St ratigrphy, Chi nese Academy of Sciences Nanjing ) (3 Guangxi Nat ural History M useum Nanning ) Key words Quelin, Guangxi, Middle2Late Eocene, Nadu Formation, A nthraco2 keryx bri m anicus Summary A new specimen of A nt hracokery x bri m anicus wit h a complete upper dentition, collected from the Nadu Formation of Quelin, Guangxi in 1990,is described in this paper. Left C,P1M3,right I2C,P3M3 are preserved on the upper jaw. Examination of the new specimen resulted in the emend of the species diagnosis : the interval beween the two incisors is relatively large. The upper canine is large (probably only in adult male),curved outwards and backwards. It extends far back to t he posterior end of maxilla, and has no enclosed root,which suggests the canine is ever growth. The dentition formula of the upper jaw is 4 3. The outer cusp of P4 is crescentic and the inner cusp is conical. The molar has a well marked parastyle,a moderate mesostyle and a weak metastyle. Cingulum is moderate. The shape of the canine of the A nthracokeryx bri m anicus,similar to that of both the male muntjac and the male musk,indicates that the canine was used as a weapon to fight for mate and territory. The finding of upper canine implies t hat A nt hracokery x bri m anicus is sexual dimorp hic. Bicuspid P4 of A nt hracokery x bi rm anicus (crescentic outer cusp and conical inner cusp) differs f rom t hat of A. kw angsiensis, A. gungkangensis and A. dawsoni. The outer cusp of P4 of A. kw angsiensis is conical and the inner cusp is flat2conical ;the anterior cingulum and t he posterior cingulum are well2developed, but absent at t he anterior part of t he outer cusp and the external of the inner cusp. In A. gungkangensis, the outer cusp is conical and the inner cusp is crescentic ;cingulum is well developed. The outer cusp and the inner cups of A. dawsoni are all conical. The age of t he Nadu Formation was considered to be Late Eocene. Comparison wit h Pondaung fauna of Burma suggests that the Nadu Fauna might partially be Middle Eocene in age. Aitchison J,1946. References Hinged teeth in mammals : A study of the tusks of muntjacs ( M untiacus ) and Chinese water deer ( Hydropotes i nermis). Zool Soc London Proc,116 (5) : Cavelier C,Pomerol C,1986. Stratigraphy of the Paleogene. Bull Soc Geol Fr,Ser 8,2 (2) : Cheng G F( ),1986. A new species of L ist riodon meuer (Suidae,Artiodactyla,Mammalia) from Xinan,Henan. Vert PalAsiat ( ),24 (4) : (in Chinese wish English summary) Chow M C( ),1957. On some Eocene and Oligocene mammals from Kwangsi and Yunnan. Vert PalAsiat ( ),3 (1) : Colbert E H,1938. Fossil mammals from Burma in the American Museum of Natural History. Bull Am Mus Nat Hist,74 (6) :259392

6 Ducrocq S,1999. The Late Eocene Anthracotheriidae ( Mammalia,Artiodactyla) from Thailand. Palaeontographica Abt A Bd,252 :93140 Li Y T( ),Li J Y ( ),Sun X Y( ) et al.,1984. Tertiary of China. Beijing : Geological Publishing House (in Chinese) Pilgrim G E,Cotter G de P,1916. Some newly discovered Eocene mammals from Burma. Rec Geol Surv India. 47 :4277 Qui Z D ( ),1977. A new species of A nthrcokeryx from Guangxi. Vert PalAsiat ( ),15 (1) :5459 (in Chinese) Sheng H L ( ),1992. The Deer in China. Shanghai : East China Normal University Press (in Chinese) Takai M,Shigehara N, Tsubamoto T et al.,2000. The latest Middle Eocene Primate fauna in Pondaung area, Myanmar. Asian Paleoprimatology,1 :728 Tang Y J ( ),Qiu Z D ( ),1979. Analyse and discussion of vertebrate fossils in Baise of Guangxi. Red Beds of Mesozoic2Cenozoic Era in South China. Beijing :Science Press (in Chinese) Tsubamoto T,Egi N, Takai M et al., A preliminary report on the Eocene mammals of the Pondaung fauna,myanmar. Asian Paleoprimatology,1 :29101 Tong Y S( ),1989. A review of Middle and Late Eocene mammalian faunas from China. Acta Palaeontol Sin ( ),28 (5) : (in Chinese with English summary) Tong Y S( ),Zheng S H( ),Qiu Z D( ),1995. Cenozoic mammal ages of China. Vert PalAsiat ( ),33 (4) : (in Chinese with English summary) Wang J W ( ),1985. A new species of A nthracokeryx from Shanxi. Vert PalAsiat ( ),23 (1) :52 59 (in Chinese with English summary) I ( Explanations of Plate I) A nthracokeryx bi rmanicus Pilgrim et Cotter,1916,V 12718, 4/ 5 1. (ventral view) ;2. (upper dentitions), (lateral view)

Perissodactyla Owen, 1848

Perissodactyla Owen, 1848 41 4 2003 10 VERTEBRATA PALASIATICA pp. 332 336 figs. 1 2 1), Q915. 876 ( 710062) 2001 8,,,,, 2, (,1959 ;,1962 ;,1976 ;,1998), 2 1, 4 11 23, : Canis variabilis Hyaenidae Homotherium sp. Megantereon sp.