7 fig'., 4 tab., 3 pis. Leiden, September molluscan fauna from LANGA, DENMARK. and. C. Beyer NORWAY STAVANGER,

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1 Contr. Contr. Tert. Quatern. Geol. 27(23) fig'., 4 tab., 3 pis. Leiden, September 1990 A Late Oligocene (Chattian B) molluscan fauna from the coastal cliff at Mogenstrup, North of Skive, Jutland, Denmark K.I. Schnetler LANGA, DENMARK and C. Beyer STAVANGER, NORWAY Schnetler, K.I., & C. Beyer. A Late Oligocene (Chattian B) molluscan fauna from the coastal cliff at Mogenstrup, North of Skive, Jutland, Denmark. Tert. Quatern. Geol., 27(23): 3982, 7 figs, 3 tabs, 3 pis. Leiden, September A rich Late very Oligocene molluscan fauna from a coastal cliff at North of Mogenstrup, Skive, Jutland, Denmark, was studied. A of the Late summary Palaeogene sedimentary sequence in NW Jutland is given, the locality and the section exposed are described. Lithostratigraphical, magnetostratigraphical and biostratigraphical correlations are suggested. A list of the molluscan species is given, comparisons with other Danish and German Late Oligocene localities are made, and palaeoecological interpretations are suggested. In the systematic part the subgenus Andersondrillia subgen. nov. is introduced within the genus Microdrillia. Several taxa are discussed and the following new species and subspecies are described: Limopsis (Pectunculina) lamellata chattica subsp. nov., Limopsis (Pectunculina) vonderhochti sp. nov., Collonia (Collonia) troelsi sp. nov., Lepetella helgae sp. nov., Lepetellajytteae sp. nov., Tubiola subangulata sp. nov., Cerithiopsis (s. lat.) antonjansei sp. nov., Laiocochlis (Laiocochlis) supraoligocaenica sp. nov., Triforis (Trituba) sorgenfreii sp. nov., Cirsotrema (Opaliopsis) subglabrum sp. nov., Searlesia ravni sp. nov., Angistoma brueckneri danica subsp. nov., Clavatula mogenstrupensis sp. nov., Microdrillia (Microdrillia) ingerae sp. nov., Microdrillia (Andersondrillia) brejningensis sp. nov., Pleurotomella (Pleurotomella) rasmusseni sp. nov., Rimosodaphnella lappanni sp. nov. and? Actaeopyramis (s. lat.) pseudopunctata sp. nov. Keywords Mollusca, Late Oligocene, Chattian B, new taxa, lithostratigraphy, magnetostratigraphy, biostratigraphy, palaeoecology, Denmark. K.I. Schnetler, Fuglebakken 14, Stevnstrup, DK8870 Langa, Denmark; C. Beyer, CBMagneto, P.O. Box 7015, Jorenholmen, N4004 Stavanger, Norway. Contents Dansk sammendrag p. 40 Material p. 45 Introduction p. 41 List of molluscan species p. 46 Late Oligocene sedimentary sequence Systematic part p. 50 in NW Jutland p. 41 Comparison with other Late Oligocene mol Lithostratigraphical correlation p. 44 luscan faunas p. 71 Magnetostratigraphical correlation p. 44 Palaeoecological interpretations p. 73 Mogenstrup section p. 44 Acknowledgements p. 75 Biostratigraphical correlation p. 45 References p This study was supported by the Danish National Science Research Council through grant

2 40 Fig. 1 Location of the Mogenstrup section. Part of the 1:25,000 mapsheet 12/6 III SV Hvalpsund (C). Kortog Matrikelstyrelsen (A8490). (Author: Schnetler). Dansk sammendrag En 0vre Oligocaen molluskfauna fra Mogenstrup, I hyppige. det foreliggende arbejde beskrives lokaliteten, og der gives en oversigt over den 0vre Oligocaene lagserie i NV Jylland. Litho, magneto og biostratigrafiske korrelationer foreslas, og en tolkning af Pectiniderne aflejringsmiljoet gives. indicerer en Chattian B alder. Faunalisten indeholder 197 arter. I den systematiske del diskuteres en raekke arter, og pa tre plancher afbildes en raekke nord for Skive, Jylland, Danmark. Fra en kystklint nord for Skive er en meget rig 0vre Oligocaen molluskfauna indsamlet. Denne fauna er karakteristiske arter. Inden for slaegten Microdrillia bemaerkelsesvaerdig ved at indeholde en lang raekke arter, der aldrig tidligere er fundet i danske aflejringer fra denne periode, ligesom en del arter, opstilles den nye underslsegt Andersondrillia. Folgende nye arter og underarter introduceres: Limopsis (Pectunculina) lamellata chattica subsp. no v., der er yderst sjaeldne pa andre lokaliteter, her er Limopsis (Pectunculina) vonderhochti sp. nov., Collonia (Collonia) troelsi sp. nov., Lepetella helgae sp. nov., Lepetella jytteae sp. nov., Tubiola subangulata sp. nov., Cerithiopsis (s. lat.) antonjansei sp. nov., Laiocochlis (Laiocochlis) supraoligocaenica sp. nov., Triforis (Trituba) sorgenfreii sp. nov., Cirsotrema (Opaliopsis)

3 Seven Wellsorted, Laminated 41 subglabra sp. nov., Searlesia ravni sp. nov., Angistoma does show affinities with unpublished assemblages from the Vejle Fjord area, especially with regard brueckneri danica subsp. nov., Clavatula mogenstrupensis sp. nov., Microdrillia (Microdrillia) ingerae sp. nov., Microdrillia (Andersondrillia) brejningensis sp. to a number of characteristic species. assemblage also shows affinities with the very rich fauna nov., Pleurotomella (Pleurotomella) rasmusseni sp. from Glimmerode, F.R.G. (R. Janssen, 1978a, b; 1979a, b). nov., Rimosodaphnella lappanni sp. nov. og? Actaeopyramis (s. lat.) pseudopunctata sp. nov. Faunasammensaetningen indicerer aflejring pa lavere vand end de ovrige kendte danske 0vre Oligocaene faunaer. Faunaen viser, isaer for en raekke THE LATE OLIGOCENE SEDIMENTARY SEQUENCE IN NW JUTLAND (AUTHOR: BEYER) Late Oligocene sedimentary sequence in NW Jutland consists of the finegrained sjaeldnere arters vedkommende, affinitet til faunaer fra Vejle Fjord omradet og til faunaen fra Branden Formation underlying a coarser grained sequence, which outcrops at several localities in the area. se coarser sediments (Fig. 2a) comprise an Glimmerode, Vesttyskland. Ud fra en palasookologisk tolkning foreslas et sublittoralt aflejringsmiljo. up to c. 120 m thick sequence of shallowmarine Introduction (Authors: Schnetler and Beyer) deposits, overlain by Miocene sands of quartz fluvial origin. Late Oligocene part may be Mogenstrup section is located in NW divided into six facies types (indicated by letters F to A, from top to bottom): Jutland, Denmark, on the east coast of Sailing, approximately 1 km South of the small village of Mogenstrup (UTM coordinates: NH , F metres of alternating mud, silt and Fig. 1). A locality near Hakkildgârd, approx sand deposited in a stormdominated tidalflat imately 1 km North of the Mogenstrup section, environment. was already mentioned by Ravn (1906, 1907). In E finegrained sand, deposited in his extensive work on the molluscan faunas from the Oligocene and Miocene Formations that nearshore, beach and backbarrier environments. thickness is up to 20 m. author considered the sediments to be of Middle D Lagoonal and protected bay deposits con Miocene age, but later (Ravn, 1924) this age sisting of black, homogeneous, organic mud assignment was questioned. Subsequently, with horizons of spherical calcite concretions. stratigraphie work was done in other localities in thickness of this facies varies from a a few the area (e.g. Sorgenfrei, 1940; Rasmussen, 1961; metres to about 60 m. Kristoffersen, 1972; Lieberkind, 1977), and a Late Oligocene or Early Miocene age C Shallowbay and shelf sediments consisting of chamositic mud, probably deposited close to a for comparable sediments suggested. Since the publication river mouth. This facies is found only in the of Ravn's papers, however, the outcrops at western part of the area (Fig. 5) where it Mogenstrup had not been studied until 1987, in reaches a thickness of 8 m. which year they were included in a facies analysis B Shelfsediments consisting of a glauconitic silt of the Late Oligocene sedimentary sequence in with layers of siderite concretions in the up NW Jutland (Beyer, 1987). That paper included a permost metre. thickness varies from less palaeomagnetic study, the results of which suggest than 1 m to 8 m. a Late Chattian age for this sequence. A silt, rich in diatoms, deposited in During field work along the coast North of Skive a thin glauconitic sand layer was discovered an anoxic environment, possibly an upwelling zone. thickness is less than one metre. in the coastal cliff at Mogenstrup. This sand was very rich in fossils, especially molluscs and spines sequence comprises at least three major of the echinoid Cidaris sp. present paper is hiatuses as indicated in Fig. 2a. At least two trans the result of a study of the rnolluscan fauna, which was found to comprise 197 taxa, 89 of which are gressions occurred during the time of deposition. During the first one, mainly finegrained material recorded from the Danish Late Oligocene for the was deposited (facies A and B). After a regression first time. a minor transgression molluscan assemblage is distinctly different from all known Danish Late Oligocene faunas, but occurred. This transgression saw the introduction of coarser material into the to the establishment of a barrier area, leading

4 42 Fig. 3. Sedimentary sequence at Mogenstrup (see Fig. 1 for location and Fig. 2b for legend) (author: Beyer). Fig. 2b. Legend to Figs 2a and 4 (author: Schnetler). Fig. 2a. Composite lithological column illustrating the Late Oligocene sequence in NW Jutland. Thicknesses which are highly variable in the area, are shown with maximum values (author: Beyer).

5 43 sediments are solely found in depressions around lagoon complex (C, D, E, and F), which was partly destroyed during a subsequent regressive salt structures and the thicknesses of the various period. This rests sequence on the Late Oligocene Branden Formation in the eastern part of the area while (Fig. 2), a primary boundary to the Palaeocene/Eocene Fur Formation is seen further to the West, where burrows extend from the facies vary greatly over short distances. Palaeotopography was probably influenced by synsedimentary halokinesis, with deposition mainly occurring in the subsiding areas between the salt structures. Erosion on top of these structures may Oligocene sediment down into the Fur Formation. have removed part of the sequence while sub deposition and preservation of this sequence was partly influenced by halokinesis. sidence between the structures led to a high preservation potential in these areas. Fig. 4. Suggested correlation of the Danish Late Oligocene with the established sequence stratigraphy of Haq, Hardenbol & Vail (1987). It should be noted that the short term changes in global sea level are not directly reflected in the Danish sedimentary facies associations: the shallow marine sequence originated in a period of relatively high global sea level, thus indicating that sedimentation was influenced by subsidence of the local basin rather than by changes in global sea level (author: Beyer).

6 44 LITHOSTRATIGRAPHICAL CORRELATION (AUTHOR: BEYER) R4a may be correlated with polarity chronozones C7a and C8, which implies an of about 27 Ma age sequence described is coeval with and very similar to the Late Oligocene Vejle Fjord Forma for the sediment that yielded here. the fauna described tion (Larsen & Dinesen, 1959) but it is clearly not directly correlative with this formation, the type locality of which is situated 200 km SE of Mogenstrup. Thus, according to standard stratigraphical procedure these two sequences should be given separate formation names. However, it is not desirable to add to the confusion over already existing Danish Late units. Oligocene lithostratigraphical That is why the Formation' is name 'Vejle Fjord also applied to the sequence described here. Similarly, facies B will be referred to as Brejning Clay Member, because of the close similarity to the Brejning Clay Member of the Vejle lithological Fjord Formation. It seems reasonable to assume that this facies was originally very widespread. Vejle Fjord Formation names will not be THE MOGENSTRUP SECTION (AUTHOR: BEYER) sequence outcropping at Mogenstrup (Fig. 3) can be divided into 4 units: Unit 1 Lightgreenish, clayey silt, rich in and glauconite, containing fragments of shells. In the uppermost metre three horizons of siderite concretions occur. se contain flattened horizontal burrows, indicating that compaction took place prior to cementation. thickness of the unit is at least 2 m. base is not visible. fauna described in the present paper was found in the uppermost 20 cm of this unit, where green sand, consisting entirely of glauconite and containing an extremely rich fauna, is associated with the concretions. used for the other facies, since these clearly have a very restricted geographical occurrence. Facies D is equivalent to the Vejle Fjord Clay and facies E and F are equivalent to the Vejle Fjord Unit 2 This unit has a sharp lower boundary and consists of 30 cm of dark brown, silt with mica and clayey Sand. gypsum precipitations, and contains organic C MAGNETOSTRATIGRAPHICAL CORRELATION (AUTHOR: BEYER) Based on magnetostratigraphical data, molluscan faunas and facies interpretations, Beyer (1987) suggested a Late Chattian age for most of the sequence (facies B to F) and correlated polarity zones R4b, N4, R5 and N5 with chronozone C7 polarity siderite and rhodocrosite. (2%), pyrite (c. 2%), Vertical vshaped cracks occur. Unit 3 About 40 cm of light brown, silty clay with mica and the jarosite precipitations, lower boundary of which is Small horizontal burrows sharp. are visible. sediment contains pyrite (8%), organic C (Fig. 4). Polarity zones Nl, R2, N2, R3, N3, and (4%), and shell fragments. Table 1. Mineralogy of the Mogenstrup sequence Legend: Qquartz, PLplagioklas, KFkalifelspar, SIDsiderite, PYpyrite, JAjarosite, RHrhodocrosite, GIgibbsite, FOfosforite, VIvivianite, GLAglauconite, MSmonosulfides. For the minerals JA, RH, GI, FO no calibration samples were available and only relative vertical changes can thus be inferred from these results. other values indicate percentage of the mineral content determined by Xray diffraction. lowermost two samples are from concretions. Metres Q. clay PL KF SID PY JA RH GI FO VI GLA MS org. C

7 45 Unit 4 Six m of silt with a variable sand and clay content. comm., 1987) examined two otolith samples (one Generally, grain size increases towards the top and from unit 1 and one from the very fossiliferous uppermost part of unit 1); both could be assigned an early Late Oligocene age. in the uppermost part lenses of longitudinal crossbedded sand occur. sediment is extensively biostratigraphy of the Chattian based on bioturbated and rich in mica and contains two horizons of scattered, spherical calcite concretions. pectinids (R. Janssen, 1979b) can be applied to the Mogenstrup fauna since a number of stratigraphically useful species were found, viz. Palliolum Also minor amounts of pyrite, vivianite and siderite occur. colour is brown but upon oxidation hausmanni hausmanni, P. limatum limatum, P. limatum it changes to black within a few minutes, which is ambignum, Pecten (Hilberia) bifidus bifidus and Pecten probably due to the presence of monosulfides. (Hilberia) soellingensis. Of these only Palliolum Based on sedimentological analyses the Mogen limatum ambignum and P. hausmanni hausmanni are strup section can be interpreted as follows (Fig. 4; Tab. 1): Unit 1, in the uppermost part of which the fauna described here was found, is thought to have been deposited in marine shelf environment. environmentof unit 2 was a depositional oxic, indicated by the content of rhodocrosite high and the occurrence of vshaped cracks which may well P. limatum limatum and Pecten preserved. (Hilberia) bifidus bifidus are represented by a few while a fragments, single nearcomplete specimen of Pecten collected. (Hilberia) soellingensis was pectinid fauna as whole indicates a a Chattian A/B most age, likely Chattian B, since the Chattian A species show signs of and this mollus reworking, indicate a supratidal marsh. An oxygendeficient can assemblage is probably not a biocoenosis. depositional environment is inferred for unit 3, because of the mineral assemblages (e.g. rich in gastropod species Hinia schlotheimi indicates, according to R. Janssen (1979b: 162), a pyrite), the facies association and burrows with a Chattian B or younger age. rather frequent very small diameter. This may have been a occurrence of this species at Mogenstrup is in shallow basin that came into existence at the start agreement with the age determination suggested of a transgression that inundated the lowlying by the pectinids. areas amongst the salt structures. Unit 4 was Two samples, one from unit 1 proper and one deposited in a brackish environment, probably a from the molluscrich upper part of unit 1 were lagoon. This interpretation is based on the facies association, the content of monosulfides, organic C, intensive bioturbation and the occurrence of small channel deposits in the of the uppermost part unit. analysed for foraminifers by Kaare Ulleberg, Norway, who concluded that both fall within samples the Angulogerina gracilis Zone, as does the Brejning Clay does 1987: (Ulleberg, 201) (K. Ulleberg, pers. comm.). geological history of the Mogenstrup section may thus be summarized as follows. Upon MATERIAL (AUTHOR: SCHNETLER) deposition in a marine shelf environment (unit 1) molluscs were collected almost exclusively followed a period of nondeposition and erosion from the uppermost part of unit 1. Large samples (unit 2). succeeding transgression flooded the of glauconitic sand were taken, dried and washed area and gave rise to a shallow, protected marine basin (unit 3). As more sandy on a 0.5 mm mesh. residues were handpicked after drying and fractioning. material was introduced into the area, barrier islands developed and the basin became a subsiding lagoon. Gradually During collecting in the field, larger specimens of e.g. Charonia flandrica, Orthosurcula regularis, the sedimentation rate exceeded the rate Scaphella siemsseni and Boreotrophon capito were of subsidence, which led to infill of the basin with coarser material (unit 4). regularly encountered, but the bulk of the material was collected by sediment processing. molluscs are generally well preserved but pyrite BIOSTRATIGRAPHICAL CORRELATION (AUTHOR: SCHNETLER) disintegration causes destruction of the rapid specimens, especially in the field. Late Oligocene age of the section is imme Very conspicuous among the nonmolluscs are diately apparent from the molluscan fauna and the very large number of spines of the echinoid this age determination is confirmed by a study of Cidaris sp. Marginal plates of asteroids, crinoid the otoliths. Dr P.A.M. Gaemers (Leiden, pers. brachials, Bryozoa and serpulids are also com

8 46 mon. Many Anthozoa are found, especially Ceratocyathus granulatus (Goldfuss, 1826), sp., and a few Flabellum sp. Many specimens brachiopod concentrations; ' Graphularia of the Terebratula sp. often in nestlike occur a few specimens of ' Terebratulina were also collected. Teleost otoliths are common; shark teeth, especially of the genus Pristiophorus, however, are less common. Foraminifers and ostracods are abundant, but decapods and balanids seem to be absent. A small molluscan fauna was collected from unit 3. molluscs are evenly sediment, species but not could be identified: distributed over the very common. following LIST OF MOLLUSCAN SPECIES (AUTHOR: SCHNETLER) In Table 2 all molluscan species from the Mogenstrup outcrop are listed. With a few exceptions, nomenclature follows R. Tanssen (1978b; 1979a, b). In column recorded from the Danish 1, species Late Oligocene for the first time are indicated with an asterisk (*). In column 2, the frequency of the specimens is indicated on a scale from 1 to 7: 1: Vil specimen 2: 23 specimens 3: 310 specimens 4: 1140 specimens Nucula (Lamellinucula) comta Yoldia (Yoldia) glaberrima Lentipecten (Lentipecten) corneus Dentalium (Dentalium) geminatum 5: specimens 6: specimens 7: more than 350 specimens Drepanocheilus (Arrhoges) speciosus number of scaphopods is estimated. Single Polinices (Euspira) helicinus valve of a bivalve = 1/2 specimen. Ancilla (Ancillus) karsteni Fusiturris selysii. In column 3, the depository of some rare species is indicated. Abbreviations: see systematic part. Table 2. Molluscan species from Mogenstrup (Brejning Clay Member, Vejle Fjord Formation) Bivalvia Leionucula peregrina (Deshayes, 1852) 2 Nucula (Lamellinucula) comta Goldfuss, Nucula (Lamellinucula) sp. 3 Nuculana (Saccella) westendorpi (Nyst, 1839) 3 Portlandia (Yoldiella) pygmaea (von Miinster, Munster, 1837) Yoldia (Yoldia) glaberrima (von Miinster, Munster, 1837) 41 3 Acar aff. dentiens (Cossmann & Peyrot, 1912) 5 PI. Pl. 1 Bathyarca Rathyarca bellula (Wiechmann, 1874) 4 Limopsis (Limopsis) aurita (Brocchi, 1814) 6 Limopsis (Pectunculina) (Peclunculina) lamellata chattica subsp. nov. 7 Limopsis (Pectunculina) retifera Semper, Limopsis (Pectunculina) vonderhochti 5 sp. nov. Glycymeris (Glycymeris) obovata (Lamarck, 1819) 2 Modiolula pygmaea (Philippi, 1843) 3 3 Musculus sp. Arcoperna 3 sp. Lentipecten (Lentipecten) corneus (Sowerby, 1818) 4 Propeamussium (Parvamussium) sp. Palliolum (s. lat.) (Speyer, venosum 1864) * 4 1 MNO Palliolum (s. lat.) limatum limatum (Goldfuss, 1833) 3 Palliolum (s. lat.) limatum ambignum (Anderson, 1958) 2 Palliolum hausmanni hausmanni (Goldfuss, 1833) 3 (s. lat.) Pecten (Hilberia) bifidus bifidus von Miinster, Münster, Pecten (Hilberia) soellingensis von Koenen, ENÁ ENA Limea (Notolimea) nysti (Speyer, 1864) 2 Anomia (Anomia) ephippium (Linne, (Linné, 1758) 3 Pododesmus (Heteranomia) squamula Linne, Linné,

9 Crassostrea cyathula cyalhula (Lamarck, 1806) 3 Cyclocardia (Cyclocardia) aff. kickxi (Nyst & Westendorp, 1839) 5 MNO Cyclocardia (Cyclocardia) depressa (von Koenen, 1884) 1 Astarte (Astarte) goldfussi Glibert, praecursor Glibcrt, Astarte (Astarte) gracilis gracilis (von Miinster, Munster, 1837) Goodallia (Goodallia) pygmaea (von Miinster, Munster, 1837) Parvicardium kochi (Semper, 1861) 1 MNO 41 4 Laevicardium (Habecardium) excomatulum Glibert Gilbert & van de Poel, Angulus posterus (Beyrich, 1868) 2 Abra (Abra) bosqueti (Semper, 1861) 4 Glossus subtransversus (d'orbigny, 1852) 2?.' Coralliophaga sp. Spaniodontella Spaniodonlella nilida nitida (Reuss, 1867) 1 MNO 2 Callista (Costacallista) (Coslacallista) beyrichi (Semper, 1861) 1 MNO Sphenia abscisa (Wiechmann, 1879) 1 MNO Corbula (Varicorbula) gibba (Olivi, 1792) 2 von von Spheniopsis depressa Koenen, MNO 4 Hiatella (Hiatella) arctica (Linne, (Linné, 1767) Teredinidae et indet. gen. sp. 4 Jouannetia (Jouannetia) aff. neuvillei Cossmann, MNO Lyonsia (Lyonsia) obovata von Koenen, AJB Thracia (Thracia) sp. 1 ISL Poromya (Poromya) hanleyana von Koenen, Cuspidaria (Cuspidaria) subcuspidata (d'orbigny, 1852) 3 3 Verticordia (Verticordia) aff. punctata Heering, 1950 sp. Pecchiolia 1 AJB indet. indct. Bivalvia 3 Scaphopoda Dentalium (Dentalium) geminatum (Goldfuss, (Goldluss, 1841) 5 Dentalium (Dentalium) polypleurum (Seifert, 1959) 4 Rhabdus sp. 2 Cadulus aff. subfusiformis (Sars, 1865) 1 MNO Gastropoda Scissurella (Anatoma) koenenia R. Janssen, Lepetella compressiuscula (Karsten, 1849) 2 Lepetella Lepelella helgae sp. nov. * 4 Lepetella jytteae nov. nov. sp. Astraea Aslraea (Lithopoma)pustulosa (von Miinster, Münster, 1844) 7 PI. 1 * (Collonia) troelsi sp. nov. Collonia 6 Homalopoma (Boutillieria) simplex (Philippi, 1843) 3 Pl.l Homalopoma (? Leptothyropsis) sp. 1 Tubiola subangulata sp. nov. 3 * Tubiola sp. 2? Skenea 3 sp. syst. part Leucorhynchia rotellaeformis rotellaejormis (Grateloup, 1828) 2 Cirsope (Cirsope) multicingulata (Sandberger, 1859) 3 Cirsope (Pseudocirsope) subeffusa subejjusa (Sandberger, 1859) 2 Cingula sp. 1 MNO Rissoa (Persephona) karsteni R. Janssen, Rissoa (Persephona) punctatissima R. Janssen, Alvania (Arsenia) semperi Wiechmann, Alvania 2 sp.

10 Ovirissoa 1 sp. Turritella (Haustator) (Haustalor) goettentrupensis goeltentrupensis (Cossmann, 1899) 1 MNO Bittium Biltium sp. 1 MNO Cerithiopsis (Metaxia) aff. degrangei (Cossmann & Peyrot, 1921) 2 henckeliusii Cerithiopsis henckeliush 3 (Nyst, 1836) 4 Cerithiopsis jutensis Schnetler, 1985 \ Cerithiopsis (s. lat.) serrula R. Janssen, Cerithiopsis (s. lat.) daphnelloides R. Janssen, 1978 * 3 Cerithiopsis (s. lat.) ariejansseni R. Janssen, Cerithiopsis (s. lat.) antonjansei sp. nov. * 2 Laiocochlis (Laiocochlis) supraoligocaenicus sp. nov. nov. * 2 Cerithiopsida boelschei (von Koenen, 1891) 41 Cerithiella bitorquata (Philippi, 1843) 4 Seila (s. lat.) angusta angusla Tembrock, Seila koeneni R (s. lat.) Janssen, MNO Triforis (Trituba) (Trituha) sorgenfreii sp. nov. * 3 Biforina (Biforina) Grundel, praeversa Griindel, 1975 * 4 Norephora (Norephora) elatior 4 (von Koenen, 1891) \ Opalia (Pliciscala) pusilla (Philippi, 1843) 1 Turriscala (Turriscala) cf. rudis (Philippi, 1843) 2 Acirsa (Plesioacirsa) leunisii (Philippi, 1843) 2 Cirsotrema (Opaliopsis) subglabrum sp. nov. 2 Cirsotrema (Opaliopsis) sp.l 1 Cirsotrema (Opaliopsis) sp.2 1 Cirsotrema (? Opaliopsis) aff. koeneni A.W. Janssen, Cirsotrema (s. lat.) crispatum Harder, JHR Amaea (Secalina) amoena amoena (Philippi, 1843) 1 ISL Archileclonica Architectonica sp. 2 Mathilda (Mathilda) sandbergeri sandbergen (Koch, 1876) 2 Aclis (Aclis) vetusta Wiechmann, Aclis (Stilbe) proneglecta R. Janssen, Aclis (Graphis) hosiusi (Lienenklaus, 1891) 4 Balcis (Balcis)? lamberti (Cossmann, 1882) * 1 MNO Balcis alba naumanni (von Koenen, 1867) 41 Balcis (Polygyreulima) pseudonaumanni R. Janssen, Eulima (Eulima) emersa Speyer, Niso (Niso) minor Philippi, Neojanacusplanatus (Speyer, 1864) 3 PI. 2 Drepanocheilus (Arrhoges) speciosus (von Schlotheim, 1820) 7 Polinices (Euspira) helicinus (Brocchi, 1814) 7 Apiocypraea? humbergi R. Janssen, 1978 * 2 Semicassis (Echinophoria) rondeleti (Basterot, 1825) 4 Charonia (Sassia) flandrica (de Koninck, 1837) 5 Ficus (Ficus) concinnus (Brongniart, 1822) 3 Pterynotus (Pterochelus) tristichus (Beyrich, 1854) 2 Boreotrophon Boreolrophon (s. lat.) capita capito (Philippi, 1843) 4 Lyrotyphis (Eotyphis) sejunctus (Semper, 1861) 5 * Trophonopsis 1'rophonopsis angustevaricata angustevancata (Gripp, 1915) 5 Coralliophila (Hirtomurex) kochi (Beyrich, 1854) 3 PI. 2 Scalaspira (Scalaspira) elegantula eleganlula aequistriata aequistnata (Speyer, 1863) 4 Scalaspira (Scalaspira)? waeli (Beyrich, 1856) 1 MNO Scalaspira (Scalaspira) sp. 5 PI. 3 Searlesia ravni sp. nov. 3 Exilioidea elatior (Beyrich, 1848) 2 Parvisipho (s. lat.) scrobiculatus (Boll, 1851) 1 ISL Coptochetus (s. lat.) danicus Schnetler, Metula (Daphnobela) scabricula (Philippi, 1843) 41 Pisanella subgranulata (von Schlotheim, 1820) 3 * Buccinidae gen. gen. et indet. 2 sp.

11 Hinia (Tritonella) schlotheimi (Beyrich, 1854) 4 Streptochetus Slreptochetus (Streptodictyon) (Streptodktyon) cheruscus cheruscus (Philippi, 1843) 6 Streptochetus (Streptolathyrus) soellingensis Tembrock, Latirus (Pseudolatirus) sp. Scaphella (Scaphella) siemsseni (Boll, 1851) Ancilla (Ancillus) karsteni (Beyrich, 1856) 5 Gibberula brevis (von Koenen, 1890) 4 PI. 3 Cancellaria (Merica) evulsa postera (Beyrich, 1856) 3 Babylonella pusilla (Philippi, 1843) 5 Vexillum (Uromitra) hastatum (Karsten, 1849) 41 Conomitra Conomilra soellingensis (Speyer, 1864) 5 Orthosurcula Orlhosurcula regularis reguláris (de Koninck, 1837) 5 Acamptogenotia Acamptogenolia morreni (de Koninck, 1837) 2 5 Stenodrillia obeliscus (des Moulins, 1842) Cochlespira volgeri (Philippi, 1843) 41 Gemmula (Gemmula) laticlavia (Beyrich, 1848) 4 Gemmula (Gemmula) trispiralis R. Janssen, Gemmula (Gemmula) pseudokonincki pseudokomncki R. Janssen, 1979 Pleuroliria konincki (Nyst, 1845) 3 Fusiturris duchastelii (Nyst, 1836) 6 Fusiturris selysii (de Koninck, 1837) 4 Fusiturris enodis R. Janssen, Glibertturricula ariejansseni (Schnetler, 1987) 4 Clavatula mogenstrupensis sp. nov. 2 Cordieria plicata (Beyrich, 1848) * 1 ISL Rathytoma Bathytoma (Bathytoma) leunisii (Philippi, 1843) 5 Asthenotoma holzapfeli (von Koenen, 1890) Splendrillia koeneni (Speyer, 1867) * 3 4 Microdrillia (Microdrillia) ingerae sp. * nov. 3 Microdrillia (Andersondrillia) brejningensis brejmngensis sp. nov. * 4 'Microdrillia' Microdrillia speyeri (Koch & Wiechmann, 1872) 3 PI. 3 Boreodrillia undatella undalella (Speyer, 1867) 4 1 Pleurotomella Pleuroiomella anderseni Schnetler, 1987 MNO Pleurotomella (Pleurotomella) margaritata margantata R. Janssen, 1978 * 3 PI. 3 * Pleurotomella (Pleurotomella) rasmusseni sp. nov. 2 Rimosodaphnella lappanni sp. * nov. 4 Conus (Leptoconus) semperi Speyer, Hastula (Haslula) (Hastula) beyrichi (Semper, 1861) 1 MNO Chrysallida sp. 1 MNO Ebala sp. 1 MNO Evalea kochi (Gorges, 1952) 3 Evalea incrassata (von Koenen, 1867) 1 ISL Odostomia Odoslomia 1 MNO sp. Syrnola (Syrnola) subcylindracea (Philippi, 1843) 5 Syrnola (Puposyrnola) laevissima (Bosquet, 1859) Turbonilla (Strioturbonilla) (Slrioturbonilla) wiechmanni Speyer, 1870 * 2 1 AJB Turbonilla (Pyrgolampros) jeffreysi jejfreysi Koch & Wiechmann, p? * Actaeopyramis Aclaeopyramis (s. lat.) pseudopunctata nov. 3 sp. Actaeon (Actaeon) punctatosulcatus (Philippi, 1843) 3 Crenilabium terebelloides (Philippi, 1843) 2 Ringicula (Ringiculina) striata (Philippi, 1843) 4 Roxania utriculus (Brocchi, 1814) 4 Philine kochi von (Ossiania) Koenen, MNO Limacina hospes (Rolle, (Rollc, 1862) 4 Limacina valvatina (Reuss, 1867) 3 Clio nielseni A.W. Janssen, 1990 * 4 Gastropoda inc. sed. 1 1

12 Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coll. Coastal Pl. Mogenstrup Most Glauconitic This 50 Stratum typicum sand, Brejning Clay Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). Derivatio nominis new subspecies of Limopsis lamellata is named after its occurrence in the Chattian stage of the Oligocene. Holotype 1, Figs 2ab, (coll. MGUH ) (leg. K.I. Schnetler). shell is small, equivalve and rather convex with a somewhat obliquely ovate to subcircular outline. It is slightly inequilateral, and the orthogyrate, dorsally projecting umbo has a slightly prosocline position. hinge line is straight, the anterior and posterior dorsal margins Fig. 5. Other Danish localities mentioned in this paper (author: Schnetler). SYSTEMATIC PART (AUTHOR: SCHNETLER) Danish localities mentioned in this section are shown in Fig. 5. depositories of the materialare indicated by abbreviations: are of almost the same length. transition of the anterior dorsal into the anterior convex margin margin is gradual; the posterior dorsal margin and the form posterior margin together an obtuse angle. posterior margin is more or less convex. transitions of both the anterior and the posterior margin into the convex ventral are also margin gradual. cardinal area is flat and triangular with a AHF S.B. Andersen and H.C. rather small triangular resilifer below the umbo. Flansen, Fredericia, DK. dentition is divided into two series of teeth by AJB A.C. Janse, Brielle, NL. an edentulous gap below the umbo. anterior DGU Geological Survey of Den series consists of three to four teeth which are mark, Copenhagen, DK. aligned almost dorsoventrally. anterior series ENA E. Nielsen, Arhus, DK. is straight. arched posterior series also consists FFIK F. von der Hocht, Kerpen of three to four teeth which are placed obliquely. Balkhausen, F.R.G. sculpture consists of flat concentric ribs ISL K.I. Schnetler, Langa, DK. which are distributed somewhat irregularly. JHR J. Hillersborg, Randers, DK. spaces between these ribs are much wider than the LJL L.B. Jorgensen, Lime, DK. ribs and finer ribs are visible in the interspaces. MGUFI Geological Museum, Univer radial ribs are weaker and the sculptural com sity of Copenhagen, DK. ponents together form a more or less reticulate or MNO M.S. Nielsen, Odense, DK. namentation. RGM National Museum of Natural shell margin has a fine crenulation which History (formerly: Rijksmuseum corresponds to the radial ribs. crenulae are van Geologie en Mineralogie), fairly regular in strength. palliai line and the Leiden, NL. adductor scars are not very distinct. anterior RLO R. Lechner, Odense, DK. impression is small and rounded, the posterior one SMF Forschungsinstitut Sencken is larger and rounded trapezoidal. berg, Frankfurt am Main, F.R.G. Measurements specimens have a height of WLH W. Lappann, Heiligenhaus about 2 mm. maximum height is c. 3 mm. F.R.G. Isenbiigel, lamellata chattica Limopsis (Pectunculina) Locus typicus subsp. nov. PI. 1, Figs 2ab, 3ab cliff at Mogenstrup, North of Skive, Jutland, Denmark (Fig. 1). Paratypes (coastal cliff): 1/2 specimen (Pl. 1, Figs 3ab) (leg. K.I. Schnetler; coll. MGUH 20 3/1 and 300/2 024); specimens (leg./coll. ISL); 1/1 and 48/2 specimens (leg./coll. JHR); 7/2 11/2 specimens (leg./coll. LJL); specimens (leg./coll. ENA); 2/2 specimens (leg. K.I. Schnetler, coll. 1/1 MNO); and 32/2 speci

13 Coastal Glauconitic Eriksen, Pl. 51 mens (leg./coll. ACJ); 292/2 specimens (leg./coll. Derivatio nominis species is named after Mr MNO); 5/2 specimens (leg. K.I. Schnetler; coll. RGM ); 5/2 specimens (leg. K.I. Fritz von der Hocht, geologist, KerpenBalkhausen, F.R.G. Schnetler; coll. SMF ). Hadsten (road excavation): 1/2 specimen (leg./coll. ISL). Brejning (beach exposure): 40/2 specimens (leg./coll. MNO); 3/2 specimens (leg./coll. ISL). Bogeskov (coastal cliff): 2/2 specimens (leg./coll. MNO). Discussion and L. lamellata new subspecies Holotype (leg. K.I. Schnetler). roundedovate, 1, Figs 4ab, (coll. MGUH ) shell is equivalve with a slightly oblique outline, inequilateral. slightly opisthogyrate umbo has a prosocline it position; projects only slightly beyond the dorsal shell is margin. strongly convex, with the maximum at about 2/3 of the convexity Lehmann, 1885 s. str. are very close with regard to size, outline, convexity and However, judging sculpture. from the numerous available specimens of the new subspecies and specimens of L. lamellata s.str. from the Miocene of Dingden and WinterswijkMiste ( Netherlands) (coll. ISL) (type locality!), Twistringen (F.R.G.), a shell height. convexity is less obvious in juvenile specimens. hinge line is straight. of the length anterior dorsal is about two thirds of that margin of the posterior dorsal margin, and these margins are almost in line. Both the anterior and the posterior margins are almost straight in their dorsal number of features seem to be different in the parts. transitions into the ventral margin are Danish material. In L. lamellata s.str. the cardinal area is higher with a more projecting and stronger umbonal region and a resilifer. number of teeth larger in L. lamellata s.str. in the anterior and posterior series is 46 and these teeth are somewhat stronger. Furthermore, the sculpture on the Miocene form is generally stronger, especially the gradual, posteriorly with a somewhat stronger curvature than anteriorly. cardinal area is flat and and subtriangular, provided with a fine striation parallel to the hinge line. rather large resilifer is placed below the umbo. Its outline is that of a rightangled triangle with the shorter cathetus placed anteriorly. resilifer is provided with a striation similar to that concentric ribs, and finally its maximum size is of the cardinal area. dentition is divided into two series of teeth by an edentulous below the gap umbo. anterior series consists of 46 teeth about 5 mm. Although not very obvious, these differences seem to be constant. Because of the difference in I age prefer to treat the Danish material as a of L. lamellata. stratigraphical subspecies nominal subspecies is distributed in the widely which are aligned almost dorsoventrally. arched series consists of 48 posterior, slightly more obliquely placed teeth, the posterior 23 of which are almost parallel to the hinge line. Middle Miocene of the North Sea Basin (Hemmoorian and Reinbekian). number of teeth is variable, but in all cases the posterior row of teeth is longer than the anterior Limopsis (Pectunculina) vonderhochti sp. nov. one. Adult specimens with 45 anterior and 56 PI. 1, Figs 47 posterior teeth are most frequent. sculpture is dominated by numerous con? 1884 Limopsis Goldfussi Nyst von Koenen, in Speyer & centric riblets which follow the growth lines. von Koenen, pi. 31, figs 910 (non Nyst).? 1937 Limopsis Goldfussi Nyst?) 1941 Limopsis Goldfussi Nyst non Nyst). p. 141 (non Nyst Gorges, 160 p. (partim, number of these riblets is about one hundred on specimens with a height of 10 mm. concentric elements alternate sculpture frequently in strength. radial sculpture is much weaker and 1952 Limopsis Goldfussi Nyst Gorges, p. 17 (non Nyst). 1979b Limopsis (Pectunculina) retifera Semper, 1861 R. Janssen, p. 30, pi. 1, fig. 16 (partim, nonsemper). almost exclusively visible in between the concentric ribs, giving the interspaces a serrated appearance. Locus typicus cliff at Mogenstrup, North inner shell margin has a coarse crenulation of Skive, Jutland, Denmark (Fig. 1). on the anterior part of the ventral margin. This Stratum typicum sand, Brejning Clay crenulation consists of 46 coarse knobs which are Member of the Vejle Fjord Formation (Late elongated in dorsal direction as radiating ribs on Oligocène, Chattian B). the inner shell surface. In the available material

14 Mogenstrup, This 52 these ribs are somewhat variable in strength. Kirstinebjerg Skov (beach exposure): remaining part of the ventral margin is usually 1/2 specimen (leg./coll. ISL). smooth, but in a few shells additional knobs are indicated, especially near the transition into the posterior margin. crenulations do not correspond with the radial ribs on the outside of the shell. KrefeldLinn (water well) (Chattian A and B): 1/2 specimen, PI. 1, Fig. 6ab, (leg. WHL, coll. SMF ), 7/2 specimens (leg.coll. FHK). KrefeldLinn water ('Maizena' well) (Chattian pallial line and the adductor scars are not A): 1/2 specimen, (leg./coll. FFIK). very distinct. A welldeveloped myophore is situ Krefeld (borehole GLA1): 1/2 specimen (Chat ated below the anterior part of the hinge. ante tian A), (leg./coll. FFIK); 1/2 specimen (Chattian rior adductor scar is subovate and much smaller B), (leg./coll. FHK). than the roundedtrapezoidal posterior one. Matzerath(Schacht S.J. 8): 1/2 specimen (310.5 Range of variation m) (leg./coll. FHK); 7/2 specimens ( Danish material demonstrates a rather wide range of variationwith regard m); 1/1 + 1/2 specimen ( m) (both leg./ to the convexity. width/height ratio varies coll. FHK). from 0.33 to 0.46, this figure being higher in larger specimens. Danish material was compared with specimens the German localities Ahnetal (coll. from SMF), KrefeldLinn and (coll. WLH) Meerbusch Dalheim (borehole 50/19): 1/2 specimen (Chattian A) (leg./coll. FHK); 1/2 specimen (Chattian B) (leg./coll. FHK). MeerbuschIlverich 1/1 and 1/2 (water well): specimens (Chattian A) (leg./coll. FHK); 4/2 Osterath (coll. FHK). Those from Ahnetal and specimens (Chattian B) (leg./coll. FHK). KrefeldLinn demonstrate a lesser degree of convexity (length/height ratio Also the ). crenulation of the anterior ventral margin is coarser, with the knobs situated more posteriorly. In addition, the sculpture is more accentuated. From MeerbuschOsterath a of sample 38/2 and 1/1 specimens was available. This population agrees more with the Danish material. closely ratio varies from 0.28 to length/height 0.39, and the crenulationof the anterior ventral margin and the external sculpture are rather similar to the Danish specimens. differences in the length/height ratios may be the result of local environmental influences and therefore I confidently refer the German populations to the new species. MeerbuschOsterath 1/1 and (water well): 38/2 specimens (Chattian A + B) (leg./coll. FHK). Willich (water well): 2/2 specimens (Chattian A) (leg./coll. FHK). NeukirchenVluyn (Schacht 5): 1/2 specimen (Chattian A) (leg./coll. FHK). (water well): 1/2 specimen (Chattian A) (leg./coll. FHK). MoersKapellen (Kieswerk): 2/2 specimens (Chattian B) (leg./coll. FHK). MoersRumeln (Schacht Rumeln): 1/1 and 3/2 specimens (Chattian A) (leg./coll. FHK); 3/2 specimens (coll. SMF); 5/2 specimens (leg./coll. AJB). Ahnetal: 3/1 and 9/2 SMF 308 specimens (coll. coll. SMF 407) (leg. J. Gorges; ). Paratypes coastal cliff: 1/2 Niederkaufungen: 1/2 specimen (coll. SMF 308 specimen (Pl. 1, Fig. 5ab) (leg. K.I. Schnetler; 409). coll MGUH ); 69/2 specimens and 21 frag Discussion large Limopsis species has ments (leg./coll. ISL); 20/2 specimens (leg./coll. 5/2 JHR); specimens (leg./coll. LJL); 1/2 specimen (leg. K.I. Schnetler; coll. MNO); 2/2 specimens (leg. K.I. Schnetler; coll.rgm 229 previously often been referred to L. goldfussi (Nyst, 1845), a restricted species the to Rupelian. A specimen of L. goldfussi was found by Harder in (1913: 52, pi. 3, the Middle fig. 18ac) 790); 2/2 specimens (leg. K.I. Schnetler; coll. SMF ); 1/2 specimen, leg. K.I. Schnetler, coll. WLH; 3/2 specimens and 2 fragments (leg./coll. AJB), 86/2 2/2 specimens (leg./coll. MNO), specimens (coll. FHK, leg. K.I. Schnetler). Oligocene fine sand at Arhus. This specimen cannot be referred to L. retifera Semper, 1861, as was assumed by R. Janssen (1979: 30). new species differs from L. goldfussi by the lower number of teeth in the posterior series, a dif Brejning (beach exposure): ferent outline, a crenulated margin, a different 1/1 and 5/2 specimens (leg./coll. MNO); 2/2 specimens (leg./coll. sculpture and a more convex shell. R. Janssen ISL); 1/2 specimen (leg./coll. RLO); 1/2 specimen (1979: 30, pi. 1, fig. 16) included specimens of the (leg./ coll. AHF). new species in L. retifera, from which it differs by

15 A Pl. Mogenstrup Coastal Coastal Glauconitic Glauconitic This This 53 several features: L. retifera has a regularly Lepetella helgae sp. nov. crenulated margin, consisting of numerous fine PI. 1, Fig. 9ab. denticles, which are almost equal in strength and correspond with the radial sculpture on the outside Locus typicus cliff at Mogenstrup, North of the shell. In the new species the posterior series of Skive, Jutland, Denmark (Fig. 1). of hinge teeth is always the longer, while in L. Stratum typicum sand, Brejning Clay retifera the anterior series is. In the new species the sculpture is dominated by the concentric ribs, Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). while in L. retifera the sculpture is more cancellate. Derivatio nominis new species is named after Finally, the new species is generally larger than L. my daughter, Helga Lund Schnetler. retifera, the auricles are less distinct and the shell is Holotype 1, Fig. 9ab, (coll. MGUH ) than that of L. more equilateral retifera. new species differs from the Miocene L. anomala auct. (non von Eichwald, 1830?) by having more convex valves, with a coarser more sculpture widely spaced concentric elements. coarse knobs on the inner ventral margin are placed more posteriorly in L. anomala and the entire ventral margin (leg. M.S. Nielsen). shell is small and capshaped. is apex highly elevated and situated precisely above the posterior apertural margin. protoconch resembles a small button. aperture is almost circular. In lateral view the anterior outline of the shell is whereas convex, is crenulate. resilifer in L. anomala has an the posterior outline is almost straight. re is no acuteangled triangular outline, while the outline external sculpture. growth lines are well of the resilifer in the new species is rightangled marked, concentric and somewhat variable in triangular. strength. On the inside of the shell the horseshoeshaped Remarks Propeamussium (Parvamussium) sp. rather limited material, consisting of muscular impression is fairly distinct. Paratypes (leg./coll. MNO). (coastal cliff): 1 specimen 11 defective valves and fragments, is available. Mr Discussion new species is related to the very A.C. Janse (pers. comm., 1988) noted that on the fragments in his collection from Mogenstrup the number of radial ribs on the inner side of the shell differs, in to P. comparison pygmaeum (von from the Miinster, German and Danish 1835) variable species L. compressiuscula (Karsten, 1849). In L. compressiuscula, however, the shell is distinctly compressed laterally and the is situated above apex the centre of the elliptical aperture. L. compressiuscula is also less elevated. relatively Late Oligocene. Mogenstrup specimens all have 11 radial ribs on the inside of both the left Lepetella jytteae sp. nov and right valve, while specimens of P. pygmaeum PI. 1, Fig. loab show 1517 ribs on the right valve and 1214 on the left one (see also R. Janssen, 1978a: 48). Locus typicus cliff at Mogenstrup, North sculpture on the outside of the right valve of Skive, Jutland, Denmark (Fig. 1). is rather similar to that of P. pygmaeum. left Stratum typicum sand, Brejning Clay valve has a cancellate sculpture, consisting of about 35 concentric ribs and numerous line radial Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). lines. Derivatio nominis species is named after Mrs there are consistent differences between the Mogenstrup material and P. pygmaeum. Apparently, More and better material is needed preserved for a proper interpretation of the Mogenstrup form. Mr A.C. referred to Janse (pers. comm., 1989) two taxa that be related: P. might duodecemlamellatum (Bronn, 1831), from the Early Pliocene of Italy and P. duodecemlamellatum bronni (Mayer, 1861) from the Middle Oligocene of Hungary. Jytte Hillersborg, who found and donated the holotype. Holotype Pl. 1, Fig. loab (coll. MGUH ) (leg. J FIR). shell is rather large and cap shaped. aperture is suboval, with its greatest width in the anterior part. shell height equals about half the diameter of greatest the aperture. apex is situated at about five sixths of the shell

16 This PI. Coastal Glauconitic A 54 length. anterior outline (lateral view) is convex, while the posterior part is almost straight. visible. aperture is subcircular with a sharp labrum. columella is concave, with a some protoconch is buttonshaped and smooth. what thickened inner lip demarcating the umbili On the juvenile shell numerous growthlines are cus. A vertical impression is visible the on columel visible, indicating that it had an almost circular la, which most probably indicates the position of the operculum. aperture. About three millimetres from the protoconch a rather strong concentric sculpture appears, whorls have a glossy appearance. consisting of strong ribs, separated by interspaces of almost equal strength. On the largest available specimen about 20 such ribs are present. concentric ribs are crossed by weaker radial ribs, the number of which increases towards the apertural margin, because secondary radialribs appear in between the primary ones. radial ribs are rather An extremely fine spiral furrow is present near the adapical suture. Prosocline growthlines are rather distinct. On the largest specimen two former apertures are visible as rather prominent prosocline lines. Paratypes Mogenstrup (coastal cliff): 3 specimens (leg./coll. ISL); 25 specimens (coll. ISL/leg. MNO); 200 specimens (leg./coll. MNO). prominent on the adapical side of the concentric Discussion new species differs from Collonia ribs, but are also visible in the interspaces. On the inside of the shell the horseshoeshaped (C.) minima (Philippi, 1843) by several features, although juvenile specimens at first look rather muscular impression is fairly distinct. similar to this species. protoconch of C. minima Paratypes Mogenstrup (coastal cliff): 25 comprises one whorl, the nucleus is rather small specimens (leg./coll. MNO); 2 specimens (coll. ISL, leg. MNO). Remarks rather large Lepetella species differs from congeners by its strong sculpture. and the transition to the teleoconch is not sharp. In C. minima the whorls increase more gently in diameter. On its base a sharp furrow is present close the to rib. demarcating spiral This spiral, as well as those within the umbilicus, are coarser in C. Collonia (Collonia) troelsi sp. nov. minima. Adapically, its aperture adapically has a Locus typicus PI. 2, Fig. 2ac cliff at Mogenstrup, North sharper edge, and it is generally new species. smaller than the of Skive, Jutland, Denmark (Fig. 1). Stratum typicum sand, Brejning Clay Homalopoma (? Leptothyropsis) sp. PI. 1, Figs 12ac. Memberof the Vejle Fjord Formation, Late Oligocène (Chattian B). rather limited material, consisting Derivatio nominis species is named after my of a single defective shell and seven fragments ofthe son, Troels Lund Schnetler. same specimen, was collected (leg. E. Nielsen, Holotype 2, Fig. 2ac (coll. MGUH ) Arhus). Unfortunately, the protoconch is missing, (leg. K.I. Schnetler). and the state of preservation is rather poor, for shell is small, rather thinwalled which reasons it has not been possible to make a and wider than high, juvenile specimens very much final interpretation of the material. so. protoconch consists of three quarters of a shell is rather small, trochiform, about as whorl, the nucleus being relatively large and slightly depressed. boundary with the teleoconch is high as wide. defective specimen consists of only 1 3/4 whorls, which are moderately convex and sharp and indicated by an almost orthocline radial separated by deep sutures. y increase very furrow. In the the teleoconch consists of largest specimen two convex whorls, rapidly increasing in diameter, rapidly in diameter. body whorl equals about five sixth of the (estimated) total shell height; the aperture is about and separated by deep sutures. last whorl half as high as the complete shell. base is only equals about nine tenths, the height of the aperture slightly convex, with a rather wide umbilicus, surrounded by a distinct spiral rib. aperture is about six tenths of the total shell height. periphery is regularly convex. base of the shell is provided with a rather narrow umbilicus, which is oval, the labrum is broken. columella is concave, and the inner lip has a rather thick callus. separated from the base by two closeset spiral ribs. In the umbilicus about four weaker spiral ribs are spiral sculpture on the middle whorls consists of a thin, sharp spiral carina, which is project

17 Pl. Coastal Glauconitic L. Mogenstrup 55 ing and situated immediately above the abapical suture. On the base of the shell three further, similar spirals are present. four spirals are regularly spaced with concave interspaces. distance between the lowermost of these spirals and the umbilical spiral is On the distinctly larger. largest fragment two secondary spirals develop between the two adapical spirals. radial sculpture consists of about 30 prosocline folds, separated by somewhat narrower interspaces. Towards the upper suture these folds are slightly accentuated, gradually becoming spiral is visible. the labrurn is aperture is almost circular; not thickened, but the columellar part of the inner lip is slightly consolidated. dominant sculptural element is a rather strong spiral rib, lying upper suture and dividing at a short distance from the the whorl into two parts: a flat adapical ramp, and a lower part which adapically is almost flat, and becomes slightly convex below. two parts of the whorl together form an obtuse angle. transition into the base is regularly rounded. growth lines are barely visible, but almost orthocline. weaker and less welldefined downwards, and they Paratypes Mogenstrup (coastal cliff): 2 disappear between the two adapical spirals. radial sculpture forms small projections on the specimens (leg./coll. MNO); 1 specimen (leg. M. S. Nielsen; coll. ISL). spirals. Distinct prosocline growth lines are pres Discussion species differs from the rather ent. From the upper suture they run slightly undulating downwards, especially well visible on the radial folds. Also between the spirals they are similar Tubiola sp. 1 (A. W. Janssen, 1984: 128, pi. 6, fig. 3ab) by having more angular whorls. Daronia dingdensis Anderson, 1964 has a concave distinct. adapical ramp and a demarcating carina which Remarks shell seems to be rather similar to projects beyond the ramp. Below this carina the Homalopoma (Lepthothyropsis) philipiana Dall (see Wenz, 1938: 341, fig. 799) in general outline, but additional material is needed. whorl is slightly concave, changing downwards to strongly convex. Its apex projects only slightly over the last whorl. Tubiola subangulata sp. nov Tubiola sp. PI. 1, Fig. 14 PI. 2, Fig. 1 Locus typicus cliff at Mogenstrup, North shell is very small and consists of Skive, Jutland, Denmark (Fig. 1). of 2 3/4 moderately convex whorls which are Stratum typicum sand, Brejning Clay Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). almost canaliculate separated by deep, sutures. shell is almost as high as wide. protoconch comprises slightly more than half a whorl, Derivatio nominis subangulata ( = near and is clearly separated from the teleoconch by a angular). species is named after the obtusely angular whorls. Holotype (leg. M.S. Nielsen). 1, Fig. 14; (coll. MGUH ) shell is very small and relatively radial orthocline rib, and by a difference in colour. nucleus is relatively large. body whorl equals about seven eighths of the total shell height. periphery is regularly convex with a transition into gradual the base. thickwalled, about 1 /% times as wide as high. It rather deep, but not very wide umbilicus is consists of about three convex whorls which rapidly increase in diameter. consists of protoconch half whorl a rather voluminous and inflated nucleus has a very finely granulated only. surface. separated by a distinct spiral rib which is granulated by the lines. One additional growth rib is visible in the umbilicus. spiral is almost aperture circular, and neither labrum nor inner lip are thickened. shell's boundary with the teleoconch is only indicated by the appearance of a spiral rib on the adapical part of the first postembryonic whorl. surface is smooth and glossy, with rather weak almost orthocline growth lines. body whorl equals total shell height. about seven eighths of the Material (coastal cliff): 2 specimens (leg./coll. MNO), 1 specimen (PI. 2, Fig. 1) (leg. base of the shell is slightly convex and MNO, coll. MGUH ). demarcated from the deep and rather wide um Discussion shell differs from Tubiola sp. 2 bilicus by a spiral rib. In the umbilicus one further from the Hemmoorian of WinterswijkMiste

18 PI. Coastal Glauconitic This 56 (A. W. Janssen, 1984: 128, pi. 6, fig. 8) by having shell is rather small and slender a lower apex and a less concave columella. turriculate; about three times as high as wide. last whorl equals slightly less than third one of the? Skenea sp. PI. 2, Figs 3ab total shell height. protoconch consists of four and a half Only five juvenile specimens are moderately convex whorls which are separated by rather distinct sutures. nucleus is small. From available. shell is very small and planispiral. the second protoconch whorl onwards the whorls protoconch consists of half a whorl, the nucleus is voluminous. transition into the teleoconch is sharp and demarcated radial rib. by a teleoconch consists of 3/4 convex whorl It has only. a and very convex periphery, an almost flat adapical part, which is separated from the lower of the part whorl by a spiral. transition into the convex base is gradual. umbilicus is wide and demarcated rib. by a spiral is subcircular and the labrum is broken. aperture the umbilicus the base of the nucleus is Through visible. sculpture consists of very fine, threadlike spiral riblets. A spiral near the suture in adapical is particular rather distinct, while those on the periphery are weaker. On the base the spiral demarcating the wide umbilicus is very distinct. growth lines are opisthocline near the adapical suture. Remarks material shows a stronger spiral sculpture than does? Skenea sp. (A.W. Janssen, 1984: 129, pi. 6, fig. 9ab). Ovirissoa sp. only specimen found is very immature and shows no teleoconch sculpture. Yet, the apical angle would indicate that the specimen is more related to closely O. gottscheana (von Koenen, than to O. 1882) westfalica (A.W. Janssen, 1967), which has a distinctly wider apical angle. Cerithiopsis (s. lat.) antonjansei sp. nov. PI. 2, Fig. 9. are provided with a radial sculpture, consisting of numerous fine riblets. Initially these riblets are almost orthocline, but they become slightly opisthocline on the last protoconch whorls. A spiral sculpture consisting of five or six threads is spiral present on the last two and half a protoconch whorls. se spirals are situated the on abapical part of the whorl and, increase in although they strength in abapical direction, remain weaker they than the ribs. two elements spiral sculptural together result in a reticulate ornament of the of the abapical part whorl. transition to the teleoconch is sharp and opisthocline. teleoconch consists of five and a half whorls in the largest specimen. se whorls are convex and sutures. separated by deep last whorl less than one third of the total shell equals slightly the with height, aperture the siphonal canal about one fourth. aperture is roundedoval, the labrum is broken and the columella is concave. two prominent spiral ribs (see below) cause an angular periphery. base is almost flat. spiral sculpture starts immediately behind the boundary with the protoconch with three of which spirals, the adapical one is threadlike and situated immediately below the suture. This does not increase spiral in strength on the following whorls, but it gradually becomes more distant from the suture. two adapical abapical spirals are prominent, especially the upper one, which is placed on the centre of the whorl. lowermost lies spiral at a short distance above the abapical suture. Inbetween the spirals the whorl is concave, especially so between the two lower spirals. A rather weak spiral rib, demarcating the base of Locus typicus cliff at Mogenstrup, North the whorl, is only visible on the last whorl. Close of Skive, Jutland, Denmark (Fig. 1). Stratum typicum sand, Brejning Clay Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). to this demarcating spiral a weaker rib is spiral present on the base. radial sculpture consists of slightly opisthocline ribs which are much narrower than the Derivatio nominis species is named after my spaces between them. number of these ribs is friend Anton C. Janse, Brielle, Netherlands. 15 on the first postembryonal whorl and 14 on the Holotype 2, Fig. 9, (coll. MGUH ) body whorl. At the points of intersection of the (leg. M.S. Nielsen). radial ribs and the two stronger spirals acute knobs

19 PL Coastal Glauconitic This = 57 occur, which are most prominent on the upper one to the teleoconch is not very distinct, but indicated by the disappearance of the spiral sculpture and a of these two spirals. adapical spiral rib continues across the radial ribs, without knobs being change from a light to a darker colour. formed. Inbetween the radial sculpture weak opisthocline growth lines are visible. rather Even the largest specimen has only one and a Paratypes Mogenstrup (coastal cliff): 1 specimen halfteleoconch whorls. y are moderately convex and separated by deep sutures. flat base 1 (leg./coll. MNO); specimen (leg./coll. JHR). Kirstinebjerg skov (beach exposure); 1 specimen (leg./coll. ISL). Remarks new species differs from the numerous other species of the genus Cerithiopsis is demarcated by a rib. labrum is spiral broken, and the inner lip is not well defined. very canal is rather short siphonal and wide, slight turned ly backwards. spiral sculpture on the teleoconch consists from the North Sea Basin by the glossy appea of two smooth spiral ribs, accompanying the upper rance of the whorls which is caused by the rather and the lower suture respectively, the shell's surface in between these spirals is smooth, except for narrow spiral and radial sculpture. protoconch of the new species about ten fine spiral furrows. radial sculpture resembles that of Cerithiopsis (s. lat.) serrula R. Janssen, 1978, but differs consists of about 12 strong almost orthocline ribs in the number of spiral threads (five to six in the per whorl which run between the two spiral ribs new species, but only two in C. serrula). Furthermore, the teleoconch sculpture is much coarser in C. serrula. re seems to be no closely related species in the North Sea Basin. Laiocochlis (Laiocochlis) supraoligocaenica without crossing them. Growth lines are visible, on the base they are prosocyrt. In the available juvenile specimens the base has no further sculpture. Paratypes specimen, (leg./coll. MNO). Mogenstrup (coastal cliff): 1 sp. nov. Discussion outline and sculpture of the pro PI. 2, Fig. 8 toconch agree with the description and illustration Locus typicus cliff at Mogenstrup, North in Gründel (1980: 251, fig. 35) of the protoconch of the genus Laiocochlis Dunker & Metzger, of Skive, Jutland, Denmark (Fig. 1). In NW Europe this genus is represented by four Stratum typicum sand, Brejning Clay species in the Lutetian of the Paris Basin (Coss Member of the Vejle Fjord Formation, Late Oli mann, 1889: 41, pi. 2, figs 2023), by the Pliocene gocène (Chattian B). Derivatio nominis species is named after its species L. woodi van Regteren Altena, 1954 ( Cerithium granosum Wood, 1848, non Borson) and occurrence in the Late Oligocene, because this by the Pliocene and Recent species L. sinistrata chronostratigraphic 'Upper' Oligocene. Holotype (leg. M.S. Nielsen). unit is often referred to as 2, Fig. 8, (coll. MGUH ) Only two were juvenile specimens collected. shell is turriculate, small and sinis (Nyst, 1835) (see Grundel, 1980: 251, erroneously cited as L. sinistria None Nyst). of these, however, seems to be related to the new for species. Except the Eocene L. which has smooth passyi (Deshayes), teleoconch whorls, all these species have a strong spiral ornament and a weaker or completely lack tral. last whorl present comprises about half ing radial ornament. the total shellheight. new species is the first representative of the relatively large protoconch consists of about genus Laiocochlis to be reported from the Late 2 x/i convex whorls which are separated by deep sutures. nucleus is and inflated. large sculpture of the larval shell consists of a group of Oligocene of the North Sea Basin. In the Oligocene (Rupelian) of the Aquitaine Basin (SW France) the species Laiocochlis lesbarritzensis four inequidistant spiral bands, lying alongside the Vergneau, 1963 occurs. A lot consisting of 23 adapical suture, and about 15 very delicate spiral specimens from the locality Gaas (Lagouarde) was grooves on the remaining part of the whorl. kindly sent on loan to me by Dr P. Lozouet upper spiral bands are separated by extremely narrow grooves. A radial sculpture, consisting of about 13 rather strong, almost orthocline ribs, is (Museum national d'histoire naturelle, Paris). French species differs from the present one by its much smaller protoconch and relatively smaller present after % protoconch whorl. transition nucleus. A radial is absent. On the teleo sculpture

20 R. R. At Coastal Glauconitic 58 conch three primary spirals are present which are stronger than the radial sculpture. Obviously L. riblets are united by a further spiral rib, below which the radial riblets are somewhat weaker, forming an angle of about 30 with the abapical lesbarritzensis is not closely related to L. supraoligocaenica. suture. number of riblets is about 24 on the penultimate protoconch whorl and it decreases to Triforis (Trituba) sorgenfreii sp. nov. PI. 2, Figs 57 about 20 on the last whorl. transition to the teleoconch is sharp and opisthocline. largest available specimen comprises 6'A teleoconch 1978b Triforis (Trituba) aff. raulini Cossmann & Peyrot 1921 Janssen, p. 171 (partim).?1978b Triforis (Trituba) aff. raulini Cossmann & Peyrot 1921 Janssen, p. 171, pi. 12, fig. 51 whorls which are low and separated by distinct sutures. se almost sutures are last straight. whorl equals less than % of the total shell height. aperture is rounded rectangular. siphonal Locus typicus cliff at Mogenstrup, North canal is broken in all available specimens, but of Skive, Jutland, Denmark (Fig. 1). seems to be directed obliquely backwards. Stratum typicum sand, Brejning Clay base is flat and provided with numerous prosocyrt Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). Derivatio nominis This species is namedin honour of the late Professor Dr Th. Sorgenfrei (Copenhagen), who contributed substantially to the knowledge of Tertiary molluscs. growth lines. spiral sculpture consists of two rather strong ribs, the adapical one being situated at about three fourths of the height of the whorl, the abapical one at about one fourth of this height. spirals are much narrower thantheir Between the interspaces. Holotype PI. 2, Fig. 5, (coll. MGUH20 038) (leg. two spirals the shell wall is concave. A third weak M.S. Nielsen). spiral is situated very close to the lower suture, but Mogenstrup a rather limited ma is usually covered by the next whorl. On the body terial consisting of five fragmentary shells and one whorl this spiral demarcates the base of the shell. juvenile individual with its protoconch preserved radial sculpture consists of about 1418 was collected. juvenile specimen is designated opisthocline ribs. At their points of intersection the holotype here, since it demonstrates the typical spiral and radial sculpture elements form solid knobs. se knobs are almost equal in strength on the two spirals. abapical knobs almost touch teleoconch sculpture on the two preserved teleoconch whorls. Additional material from Kirstinebjerg Skov, consisting of three fragmentary shells the suture. spiral demarcating the base is and three fragments, was also used for the follow smooth. Opisthocline growth lines are especially ing description. dextral shell is rather small, turriculate to well visible on the concave part of the whorl and on the base, which is otherwise smooth. cylindrical and slender, about four times as high as Paratypes Mogenstrup (coastal cliff): 1 defective wide. shell is very fragile because of the high numberof whorls, for which reason only fragmentary or juvenile shells could be collected. A reconstruction of the fragments suggest a shell height of up to 20 mm for adult specimens. protoconch comprises about six low and moderately convex whorls. se whorls are sutures. separated by deep nucleus is small. last protoconch whorl equals about one third of the total protoconch height. last five larval specimen; PI. 2, Fig. 6, (coll. MGUH ) (leg. K.I. Schnetler); 1 defective specimen (leg./coll. 1 ISL); defective specimen (leg./coll. MNO); 2 defective specimens (leg./coll. JHR). Kirstinebjerg Skov (beach exposure): 1 defective specimen, PI. 2, Fig. 7 MGUH (coll (leg. K.I. Schnetler); 2 defective 3 specimens, fragments (leg./coll. ISL). Freden: 1 specimen (coll. SMF ), mentioned in R. Janssen, 1978b, p. 171 (erroneously whorls are provided with a radial sculpture, con two specimens were mentioned there). sisting of numerous threadlike riblets which run Volpriehausen:? 2 specimens (one of which illustrated in R. Janssen, 1978b, pi. 12, fig. 51) (leg. from one suture to the other. On the second protoconch whorl these riblets are almost orthocline, but on the last protoconch whorl they become opisthocyrt. At the same time a spiral rib appears above A. von Koenen, coll. GeologischPalaontologisches Institut, Gottingen). KrefeldLinn (water well): 2 defective specimens the the abapical suture. adapical parts of the (leg./coll. WLH).

21 Schnetler, Mogenstrup 1 Since At Norre Schnetler, 59 Discussion protoconch sculpture of the new 171; two specimens were mentioned erroneously), species shows a similar development as in Triforis (Trituba) aff. bitubulatus Baudon, 1856 from the Hemmoorian of WinterswijkMiste kindly loaned to me by Dr Janssen. This specimen agrees with the Danish material and consequently it is referred to the new species. in Netherlands (see A.W. Janssen, 1984: 160, pi. 7, fig. 6), but it differs by having a higher number of radial riblets. Furthermore, in the new species the Cirsotrema (Opaliopsis) subglabrum sp. PI. 2, Fig. 10 nov. protoconch whorls have their greatest convexity about halfway between the sutures, whereas it is 1987 Cirsotrema (Opaliopsis) sp. in Schnetler & situated more abapically in the species from Beyer, p. 211, pi. 1, fig. 12 WinterswijkMiste. teleoconch whorls of the Locus typicus Vissing, claypit of Galten new species have a smooth peripheral spiral, in the Brickworks, Jutland, Denmark. species from Miste this spiral is provided with knobs. In other features the two rather are species similar. I have compared the Danish material with topotypical specimens of raulini Cossmann & Peyrot, 1921 from PeyrehoradePeyrere Triforis (Trituba) (France), kindly sent on loan to me Mr A.W. by Janssen (coll. ROM), and with two such topotypes in coll. ISL (leg. R. Janssen). protoconch in the French species consists of three whorls only. Its nucleus is relatively large, the whorls are highly Stratum typicum Member of the Brejning Clay = almost Vejle Fjord Formation, Late Oligocene (Chattian B). (L.) subglabrum Derivatio nominis smooth. Holotype in Schnetler & Beyer, 1987, pl. 1, fig. 12, MGUH (coll ). first of this my description species additional material has been collected, consisting of three defective shells and a protoconch from and Mogenstrup, a juvenile specimen with convex, especially so the second and the third, and its protoconch preserved from Kristinebjerg Skov. provided with a collabral sculpture. On the initial new material matches the specimen from protoconch whorl 12 rather weak orthocline ribs are present, which number increases to 16 on the following whorls. radial sculpture becomes stronger downwards, on the last half protoconch whorl the ribs become opisthocline. teleoconch of T. raulini differs in several respects. Danish species has relatively higher whorls and the two spirals are further apart. two rows of knobs are almost equal in strength in the Danish species, while in the French form the abapical knobs are stronger and also more pointed. Finally, the peripheral spiral of the Danish species is smooth, while the corresponding spiral in the French species has distinct knobs, causing a more undulated suture. Thus, the new Norre Vissing perfectly. largest specimen from consists of 3 A teleoconch Mogenstrup whorls, including the transition from the protoconch to the teleoconch. number of radial ribs is about 20 on the last whorl. A very fine spiral ornament, of consisting eight spiral bands, separated by delicate spiral grooves, is faintly discernible. For detailed a more description the reader is referred to the abovementioned paper. Paratypes (coastal cliff): 1 defective coll. specimen (Pl. 1, Fig. 13) (leg. J. Hillersborg; MGUH ); 2 defective specimens (leg./coll. 1 JHR); juvenile specimen (protoconch only) (leg./coll. MNO). Kirstinebjerg Skov (beach exposure) specimen (leg./coll. ISL). species demonstrates a closer relationship to the species from WinterswijkMiste than to the one Cirsotrema (? Opaliopsis) aff. koeneni from Peyrehorade. From the German Late Oligocene R. Janssen A.W.Janssen, 1967 PI. 2, Fig. 11 (1978b: 172, pi. 12, fig. 51) illustrated a specimen from Volpriehausen, indicated as Trifora (Trituba) aff. raulini Cossmann & Peyrot, This Mogenstrup a single specimen was found, of consisting two protoconch whorls specimen might be conspecific with T. sorgenfreii, and 4 3/4 teleoconch whorls. but the specimen was not yet available for study. I have compared a rather defective specimen (about two teleoconch whorls) from Freden (coll. shell is slender turriculate, about three times as high as wide. last whorl equals about one third of the total shell height. preserved SMF ), mentioned in R. Janssen (1978b: protoconch whorls are slightly convex and sepa

22 A I A 60 rated by deep sutures. y increase gradually in of very delicate, closeset spiral threads can be diameter. whorls are provided with about 30 noted. On the lower part of the whorls about ten opisthocline radial riblets. transition to the extremely fine spirals are discernible. spiral teleoconch appears to be sharp. rib demarcating the basal disc is visible in the teleoconch consists of 4 3/4 convex, some suture. what angular whorls which are separated by deep sutures. shell has a smooth basal disc, the radial sculpture consists of 1820 prominent, almost orthocline ribs which run across the aperture is rounded oval, the labrum is broken whorls from one suture to the other. se ribs are and the inner lip is concave. callus is thin. almost as wide as their interspaces. Inbetween the spiral sculpture is visible from the second ribs growth lines are visible. On the adapical part teleoconch whorl onwards. It consists of three of the whorls they cross the delicate spiral threads, spiral ribs, of which the adapical one is the resulting in a fine granulation (magnification strongest. This rib is situated at about two thirds of whorl three height. spirals are regularly spaced. A further, smooth demarcates spiral the x 50). Remarks shell has an overall Turbonillalike appearance because of the slender outline, the basal disc, it is partly covered by the subsequent radial sculpture and the weak spiral sculpture. volution, but visible on all teleoconch whorls. No Opalia (Pliciscala) pusilla (Philippi, 1843) (see R. secondary spirals are inserted. radial sculpture consists of about 16 ribs which are stronger than the spirals. y are almost as wide as their Janssen, 1978b, pi. 12, fig. 57) from the Lower Rhine area, F.R.G., has a rather similar radial sculpture, but no spiral on the base at the transi interspaces. On the first teleoconch whorl the tion from the base to the canal. Cirsotrema (Opaliop radial ribs are opisthocline, but they gradually sis) turbonillaeforme A.W. Janssen, 1967 is quite become almost orthocline on the younger whorls. similar in many respects, but differs by its lower At the points of intersection between the two number of radial ribs (14 instead of 1820 ribs per sculptural elements rounded knobs form, most whorl). prominently on the adapical spiral rib. Remarks Late Oligocene specimen de Cirsotrema (Opaliopsis) sp. 2. scribed here differs from the Miocene C. koeneni by PI. 2, Fig. 13 having three primary spirals instead of two. single protoconch was found which Cirsotrema (Opaliopsis) sp. 1 PI. 2, Fig. 12 differs from Cirsotrema (Opaliopsis) subglabra sp. nov. specimen comprises five moderately convex whorls, separated by deep sutures. nucleus is relatively large and inflated. last single specimen was collected at three whorls have a very fine radial sculpture, con Mogenstrup. protoconch sisting of numerous flexuous riblets (about 60 on and the first teleoconch whorls are missing and the aperture is very the penultimate whorl). In between this radial poorly preserved. shell is slender turriculate, sculpture numerous extremely fine spiral grooves about four times as high as are suggested. base of the shell is convex. wide. whorls expand regularly in diameter. specimen consists of 4 3/4 whorls which are separated by deep, aperture is rounded oval, the inner lip somewhat thickened. slightly undulating sutures. height of the body Remarks refer this larval shell to Cirsotrema whorl measures about one third of the total shell (Opaliopsis) height. flat basal disc is demarcated by a smooth spiral rib which is thickened in adapical on account of its similarity to the protoconchs of C. (O.) subglabrum sp. nov., as described above, and of C. (O.) turbonillaeforme A.W. direction in between the radial ribs. aperture is roundedoval, the lab rum and the abapical part of the aperture are broken off. A further spiral lies Janssen, 1967 from the Reinbekianof Dingden. In fact, it cannot be excluded that the present specimen does belong to the same species as Cir at the transition from the base to the canal. sotrema (Opaliopsis) sp. 1, the protoconch of which At about three quarters of their height the whorls show a faint angulation. Above this angulation, in the adapical part of the whorls, a number is unknown. This assumption seems to be supported by the very weak teleoconch sculpture of the lastnamed species.

23 A Tembrock, Rasmussen, I 61 Apiocypraea (Apiocypraea) cf. humbergi R. Janssen, 1978 PI. 3, Fig. 4ab protoconch is paucispiral and consists of two whorls, separated by deep sutures. nucleus is and vesicular. large On the last % whorl rather limited material, consisting of one one protoconch, juvenile shell with half a teleoconch whorl preserved and one isolated lab rum, is available. five orthocline radial ribs are present. boundary with the teleoconch is sharp and marked by a slightly flexuous radial rib. protoconch surface shows an extremely delicate microsculpture, consisting of about 20 furrows whorl. spiral per protoconch is conical and comprises about five moderately convex whorls which are separated This microsculpture is specimens only. visible on wellpreserved by rather shallow sutures. nucleus is relatively large and inflated, the following whorls are provided with a cancellated sculpture, produced by numerous fine opisthocline and prosocline threadlike riblets. transition to the teleoconch cannot be studied on the juvenile specimen. largest specimen available has 3 3/4 teleoconch whorls which are angular because of a ring of projecting above midwhorl. adapical part of the whorl is almost spines placed just flat, whereas the lower part is slightly convex. base of the body whorl is regularly constricted. sculpture on the preserved half teleoconch columella is slightly flexuous, the aperture is whorl consists of ribbonlike spiral ribs which are narrower than their interspaces. Secondary spirals roundedoval and constricted below to the long, narrow and straight siphonal canal. inner lip develop rapidly. radial sculpture consists of is simple, smooth and well defined; the labrum is prosocyrt riblets which are weaker than the spiral ribs. combination of the two sculptural somewhat thickened. spiral sculpture starts with three weak elements results in a reticulate ornament. radial soon sculpture disappears in apertural direction. isolated labrum lacks its adapical part. spirals, the upper one of which is the strongest. spirals are separated by much narrower interspaces. On the body whorl five spirals are present below the strong adapical one. outline of the labrum, its size, and also the number radial sculpture consists of laminate ribs and the shape of the teeth, agree very well with the description and illustration of Apiocypraea (A.) which are almost orthocline. On the first teleoconch whorl there are 15 such ribs, increasing to humbergi in R. Janssen (1978b: 197, pi. 14, fig. 18 to 20 on the body whorl. At the points of intersection with the spirals the ribs become crispate. 86). protoconch closely resembles the description and illustration of the species A. (A.) subphy sis On the strong upper spiral spines are formed that septemtr ionalis (Schilder, 1929) from the Hem project somewhat in adapical direction. On the moorian of WinterswijkMiste, Netherlands base a similar sculpture is present, but it becomes gradually weaker downwards. Inbetween the (A.W. Janssen, 1984: 193, pi. 82, fig. 2). I therefore refer the Danish material, with a query, humbergi. to A. radial sculpture fine, slightly flexuous growth lines are visible. Remarks compared the fairly rich material Trophonopsis (Pagodula) angustevaricata from Mogenstrup with specimens of the genus (Gripp, 1912) Trophonopsis from a number of localities. From the PI. 2, Fig. 14ab Danish Late Oligocene I studied material from Skanderborg (1 specimen, coll. ISL) and Brejning 1912 Murex angustevaricata nov.sp., Gripp, in Koch, Gripp & Franke, p. 22, fig Trophonopsis (Pagodula) semperi angustevaricata (Gripp, 1912) p. 314, pi. 5, fig Trophonopsis semperi (Koenen, 1872) p. coll. (2 specimens, ISL, and 2 coll. specimens, MNO). se lots seem to agree with the material from Mogenstrup with regard to the protoconch (worn on the two specimens from Brejning). 125 (partim, non von Koenen). number of radial ribs is lower on the Brejning specimens (10 to 12 ribs, but 18 on those from shell is slender and fusiform; its Mogenstrup). In addition, these ribs are height equals about 2 la times the width (estimated, because the siphonal canal is broken in opisthocline abapically. Thus, the Brejning material probably does not belong to Trophonopsis all specimens). angustevaricata, but seems to be closer to T. semperi

24 = Angistoma Glauconitic Ravn, Schnetler, Ravn, This 62 (von Koenen, 1872) (compare Tembrock, 1963, Derivatio nominis new species is named in pi. 5, fig. 5). From the Danish Late Miocene one specimen of honour of the late Dr J.P.J. Ravn, first to describe this species. who was the Trophonopsis semperi from Gram was studied (coll. Holotype 1907, pi. 5, fig. 14ab (coll. ISL). Its protoconch differs from the Mogenstrup MGUH 505). shell is large, fusiform and rather form by having a smaller nucleus, a slightly angular last whorl and a microsculpture consisting of thickwalled. height equals two times the prosocline radial threads and spirals that cross the diameter, the last whorl about three quarters of the radials at right angles (Rasmussen, 1968: 123). On total shellheight, and the aperture and the canal the specimens from Mogenstrup the microsculpture consists of spiral furrows only. On the teleo slightly more than half of the total shell height. adult shell may have up to eight whorls, but conch of the Gram specimen the angulation of the no complete specimen was found. On one defec whorl is situated on the middle of the whorl, and tive shell the protoconch is present, but it is rather there are only two spirals present. number of poorly preserved. It consists of two relatively large radial ribs is about 12 the shell from Gram and on whorls, of which the first one is flattened and the spines are projecting more adapically than in the Mogenstrup material. From Made (Danish domeshaped, while the second one is more convex. se whorls are separated by deep sutures, Gram Clay, Late Miocene) Rasmussen (1968: the nucleus is large and apparently vesicular. 124) mentioned a large specimen having only eight or nine radial ribs. Rasmussen (1968: 125) united T. semperi semperi (von Koenen, 1872) and T. semperi angustevaricata (Gripp, 1912) since the number of spirals variable in his material from Gram. In my opinion was T. second protoconch whorl is with a provided spiral ornament, consisting of about 11 weak spiral bands. transition to the teleoconch is distinct and marked by five closeset orthocline very growth lines. Immediately behind this boundary the radial ribs and the occur, number of spiral semperi and T. angustevaricata should be considered bands increases markedly on the first teleoconch two separate species because of the difference in whorl. protoconch and the considerably higher number of teleoconch consists of up to about seven radial ribs on the specimens from Mogenstrup. Tembrock (1963: 314) treated T. angustevaricata whorls which are moderately convex and separated by deep sutures. presence of radial ribs (see below) makes the suture line undulate on as a subspecies of T. semperi and stated that both occur in the 'Upper Miocene' of Liineburg, F.R.G. the first teleoconch whorls. This course of action does not comply with the aperture is oval and constricted into the presentday subspecies concept and seems to con welldefined siphonal canal which is relatively firm that T. semperi and T. angustevaricata are short and somewhat twisted backwards. separate species. labrum is strongly and regularly thickened, both specimens from Brejning seem to belong to T. semperi and thus T. and semperi T. angustevaricata are in the North Sea Basin from the Late present interiorly and exteriorly. Exteriorly the apertural margin is abruptly constricted. inner lip is well defined, and the columella is smooth, slightly Oligocene to the Late Miocene. thickened at the transition to the canal. A solid, rounded parietal tooth is present. transition to Searlesia ravni sp. nov the base with the canal is gradual and slightly PI. 2, Figs 16, 17ab concave. spiral of the teleoconch starts with sculpture 1907 Fusus aff. Koninckii Nyst p. 119, pi. 5, fig flat spiral bands which are separated by nar 14ab Angistoma brueckneri (Beyrich, 1856) in Schnetler & Beyer, p. 213 (partim, non Beyrich, non pi. 2, fig. 3 brueckneri danica subsp. nov.). row furrows. number of these spirals is constant, but on the body whorl a generation of secondary spirals develops. 34 adapical bands are weaker than the lower primary spiral Locus typicus Cilleborg (former claypit), ones. base and the canal have a similar spiral Jutland, Denmark. Stratum typicum clay, Late Oligocene ornament. radial sculpture consists of 16 radial ribs on (Chattian A). the first teleoconch whorl, decreasing to 1315 on

25 I 1 63 the last whorl. se ribs are about as wide as their and Fakkegrav specimens show a number of dif interspaces. On the younger whorls the radial ferences with as well as similarities to typical A. sculpture gradually becomes weaker, and on the brueckneri, for which reasons I treat this material as penultimate whorl it fades completely. Weak, al a geographical subspecies of A. brueckneri. most orthocline growth lines are visible. Below the Affinities with some other species of the 'Fusus' group adapical suture they are slightly curved backwards. Rupelian Fusus koninckii Nyst, 1845 (sensu Paratypes Mogenstrup (coastal cliff): 1 specimen Beyrich) has more angular and more convex (PI. 2, Fig. 16ab, (coll. MGUH , leg. S.B. whorls, and radial ribs on the adapical part of all 1 Andersen); specimen (PI. 2, Fig. 15) (coll. MGUH , leg. K.I. Schnetler); 1 specimen (leg./coll. ISL); 1 specimen (leg./coll. MNO). 1 Brejning (beach exposure); specimen (leg./coll. MNO). Discussion have recently discussed the Danish whorls. Its size, aperture and tooth parietal are similar to Searlesia ravni sp. nov., which seems to be closely related. In my opinion Fusus koninckii should also be considered to belong to the genus Searlesia, rather than to Euthria, as was suggested by R. of Euthria Janssen (1979a: 290). Species material of the genus Angistoma and pointed out have a concave subsutural depression without dis differences between the German A. brueckneri some (Beyrich, 1856) and the Danish material (Schnetler, in Schnetler & Beyer, 1987: More mate 213). tinct spiral sculpture, no parietal tooth and a canal. longer, narrow siphonal Angistoma from the Ger feldhausi (Beyrich, 1856) rial is available now, and it seems justified to man Middle Oligocene (NeustadtMagdeburg) separate the Danish material from Cilleborg, Brej has 1013 spirals, 34 denticles on the columella ning and Mogenstrup from A. brueckneri. and is smaller than Searlesia ravni sp. nov. specimens from Arhus, Norre Vissing and Fakke Searlesia mitgaui (von Koenen, 1867) from the grav Badehotel, however, should be treated as a geographical subspecies of A. brueckneri. In my German Late Oligocène (Söllingen) has radial ribs on all whorls and two generations of secondary opinion the new species is better referred to spirals that do not reach the strength of the Searlesia, in view of its protoconch, spiral sculpture and aperture. Searlesia ravni sp. nov. differs from A. typical brueckneri by a number of considerably higher primary ones. On the inner two weak denticles lip are present, that is, on the parietal wall and at the transition to the canal. For the material from Arhus, Norre Vissing and spiral bands which are rather equal in strength, by Fakkegrav Badehotel I here introduce the name the absence of knobs on the columella and the Angistoma brueckneri danica subsp. nov., and presence of a solid parietal tooth. whorls are designate the specimen illustrated by Schnetler (in more regularly convex and the shell attains a Schnetler & Beyer, 1987, pl. 2, fig. 3; coll. MGUH larger size. Finally the protoconchs are very different: A. brueckneri has a relatively small, ) holotype. following paratypes are available: Arhus (former railway excavation) 1 multispiral protoconch, the larval shell is large whorls. whereas in Searlesia ravni and consists of only two specimen, coll. D.G.U. (leg. P. Harder); Norre Vissing (claypit of Galten Brickworks) 1 specimen (leg./coll. ISL), 1 specimen (coll. MNO, specimens from Arhus (Harder, 1913, p. 78, pi. 6, Figs 910), from Norre Vissing leg. K.I. Schnetler) 2 specimens (leg./coll. MNO); Fakkegrav Badehotel (coastal cliff) specimen (Schnetler, in Schnetler & Beyer, 1987, p. 213, pi. (leg./coll. MNO). 2, Fig. 3) and from Fakkegrav Badehotel (coll. MNO) agree with the German species A. brueckneri Buccinidae gen. et sp. indet. in size and general outline, and in the absence of TextFig. 6 a parietal tooth. Unfortunately, no Danish specimen has its protoconch preserved. y have Only two defective juvenile slightly angular whorls with about 16 spiral bands of almost equal strength and two denticles on the specimens are available (leg. M.S. In the Nielsen). better one of these the and the first protoconch columella, whereas the German specimens have teleoconch whorl are preserved. second nonangular whorls with about 6 to 9 spiral bands, specimen, demonstrating a part of the second somewhat variable in strength, and four denticles teleoconch whorl, is very defective because of a on the columella. Thus, the Arhus, Norre Vissing split parallel to the axis of the shell.

26 One 64 protoconch comprises 1 % whorls, separated by a deep suture. nucleus is large, from my private collection. All these specimens are fullgrown and therefore their apical parts are globular and slightly vesicular. On the last half worn. Mogenstrup material seems to agree protoconch whorl fine, irregularly distributed, almost orthocline growthlines are present, crossed by some ten very delicate spiral furrows, which are best with L. rarus with regard to the convexity of the whorls and the number of primary spirals. On the Brejning specimen the number of spirals is 12, merely suggested. se furrows are most promi while the specimens of L. ventrosus from Gram nent on the upper part of the whorl. transition have only 7 primary spirals. very juvenile to the teleoconch is gradual, indicated by about 15 state of the material, however, prevents a final in closeset growthlines. terpretation. Dr R. Janssen (SMF) kindly examined the illustrated specimen. In his opinion it cannot be referred to Liomesus rarus (pers. comm., 1990), which differs by a much larger protoconch (diameter 2.5 mm), which is flat and domelike. Furthermore the spiral sculpture of that species is much coarser. Latirus (Pseudolatirus) sp. Pl. 3, Fig. 2 juvenile specimen, consisting of the protoconch and one teleoconch whorl, was collected at Mogenstrup (leg./coll. JHR). In addition, four juvenile specimens are known from Kirstinebjerg Skov (leg./coll. ISL). shell is fusiform, about three times as high as wide. height of the whorl is about six body tenths of the total shell height. protoconch is conical, it comprises 44 moderately convex whorls which are separated by deep sutures. nucleus is small. whorls are Fig. 6. Buccinidae sp., Mogenstrup; a: frontal view, b: apical view. Coll. MGUH (A.W. Janssen del., 1990; bar length is 1 mm). smooth and glossy, except for the last half whorl, on which opisthocyrt growth lines are rather distinct and three rather weak spiral ribs appear. one of these is the weakest. A further adapical preserved teleoconch whorls are moderate fine spiral is present on all protoconch whorls in ly convex and separated by a deep, almost canali the abapical suture. Immediately in front of the culate suture. aperture is ovoid, abapically boundary with the teleoconch 2 or 3 slightly passing into a short, rather wide and oblique siphonal canal. opisthocline radial ribs are present. to the teleoconch is gradual. transition Only one convex teleoconch whorl is present in spiral sculpture consists of about ten primary spiral bands, which are separated by narrower the available specimens. height furrows. adapical spiral is somewhat more prominent. A radial sculpture is absent, except for weak orthocline growthlines. of the rounded oval aperture is about halfthe total shell height. siphonal canal is long and rather narrow. inner lip is smooth and not very strongly protrud Discussion features of protoconch and ing. A weak oblique fold is present on the col teleoconch agree rather well with the description and illustration of Liomesus rarus (Beyrich) (see umella. of the teleoconch consists of spiral sculpture Beyrich, 1856: 250, pi. 17, figs 6ab). I compared three rather narrow spiral bands which are the the present materialwith a specimen of this species continuations of the spirals on the protoconch. from Brejning and with two specimens of L. ventrosus (Beyrich, 1856) from Gram, both samples Below the adapical suture a weak spiral band appears on a quarter of a whorl. radial orna

27 Coastal Glauconitic This One 65 ment comprises nine ribs strong on the first whorl which distinctly weaken downwards, the transition to the canal. base and the siphonal canal on are provided with a similar spiral sculpture. Discussion limited Danish material consists separated by a rather distinct spiral furrow; four to five very weak, merely indicated bands are spiral present above the two lower ones. transition to the teleoconch is difficult to detect, but suggested by closeset growth lines. On the two specimens found the teleoconch consists of up to 2 Vi whorls which are slightly convex, having an almost flat part below the adapical of juvenile specimens only, for which reason a specific identification is impossible. Yet, the protoconch characteristics differ from those of the Miocene North Sea Basin species Latirus (Pseudolatirus) suture. whorls are separated by deep sutures. rothi (Beyrich, 1856) by a number of features. base is strongly constricted below the peri protoconch of the Danish specimens has a smaller phery. aperture is roundedoval and rather apical angle and only two or three radial ribs on its terminal part while in L. rothi the complete ultimate abruptly constricted at the transition to a rather short, wide and open siphonal canal. labrum protoconch whorl is provided with radial ribs. Fur is broken, the inner lip and the left margin of the thermore, the columellarfold seems to be weaker in canal are Sshaped. callus is thin, lying deeper the Danish specimens. Sorgenfrei (1958: 232, pi. and thus showing traces of resorption of calcareous matter. 50, fig. 157) described and illustrated the protoconch of L. rothi from the Arnum Formation, and he also found the radial ornament of the protoconch spiral sculpture starts with six to seven primary spiral bands which are separated by to be restricted to the last one third or half whorl. rather deep furrows. adapical spiral band is separated from the second one by a somewhat This feature was also noted in a sample of 20 juvenile specimens of L. rothi from a water well at Ny Lifstrupvej (South ofvarde, Denmark). Thus, it might well be possible that a more extensive material from Denmark allows ataxonomic subdivision. deeper furrow and develops into a sutural band on the second teleoconch whorl. spiral bands number two and seven are weaker than the others. number of spirals increases by one on each of the teleoconch whorls because of further spirals ap Clavatula mogenstrupensis sp. nov. pearing from below the abapical suture. No secon Locus typicus PI. 3, Fig. 5 cliff at Mogenstrup, North are inserted. base and the canal dary spirals are provided with a similar spiral sculpture. radial ribs are On the first opisthocyrt. of Skive, Jutland, Denmark (Fig. 1). teleoconch whorl they run from one suture to the Stratum typicum sand, Brejning Clay other, but on the second whorl they become weaker, especially so below the adapical suture Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). Derivatio nominis type locality. species is named after the where the sutural band is radial developed. sculpture is rather similar to that of the wellknown species Fusiturris duchastelii (Nyst, 1836). Holotype PI. 3, Fig. 5 (coll. MGUH ) (leg. number of ribs is about 18 on the last whorl. K.I. Schnetler). radial ribs follow the course of the growth shell is rather small, slender lines which are visible between the radial ribs, fusiform, about two and a half times as high as with the deepest point of the sinus situated be wide. height of the last whorl equals slightly tween spiral band number three and four, as than half the total height, the height of the aper counted from the adapical suture. ture is about two fifths of the total height. Paratypes specimen from the type locality protoconch comprises about 5 % slightly to moderately convex whorls which are separated by (leg./coll. Discussion MNO). deep sutures. nucleus is small and slightly heterostrophic. protoconch whorls are smooth, new species shows some resemblance with C. chattica R. Janssen, 1979, but differs by having more convex protoconch whorls except for the last one which is provided with about 18 opisthocyrt radial ribs. last protoconch whorl is more convex and has a weak spiral and stronger opisthocyrt radial ribs on the last protoconch whorl. On the teleoconch the radial sculpture is stronger. Finally, the number of primary ornament, consisting of two weak spiral bands above the abapical suture. se bands are spirals is six to seven in the new species, whereas C. chattica has only five primary spirals.

28 A Beach PI. This 66 Microdrillia (Andersondrillia) subgen. nov. Stratum typicum Brejning Clay Member of the Vejle Fjord Formation, Late Oligocene (Chattian Diagnosis subgenus of Microdrillia showing the B). same protoconch features as the genus. On the te Derivatio nominis species is named after its leoconch the sickleshaped riblets on the subsutural flat or concave of the whorl absent. part are On the lower part of the whorls the combinationof spiral and radial ornament results in more or less prominent knobs, especially so on the carina. Type species Microdrillia (Andersondrillia) grippi Anderson, 1964 (Miocene, North Sea Basin). Discussion type locality. Holotype (leg. M. S. Nielsen). 3, Fig. 8ab, (coll. MGUH ) shell is rather small and slender fusiform, about 2.8 times higher than wide. height of the last whorl equals about two thirds of the total shell the height, height of the aperture is about half the total shell height. largest new subgenus resembles the genus Asthenotoma Harris & Burrows, 1891 and especially the species A. obtusangula (Brocchi, 1814) in specimen consists of about seven 3% belong to the protoconch. whorls of which outline general and of the sculpture teleoconch, as pointed out by A.W. Janssen (1984: 299). protoconch, however, with Microdrillia agrees entirely protoconch is broadly conical with convex whorls which are separated by deep sutures. nucleus is small, slightly inflated and apparently s.str. From typical representatives of Microdrillia s.str., such as M. bicingulata (Sandberger, 1860), M. smooth. Succeeding protoconch whorls are provided with a radial sculpture which consists of fine, serratula (Bcllardi, 1878) and M. teretiaeformis A.W. opisthocyrt ribs, numbering about twenty on the Janssen, 1972, the new subgenus differs markedly by the complete absence of the sickleshaped riblets first whorl. ribs run backwards below the adapical suture and form an opisthocyrt curve across in the subsutural excavated part of the whorl. se the whorl. Near the abapical suture the ribs bend species generally are also more slender and their spiral ornament is more prominent. forwards and disappear. radial sculpture gradually increases in strength; on the last protoconch new subgenus is represented in the North whorl the ribs are more widely spaced and prominent. On the terminal half protoconch whorl about Sea Basin by two species, viz. Microdrillia (Andersondrillia) grippi Anderson, seven such ribs below the are present. Immediately 1964 and M. (A.) brejningensis sp. Nov. Judging from the description adapical suture lies a fine spiral rib; on the middle and illustration in Peyrot (1932: 71, pi. 7, figs 67) the spe part of the whorl a few spiral furrows are indicated. cies M. venusta (Peyrot, 1932), described from Late On the abapical part of the protoconch whorls an sediments at Oligocene Peyrere (SW France), probably also belongs to the new A. subgenus. W. Janssen (1984: 299) considered this taxon to be closely related to, or even conspecific with M. grippi. extremely fine sculpture is present, consisting of thinlines crossing the radial ribs, exclusively visible in in front wellpreserved specimens. Immediately of the transition to the teleoconch several flexuous Microdrillia speyeri (Koch & Wiechmann, 1872) ribs are present. has a teleoconch rather similar to Microdrillia s.str., largest specimen has 3 3/4 teleoconch whorls but the paucispiral protoconch excludes a close which are carinate due to a prominent, central relationship. According to A.W. Janssen (pers. comm., 1990) that should better be species referred to another, possibly new genus. spiral rib. This rib divides the whorl into a distinctly concave adapical part and an almost vertical base abapical part. is slightly convex and Derivatio nominis new subgenus is named rather abruptly constricted to a siphonal canal of afterprofessor H. J. Anderson (Marburg, F.R.G.) for his substantial contributions to the knowledge of Tertiary molluscs. medium length. aperture is elongatedoval with a gradual transition into the canal. lab rum is broken on all available specimens. inner lip is smooth, the callus is thin and excavated, Microdrillia (Andersondrillia) brejningensis thus showing traces of resorption of calcareous sp. nov. matter. PI. 3, Figs 9ab, 90 spiral sculpture starts with three primary Locus typicus exposure at Brejning, South spirals, of which the adapical one is the strongest. This accentuated spiral rib, or better, carina, is of Vejle Fjord, Jutland, Denmark. situated at midwhorl, and the two other spiral ribs

29 This PI. Coastal Glauconitic This 67 are regularly distributed on the abapical part of Microdrillia (Microdrillia) ingerae sp. nov. the whorl. Immediately above the abapical suture PI. 3, Figs 7ab a weak spiral rib is seen from the second teleoconch whorl onwards. spirals are distinctly Locus typicus cliffat Mogenstrup, North narrower than their interspaces. A large specimen of Skive, Jutland, Denmark (Fig. 1). Stratum typicum sand, Brejning Clay from Mogenstrup shows an aberrant spiral sculpture in that the third spiral from above is completely absent. In all other features the shell agrees Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). completely with the remaining material. In the Derivatio nominis species is named after my concave subsutural part of the whorl four spiral wife Inger Lund Schnetler. bands are present on the first teleoconch whorl, Holotype 3, Fig. 6ab, (coll. MGUH ) separated by narrow furrows, but they soon disappear. On the body whorl only one very weak spiral is indicated immediately above the adapical spiral (leg. K.I. Schnetler). shell is rather small, fusiform, about 2.4 times higher than wide, rather solid, the rib. No secondary spirals are inserted. base is height of the aperture equals slightly less than half provided with a similar sculpture to the end of the the total shell height. canal, a further seven spirals are present. protoconch is broadly conical and consists radial sculpture consists of 1518 ribs which of five moderately convex whorls which are are narrower than their se ribs interspaces. are almost orthocline and visible below the only carina. At the points of intersection of and spiral radial sculpture knobs are formed, most prominently on the carina. growth lines are especially well visible above the carina, where radial sculpture is absent. rather sinus is deep separated by deep sutures. nucleus is small and inflated. initial one and a half protoconch whorls are smooth, the following ones are provided with a weak rib above spiral the suture abapical and with numerous narrow radial ribs which gradually increase in Near strength. the adapical suture these ribs run strongly backwards, but they situated immediately above the carina. On the become opisthocyrt on the last protoconch whorl base the growth lines form a very flat prosocyrt and here about eight fine spiral grooves curve. are indicated. An oblique microsculpture, consisting of Paratypes Brejning (beach exposure): 3 specimens (leg./coll. MNO); 1 specimen (leg. M.S. Nielsen; coll.isl). Mogenstrup (coastal cliff): 1 specimen, PI. 3, Fig. 9, (coll. MGUH ) (leg. K.I. Schnetler); 1 specimen (leg./coll. LJL); 5 specimens (leg./coll. very fine divergent lines crossing the radial ribs at more or less right angles is present in between the ribs (preserved on one shell only). In very juvenile specimens it can be seen that the base of the protoconch is demarcated by a strong, smooth spiral rib followed by a weaker rib below. spiral On the ISL); 4 specimens (leg./coll. MNO); 1 (leg./coll. JHR). specimen base of the protoconch the growth lines form a prosocline curve. transition to the teleoconch is Discussion species resembles Microdrillia distinct but not very sharp. grippi Anderson, 1964, but differs by a number of teleoconch of the largest specimen consists of three whorls which are separated by deep sutures. body whorl equals about two thirds features. new species has a more slender protoconch with the radial sculpture starting immediately after the nucleus. teleoconch has a distinct concave adapical part of the whorl without of the total shell height. whorls are divided into a slightly concave adapical part and an almost spiral sculpture. spirals below the carina are ilat abapical part by a carina which lies at mid narrower and there is no sutural spiral band. whorl. base is constricted to a rather short radial sculpture consists of almost orthoclinic ribs, canal. callus is thin and the columella is while M. grippi has prosocline ribs. sinus in smooth, showing traces of resorption of calcareous the growth lines of the new species is situated below the middle of the concave part of the whorl, while in M. grippi the sinus lies at midwhorl. matter. aperture is oval and narrows gradually into the siphonal canal. spiral sculpture starts with a strong carina shell has a glossier appearance because of the on the middleof the whorl. Immediately below the sharper sculpture, the weak radial sculpture and adapical suture a relatively broad sutural band is the absence of secondary spirals. situated and above the abapical suture a weak

30 PI. Coastal Glauconitic 68 spiral rib is visible. From the second teleoconch whorl onwards the carina consists of two closeset spirals. Both the sutural band and the spiral rib Pleurotomella (Pleurotomella) rasmusseni sp. nov. PI. 3, Figs llab below the carina gradually increase in strength. A secondary spiral rib develops in the centre of the Locus typicus cliff at Mogenstrup, North space between the carina and the adapical spiral. of Skive, Jutland, Denmark (Fig. 1). This secondary spiral soon becomes stronger than Stratum typicum sand, Brejning Clay the rib. On the base and the canal abapical spiral a further eleven spirals are present. A generation of secondary spirals develops on the base and on the neck of the canal. radial sculpture is rather From complex. the carina, on which they form accentuated Member of the Vejle Fjord Formation, Late Oligocène (Chattian B). Derivatio nominis species is named in honour of State Dr Geologist Leif Banke Rasmussen, Geological Survey of Denmark, Copenhagen, who has contributed substantially to the knowledge of knobs, thickened opisthocline ribs (following the the molluscs and the stratigraphy of the Danish direction of the growth lines) are present, quickly Neogene. diminishing in strength downwards and disap Holotype 3, Fig. loab, (coll. MGUH 20 pearing already near the second spiral below the 056) (leg. K.I. Schnetler). carina. ir number increases from about 18 on shell is rather small and thin the first teleoconch whorl to 24 to 26 on the body walled, subfusiform, about two and a half times whorl. growthlines are distinct, especially so higher than wide. height of the last whorl on the concave subsutural part of the whorls, equals three fifths of the total shell height. where they form numerous sharp and regularly largest specimen has about seven whorls, of which spaced, sickleshaped riblets (curved backwards). Below the carina are they curved in an anterior direction and, gradually becoming weaker, they continue downwards in a flat prosocyrt curve. 4% belong to the protoconch. This larval shell is broadly conical with convex whorls separated by deep sutures. nucleus is small, slightly heterostrophic and provided with an extremely Paratypes Mogenstrup (coastal cliff): 4 juvenile fine microgranulation arranged in eight spirals. specimens (leg./coll. ISL); 1 juvenile specimen last three whorls of the protoconch have a (leg./coll. JL). 2 specimens (leg./coll. MNO); 2 cancellate ornament on the abapical two third of juvenile specimens (leg./coll. JHR). the whorl, while the abapical third of the whorl has Kirstinebjerg Skov (beach exposure): 2 collabral riblets only. On the last protoconch specimens (leg./coll. ISL). whorl carina a develops on the middleof the whorl Discussion new species is obviously closely and immediately in front of the transition to the related to Microdrilliaserratula (Bellardi, 1878) but teleoconch a spiral rib appears on both sides of the differs by a number of features. M. ingerae has a carina. On the last % whorl numerous higher number of radial ribs on the protoconch. opisthocline growth lines are sutural band more strongly developed, the carina is less sharp and the two spirals below the visible. transition to the teleoconch is sharp but not very prominent. diagonal sculpture fades away and a carina are rather weak. In M. serratula the carina is sharp and consists of one spiral only. More fine granulation appears at the same time (magnification x 50). over, this species has a higher number of spiral largest specimen has 2 3/4 teleoconch ribs on the base and its general outline is more whorls which are very convex and separated by slender. deep, undulating sutures. Below the adapical In Microdrillia bicingulata (Sandberger, 1860) the suture the whorls are slightly concave. last two spirals below the carina are prominent and whorl comprises about three fifths of the total shell more widely spaced. Its radial sculpture is much height. base is convex and abruptly con weaker. M. teretiaeformis A.W. Janssen, 1972 has stricted into a rather short siphonal canal. a smooth carina consisting of only one spiral. aperture is roundedoval. labrum is broken Microdrillia speyeri (Koch & Wiechmann, 1872), on all specimens and the columella is smooth with which is also present at Mogenstrup, has a paucispiral protoconch. a welldefined inner lip. callus is thin, showing traces of resorption of calcareous matter.

31 This Fakkegrav This 69 spiral sculpture starts with three primary spirals which are continuations of the carina and the two spiral ribs on the last 3 A whorl of the protoconch. adapical spiral rib remains weak on all concave of the whorls is part more excavated on the new species, and finally the granulation is weaker than in P. mioweberi. Pleurotomella sp. from the German Late Oligocene (R. Janssen, 1979a: whorls, whereas the middle one, being the continuation of the carina, remains the strongest. 329, pi. 18, fig. 77) differs by the absence of secondary spirals and by its more angular whorls. abapical spiral increases in strength but remains weaker than the middle one. Immediately above Rimosodaphnella lappanni sp. nov. PI. 3, Fig. 13ab the abapical suture a further weak spiral rib appears on a quarter of a whorl. In the upper concave part of the whorl four narrow spirals are present from the first teleoconch whorl onwards. ir number increases to seven on the whorl. In body between the primary spirals a generation of secondary spirals develops from the second teleoconch Locus typicus Badehotel (coastal cliff), North of Vejle Fjord, Jutland, Denmark Stratum typicum Member of the Brejning Clay Vejle Fjord Formation, Late Oligocene (Chattian B). Derivatio nominis new species is named after whorl onwards. base and the canal are provided with a similar spiral ornament. my friend, Mr. Werner Lappann, Heiligenhaus Isenbügel, F.R.G. radial sculpture consists of nine to ten slightly opisthocline ribs separated by about equally wide interspaces. se ribs are almost invisible on the concave part of the whorls but prominent on Holotype PI. 3, Fig. 12ab; MGUH (coll ) (leg. M.S. Nielsen). shell is rather small and the convex part. At the points of intersection of fusiform, about 2.5 times higher than wide. last whorl comprises three fifths, the aperture slightly less than half the total shell height. protoconch is highconical and multispiral, spiral and radial ornament elongated knobs form. radial ribs on the base. gradually disappear growth lines are very prominent in the concave part of the whorls where together with the consisting of 3 Zi 4 moderately convex whorls spiral ribs they form a regular reticulate pattern. which are separated by deep sutures. whorls Downwards they decrease in strength. On the base have their greatest width abapically. nucleus is small and apparently smooth, while the last three they form a very shallow prosocyrt curve. entire teleoconch has a fine granulation between the whorls are provided with a diagonal sculpture. On sculptural elements. sinus is rather shallow the last half whorl of the protoconch a carina is and has its deepest point slightly of the concave part of the whorl. below the middle present, lying somewhat below the middle of the whorl. On the last % whorl before the transition to Paratypes (leg./coll. ISL). Mogenstrup (coastal cliff): 1 specimen the teleoconch a weak spiral rib develops between the carina and the abapical suture. in Immediately Jensgard (coastal cliff): 2 specimens (leg./coll. front of the transition a further weak spiral occurs ISL); 1 specimen (leg./coll. MNO). above the carina. se three spirals continue as Discussion new species is closely related to the primary spirals on the teleoconch. transi P. mioweberi (Nordsieck, 1972), as described and illustrated by A.W. Janssen (1984: 325, pi. 74, fig. 10; pi. 82, fig 5). I have compared the new species tion to the teleoconch is sharp, sigmoidal rib. teleoconch consists of 3 demarcated by a 3/4 whorls which with materialof the latter species from Gram (Late Miocene, coll. ISL) and Miste (Middle Miocene; have a concave adapical and a convex lower part. whorls are separated by deep sutures. coll. ISL). A number of differences seem to be constant. protoconch of the new species consists of base of the shell is convex, but rather abruptly constricted to a relatively long siphonal canal. 4 Yi whorls, whereas 5 such whorls are present in callus is thin and the inner lip is not very sharply P. mioweberi. protoconch of P. rasmusseni is demarcated (which could indicate that the avail more broadly conical. sculpture in the concave adapical part of the whorl is much more prominent able specimens are not yet fullygrown). aperture is oval, rather suddenly narrowed at the in the new species and the sinus of the growth lines is situated more abapically than in P. mioweberi. transition to the siphonal broken. canal. lab rum is

32 PI. At Water 70 spiral sculpture starts with three spirals on continues as the second spiral. Furthermore, the German species has a somewhat more thickset outline, a more broadly conical protoconch, a the convex part of the whorl which are continuations of the carina and the two spirals on the protoconch. On a quarter of a whorl a further, rather wider concave zone, a less sharp sculpture and is weak spiral rib appears immediately above the generally larger. abapical suture. second spiral from above is situated on the middle of the whorl. second? Actaeopyramis (s. lat.) and third spirals are stronger than the others. On pseudopunctata sp. nov. the second teleoconch whorl secondary spirals are PI. 3, Figs 1415 inserted between the primary ones, and on the younger whorls two further spiral generations are Locus typicus well at KrefeldLinn, present. Ultimately, the spirals become nearly equal in strength. base and the canal are furnished with a similar sculpture. All spirals are NordrheinWestfalen, F.R.G. Stratum typicum Grafenberg Sands, Late Oligocène (Chattian A or B). threadlike and narrower than their interspaces. Derivatio nominis species is named after its On the columella the spirals are visible below the very distinctive sculpture (pseudopunctata = falsely callus. On the concave upper part of the whorls punctate). no spirals are present. radial sculpture is restricted to the convex Holotype 3, Figs. 15 (coll. MGUH ) (leg. F. von der Flocht). part of the whorls. It starts on the first teleoconch Mogenstrup six juvenile speci whorl with about 14 opisthocline ribs which are mens were collected. An adult specimen from about as wide as their interspaces. number of ribs is constant on all the whorls. On the base of KrefeldLinn is designated holotype. shell is very small and rather thickwalled, the body whorl the radial sculpture gradually ovoidconical. In juvenile specimens the height disappears downwards. At the points of intersection of spiral and radial ornament small, elongate equals about 1.3 times the width. In the presumably adult holotype the shell height is about 1.9 knobs form, most prominently on times the width. height of the aperture equals the first teleoconch whorl. On the concave of the adapical part whorls the growth lines are accentuated and form sickleshaped riblets with a backward curvature. about one third of the total shell adult height. specimen comprises about four whorls, one of which belongs to the heterostrophic protoconch, On the convex part of the whorls they curve for of which only the terminal 3/4 whorl is visible, wards to opisthocline, in the same direction as the projecting only slightly above the teleoconch. radial sculpture. An extremely fine granulation is between present the elements all over sculptural the teleoconch, especially well visible (magnification x 50) on the concave part of the whorls. Paratypes transition to the teleoconch is sharp and slightly prosocline. three teleoconch whorls are slightly to moderately convex and separated by deep, somewhat canaliculate sutures. body whorl equals Mogenstrup (coastal cliff): 15 specimens (leg./coll. ISL); 13 specimens (leg./coll. about 6/10 of the total shell height. base is MNO); 1 specimen (leg. ISL/coll. WLH); 1 convex, the umbilicus is rather narrow. aper specimen (leg. ISL/coll. SMF ); 1 ture is oval, with a regularly rounded lower ISL/coll. RGH specimen (leg ). Kirstinebjerg Skov (beach exposure): 1 specimen (leg./coll. MNO). Discussion new species differs from Rimosodaphnella sp. [described and illustrated sub nomina labrum is with margin. sharpedged a transition into the columellar gradual part. columella is concave and bears a rather weak, oblique fold the umbilicus. opposite spiral sculpture starts after immediately Philbertia sinuosula Sorgenfrei, 1958 by Anderson the boundary with the protoconch and consists of (1964: 312, pi. 42, figs 268, 268a) nine to ten ribbonlike spirals which are separated from the Miocene of Dingden, F.R.G.] by the development of by narrower furrows. A weak subsutural ramp is the primary spirals. In the species described by demarcated by the adapical spiral band; on this Anderson, the carina on the last protoconch whorl develops into the uppermost primary spiral of the ramp two very fine spiral furrows are visible. base of the shell has a similar spiral sculpture. teleoconch, whereas in the new species this carina radial sculpture consists of about 30 opis

33 I 71 thocline, flat riblets which are weaker than the spiral bands. combination of the two is wide and open, very much shaped like the concave apical side of the shell. base of the sculptural elements results in a pattern punctate appearance. depressions of almost nucleus is visible in the umbilicus. spiral sculpture consists of barely visible are elliptical, having their greatest diameter in the direction of the spirals. growth lines are very fine furrows. On the upper side of the whorl one furrow can be seen, on the periphery three, and on and can be observed few on a specimens only. y are almost orthocline and appear to have a very flat sinus under the adapical suture. Paratypes Mogenstrup (coastal cliff): 1 specimen (PI. 3, Fig. 14) (coll. MGUH , M.S. leg. 4 Nielsen); specimens (leg./coll. MNO); 1 the base two. growth lines sometimes visible as are sigmoidal lines, especially so on the periphery. COMPARISON WITH OTHER LATE OLIGOCENE FAUNAS (AUTHOR: SCHNETLER) specimen (leg. M.S. Nielsen, coll. ISL). Danish Late Oligocene molluscan faunas were Discussion refer the new species to the genus published by Ravn (1907, 1909), Harder (1913), Actaeopyramis (s. lat.) with some hesitation, since Madsen (1918), Eriksen (1937) and Schnetler & the sculpture is somewhat untypical. In Actaeopyramis the spirals are smooth and and in projecting, the interspaces the radial lamellae give rise to a Beyer (1987). se faunas are characterised by a high frequency of species like Drepanocheilus (Arrhoges) speciosus, Polinices helicinus and Fusiturris punctation. Moreover, the growth lines are prosocline. general outline of the new species corresponds well to the genus Actaeopyramis, so do the shape of the protoconch and the aperture, but it closely resembles the species referred to as Pyramidellidae indet. and described from Norre in: Schnetler Vissing (Schnetler, & 1987: Beyer, 220, pi. 2, From this form the figs 1617). new species differs by its weaker subsutural ramp, less convex whorls and a less regular punctate sculpture. In all other features the two species appear closely related. y should be probably united in a new genus. Introduction of such a new genus is deferred to another occasion. Gastropoda inc. sed. TextFig. 7 Only a single (? juvenile) specimen is available (leg./coll. M.S. Nielsen). shell is very small and discoidal, 2.8 times wider than high. protoconch consists of 1 % whorls which are separated by a distinct suture. nucleus is small. transition to the teleoconch is sharp, demarcated by a distinct, slightly flexuous rib. teleoconch consists of somewhat more than 1 % whorls, slowly expanding in diameter and circular in transverse section, separated by a distinct suture. upper junction of the whorls lies on the periphery of the preceding whorl, the lower one touches below the periphery, which results in concave and lower sides of the shell. upper transition into the base is gradual. aperture is Fig. 7. Gastropoda inc. sed., Mogenstrup; a: apical view, b: frontal view, umbilical c: view, d: lateral view. Coll. MGUH (A.W. Janssen del., 1990; subcircular, the labrum is broken. umbilicus bar length is 1 mm.

34 72 duchastelii. Other very common species are Nucula comta, Limopsis aurita, aff. kickxi and Cyclocardia Gemmula spp. In to comparison the Arhus fauna (Harder, 1913) the following species are absent at Mogenstrup: [Brejning, Fakkegrav Badehotel, Bogeskov, Kirstinebjerg Skov, and Jensgârd Skanderborg or even (glacial flow)], absent from other Danish Late Oligocene localities (e.g. Arhus, Cilleborg and Norre Vissing). Thus, these rare species point out certain affinities with the faunas from the Vejle Fjord area, rather than with the faunas from Nucula harderi Schnetler, in Schnetler & Beyer, 1987 Arhus, Cilleborg and Norre Vissing. When compared to the German Late Oligocene Emarginula punctulata Philippi, 1843 faunas the Mogenstrup association shows affinities Unitas granulata (Nyst, 1845) Conomitra ravni (Harder, 1913) with those from the Lower Rhine area, Glimmerode, Freden and the Sternberger Microdrillia bicingulata (Sandberger, 1860). In comparison to the recently described fauna Gestein. Associations described from Sollingen, Niederkaufungen, Ahnetal and Doberg, however, appear to be different. from Norre Vissing (Schnetler & Beyer, 1987) the fauna from the Lower Rhine area has 110 following characteristic species are absent: species in common with Mogenstrup, among which are the following characteristic species: Nucula harderi Schnetler, in Schnetler & 1987 Beyer, Solariella spp. Unitas granulata Pleurotomoides naumanni (Speyer, 1867). Limopsis aurita Limopsis vonderhochti Lepetella compressiuscula Cirsope multicingulata Cirsotrema crispatum faunas from Norre Vissing and Mogenstrup have 97 species in common, but frequencies Conomitra soellingensis Microdrillia speyeri of common species differ. A fairly large number of species found at Mogenstrup is absent at Norre Moreover, Emarginula spp. (except E. punctulata Vissing, e.g. : Philippi, 1843) are absent from both faunas. Obvious differences are the absence of Astraea spp. Limopsis lamellata chattica and the lower diversity of the Cerithiopsidae in the Limopsis vonderhochti Acar aff. dentiens Pecten bifidus bifidus Pecten soellingensis Lower Rhine area. fauna from Glimmerode has 105 species in common with the Mogenstrup association, among which are: Astraea pustulosa Cocculina dittmeri Cirsope subeffusa Triforis sorgenfreii Neojanacus planatus Trophonopsis angustevaricata Pterynotus tristicus Acar aff. dentiens Astraea spp. Scissurella koeneniana Gibberula brevis Cerithiopsidae Muricidae. Coralliophila kochi Gibberula brevis Microdrillia spp. At Glimmerode the following species common at Mogenstrup, are missing: which are Pleurotomella margaritata Pleurotomella rasmusseni Rimosodaphnella lappanni pteropods. Limopsis aurita Limopsis vonderhochti. Many of these species are extremely rare in the Additionally, the Mogenstrup fauna did not yield any Amphineura or Emarginula spp. Brejning Clay of localities in the Vejle Fjord area fauna from Freden has 100 species in common with Mogenstrup, among which are:

35 73 Limopsis aurita Astraea pustulosa Cirsope subeffusa. Vexillum hastatum Gemmula trispiralis Turbonilla spp. following species from Mogenstrup are at Freden: missing Limopsis vonderhochti Emarginula spp. are present in the Söllingen fauna. Doberg fauna has 68 species with Mogenstrup: e.g. : in common almost all Cerithiopsidae Microdrillia speyeri. Acar aff. dentiens Astraea spp. An obvious difference is the presence of Emarginula spp. in the Freden fauna. following species, common at Mogenstrup, are absent from the Doberg fauna: fauna from the typical 'Kasseler Meeressand' (Niederkaufungen and Ahnetal) has 80 species in common with Mogenstrup, among which are: Limopsis aurita Limopsis vonderhochti Rissoa spp. Exilioidea elatior Metula scabricula Streptochetus soellingensis Conomitra soellingensis. many Cerithiopsidae Muricidae, except Lyrotyphis cuniculosus (Nyst, 1836) Vexillum hastatum Emarginula spp. are absent from both faunas. In Conomitra soellingensis Gemmula trispiralis comparison to the Mogenstrup fauna, a large Gemmula pseudokonincki. number of species are absent, e.g. : Doberg fauna comprises a large number of Acar aff. dentiens bivalve species, especially Pectinidae. Emarginula Limopsis aurita Palliolum hausmanni hausmanni Astraea pustulosa Cirsope spp. Rissoa spp. spp. are also present. 'Sternberger Gestein' fauna has 100 species in with common As Mogenstrup. this fauna originates from boulders of different ages (Chattian A, B and C) further analyses were not made. almost all Cerithiopsidae. PALAEOECOLOGICAL INTERPRETATIONS (AUTHOR: SCHNETLER) fauna from Sollingen has 75 species in common with Mogenstrup, among which are: In unit 1 molluscs are generally not very common and regularly distributed in the sediment. In the Acar aff. dentiens Scissurella koeneniana Astraea pustulosa uppermost part of unit 1 molluscs and other fossils are abundant. molluscan assemblage is most probably not a biocoenosis, as the fossils are con Pterynotus tristrichus. centrated in a thin horizon which has a sharp lower boundary; Fusiturris duchastelii, Gemmula Among the many Mogenstrup species that do spp., Polinices helicinus and Streptochetus cheruscus are found almost exclusively as juvenile specimens. not occur at Sollingen the following may be mentioned: However, of many molluscs both juvenile and adult specimens are found, e.g. the gastropods Limopsis aurita Limopsis vonderhochti Rissoa spp. Triphoridae Ancilla karsteni Astrea pustulosa (dominant), Lyrotyphis sejunctus, Hinia schlotheimi, Charonia and the bivalves flandrica Limopsis spp. abundant occurrence of Astraea pustulosa is very conspicuous. According to R. Janssen

36 74 (1978a: 161) the genus Astraea littoral con suggests gastropod species Charonia flandrica, on the ditions. This is also suggested by the of occurrence other hand, would indicate deeper water and the Alvania semperi, Cirsope spp., Gibberula brevis and occurrence of Cocculina dittmeri might even indicate Vexillum hastatum. common species Fusiturris rather deep water. genus Cocculina and related duchastelii and Polinices helicinus seem to be rather forms live on waterlogged wood (Marshall, 1985) facies independent since they are common in all at depths below 150 m together with Teredo sp., facies. and since Teredo sp. is rather common at Mogenstrup this might indicate deeper water. Table 3. Frequencies of bivalve and gastropod species. Very remarkable is the high diversity of Murici number of specimens % Bivalvia 1. Limopsis lamellata chattica Limopsis retifera Limopsis aurita Cyclocardia aff. kickxi dae and Cerithiopsidae, since representatives of these groups are very rare at other Danish Late Oligocene localities, e.g. Norre Vissing, Arhus and Skanderborg. We may assume that, generally speaking, the gastropod assemblage indicates littoral to higher sublittoral conditions Limopsis vonderhochti vonderhochli bivalve assemblage is dominated by Limopsis spp.: species of this genus make up 63.9% of 6. Astarte goldfussi praecursor Teredinidae et indet. gen. sp Portlandia pygmaea the bivalve total number of specimens. Limopsis is epibenthic and lives in a sublittoral environment. 9. Acar aff. dentiens L. retifera is very 10. Bathyarca Balhyarca bellula often found at Mogenstrup articulated specimens, indicating low current Gastropoda 1. Collonia troelsi Astraea pustulosa Fusiturris duchastelii Polinices helicinus Streptochetus cheruscus Lyrotyphis sejunctus Drepanocheilus speciosus Stenodrillia obeliscus Gemmula trispiralis Orthosurcula regularis reguláris velocities and hardly any transport. species Nucula spp., Nuculana westendorpi and Yoldia glaberrima are rather rare at Mogenstrup, but very common in the clayey sediments of the Danish Late Oligocene, e.g. the localities Àrhus, Norre Vissing and Skanderborg. se specimens are endobenthic. equally endobenthic species Astarte goldfussi praecursor and Cyclocardia aff. kickxi are rather common. Very interesting is the relatively common occurrence of Acar aff. dentiens, which is Table 4. UTM coordinates of Danish localities mentioned in this paper No. Locality Kind of outcrop Mapsheet Coordinates 1. Mogenstrup coastal cliff 1216 III SV NH Cilleborg former claypit 1216 II S0 NH Hadsten former roadexcavation 1315 III NV NH Arhus former railwaycut 1314 I VN0 NH Norre Vissing working claypit 1214 I N0 NH Skanderborg former motorwayexcavation 1214 I S0 NH Jensgârd coastal cliff 1313 I NV NG Fakkegrav Badehotel coastal cliff 1213 I S0 NG Brejning beach exposure 1213 I SV NG Bogeskov coastal cliff 1213 I S0 NH Kirstinebjerg Skov beach exposure 1213 II N0 NG teresting, since remains of these groups are very also common in the nearshore sediments at Glimmerode. In all, the bivalve assemblage indicates rare at other Danish Late Oligocene localities (for higher sublittoral conditions. instance Norre Vissing, Arhus and Skanderborg). abundance of spines and other remains of brachiopod Terebratula sp. is common. Ser the regular seaurchin Cidaris sp., of marginal pulids are also common, but balanids are absent. plates of asteroids and of Anthozoa is very in Teeth of some shark species were found: Pristiopho

37 Zool. Ann. Danm. Bull. Science, 75 rus sp., Synodontaspis acutissima (Agassiz, 1843) and Mr K. Ulleberg (Galleberg, Norway) is kindly Odontaspididae gen. et indet. sp. (F. von der Hocht, pers. comm., 1987). se species might indicate deeper water, but as the shark fauna is thanked for information on the Foraminifera zonation. We furthermore like to thank Dr C. Heilmann very limited at Mogenstrup and the state of preser Clausen (Geological Institute, University of vation is rather poor the teeth are possibly Arhus, Denmark) for stimulating critical remarks reworked. and helpful advice. A number of teleost otoliths were collected (coll. ISL, AJB). This material was examined by P.A.M. Gaemers (Leiden), who identified 17 species, of which Gadichtys the most common. According (pers. comm., 1987, attenuatus (Koken, Dr is 1891) to Dr Gaemers the otolith fauna indi 1989) cates rather deepwater ( m). two deep water genera Diaphus (lanternfishes) and Coelorinchus (grenadierfishes) are represented by a few specimens and all other species, except for Colliolus spp., indicate rather deep water. state of preservation indicates that this material is not reworked and is thus a good indication of deposition in a sublittoral environment. Mr S.L. Jakobsen and Mrs M. Bjerreskov (Geological Museum, University of Copenhagen, Denmark) were very helpful with regard to the MGUH collections. Mr O.B. Berthelsen (Geological Institute, Copenhagen) is thanked for doing part of the photographic work. MrJ.W.M. Jagt (Venlo, coeditor Netherlands), of this periodical, kindly corrected the English text. substantial assistance of Mr S.B. Andersen (Geological Institute, University of Arhus, Denmark) in critically reading and correcting the manuscript, providing material and, especially, for doing the photographic work, is gratefully acknowledged. ACKNOWLEDGEMENTS REFERENCES authors are very grateful to the following persons for (information on) molluscan material in Anderson, H.J., Die miocane ReinbekStufe in Nordund Westdeutschland und ihre Molluskenfauna. their collections: Mr Sten Bo Andersen (Frederi Fortschr. Geol. Rheinld. u. Westf., 14: cia, Mr Denmark); H.C. Hansen (Fredericia, Denmark); Mrs Denmark); Mr A.C. Janse (Brielle, Netherlands); J. Hillersborg (Randers, Mr F. von der Hocht (KerpenBalkhausen, F.R.G.), who also information on shark supplied material; Mr A.W. Museum of Janssen (National Natural History, Leiden, who Netherlands), also kindly examined some problematic molluscs, prepared textfigs 6 and 7, an earlier improved version of the and manuscript studied the Mogenstrup pteropods for a Dr R. separate paper; Janssen (Senckenberg Museum, Frankfurt am Main, F.R.G.), who also examined some kindly problematic molluscs; Mrs L.B. Jorgensen (Lime, Mr Denmark); W. Lappann (HeiligenhausIsenbiigel, F.R.G.); Lechner Den (Odense, Mrs R. mark); Dr P. Lozouet national d'histoire (Musee naturelle, Paris, Mrs E. France); Nielsen (Arhus, Denmark) and Mr M.S. Nielsen Denmark), who kindly placed his important collection (Odense, from Mogenstrup at my and in this disposal way contributed substantially to this paper. Dr P.A.M. Gaemers Museum of (National Natural History, Leiden, Netherlands) supplied information on otolith material. Beyer, C., OligocasnN. Miocaen i Nordvest Jylland. Faciesanalyse and Arhus magnetostratigrafi. (thesis University of Arhus), 165 pp. (unpublished). Beyrich, E., Die Conchylien des norddeutschen Tertiargebirges, 4/5. Berlin (Hertz): , pis Cossmann, M Catalogue illustre des coquilles de l'eocene des environs de Paris. Belg., 24: 3381, 12 pis. Soc. r. Malacol. Eriksen, K., En forekebig meddelelse om Tertiaeret ved Brejning pa sydsiden af Veile Fiord. Denmark, 9(2): geol. Soc. Gorges, J., Die Oberoligocanfauna Rumeln von am Niederrhein. Decheniana, 100(A): , 3 pis. Gorges, J., Die Lamellibranchiaten und Gastropoden des Oberoligocanen Meeressandes von Kassel. Abhandl. hess. L.Amt Bodenforsch., 4: pp, pis. Griindel, J., Bemerkungen Uberfamilie Cerithiopsacea zur (Gastropoda) sowie zur zur Fassung einiger ihrer Gattungen , 41 figs. Anz., lena, 204(34): Haq, B.U., J. Hardenbol & P R. Vail, Chronology of fluctuating levels since the Triassic. sea : Harder, P., De oligocaene Lag i Jaernbanegennem skaeringen ved Aarhus Station. Geol. Unders., (2)22: tabs, 4 pp., figs, 9 pis. Janssen, A.W., Mollusken uit het Mioceen van WinterswijkMiste. Een inventarisatie, met beschrijvingen en afbeeldingen van alle aangetroffen soorten. Amsterdam (K.N.N.V., N.G.V., R.G.M.), pp., pis.

38 Geol. Danm. Danm. Arch. Arch. Contr. Actes Danm. Kgl Bull. belyst Danm. New Bull. 76 Lieberkind,K., Ostracodstratigrafi i 0verste Eocaen og Janssen, A.W., Pteropod species (Mollusca, Gastropoda, Euthecosomata) from the Late Oligocene of Mo Oligocaen i Danmark ud fra Viborg 1 boringen og genstrup, Jylland, Denmark. Tert. Quatern. enkelte daglokaliteter. Arhus (thesis University of Arhus), Geol., 27(23): 8392, 10 figs, 1 pi. (printed in this issue). 112 pp, 23 pis. (unpublished) Janssen, R., 1978a. Die Scaphopoden und Gastropoden des Madsen, V., Om Tertieeret ved Mariager Fjord. Kasseler Meeressande von Glimmerode (Niederhessen). Danm. geol. Unders. (4)1(8): 41 pp. Jb., (A)4: 3195, 3 figs, 3 tabs, 7 pis. Marshall, B.A., Recent and Tertiary Cocculinidae and Janssen, R., 1978b. Die Mollusken des Oberoligozans (Chattium) in Nordseebecken, 1. Scaphopoda, Archaeogastro Pseudococculinidae (Mollusca: Zealand and New South Wales. Gastropoda) from New Zealand Journal poda, Mesogastropoda. Moll., 109(1/3): of Zoology, 12(1985): , 24 tabs, 14 figs , 1 fig., pis 914. Peyrot, A., Conchologie neogenique de l'aquitaine, 6. Soc. linn. Bordeaux, 84: 5128, 10 pis. Janssen, R., 1979a. Die Mollusken des Oberoligozans (Chattium) im Nordseebecken, 2. Neogastropoda, Euthyneura, Rasmussen, L.B., De miocasne formationer i Dan Cephalopoda. Moll., 109(4/6): , 5 pis. mark. geol. Unders., 4(5), 45 pp. Janssen, R. 1979b. Revision der Bivalvia des Oberoligozans (Chattium, Kasseler Meeressand). Geol. Abhandl. Rasmussen, L.B., Molluscan faunas and biostratigraphy of the marine younger Miocene Hessen, 78: 1181, 1 fig., 4 pis. formations in Denmark, 2. Palaeontology. 265 pp., 27 pis. geol. Unders., 2(92): Koch, E., K. Gripp & A. Franke, Die staatlichen Tiefbohrungen XIV, XV, XVI, XVII in den Vierlanden bei Hamburg. Jb. Hamb. wiss. Anst. 29, 4. Beih.: 33 pp., 8 figs, 1 pi. Ravn, J.P.J., Nogle bemaerkninger om de Oligocsene Geol. Soc. Den og Miocsene aflejringer i Jylland. mark, 12: 16. KristofTersen, F. N., Oligocaen og Miocaen i Novling, 1. geol. Unders., 3 (40): Larsen, G., & Dinesen, A., Vejle Fjord Formationen Ravn, J.P.J., Molluskfaunaen i Jyllands Tertiaeraflejringer. 3(2): 180 pp., 8 pis. Danske Vid. Selskab Skr., 7 (nat. mat.) ved Brejning. Sedimenterne, Foraminiferfaunaen (Olig Ravn, J.P.J., Om nye findesteder for Tertiasrforstenin Mioc.). geol. Unders., (2)82, 114 pp. ger i Danmark. geol. Soc. Denmark, 3: PLATE 1 Fig. 1. Acar aff. dentiens 022. (Cossmann & Peyrot, 1912); a: left valve, interior; b: left valve, exterior; x 10. Coll. MGUH 20 Fig. 2. Limopsis (Pectunculina) Coll. MGUH lamellata chattica subsp. left nov., holotype; a: valve, interior; b: left valve, exterior; x 9.5. Fig. 3. Limopsis (Pectunculina) lamellata chattica subsp. nov., paratype; a: right valve, interior; b: right valve, exterior; x 9.5. Coll. MGUH Fig. 4. Limopsis (Pectunculina) vonderhochti sp. nov., holotype; a: right valve, interior; b: right valve, exterior; x 3. Coll. MGUH Fig. 5. Limopsis (Pectunculina) vonderhochti sp. nov., paratype; a: left valve, interior; b: left valve, exterior; x 3. Coll. MGUH Fig. 6. Limopsis (Pectunculina) vonderhochti sp. nov., paratype; a: left valve, interior; b: left valve, exterior; x 3. Coll. SMF Fig. 7. Limopsis (Pectunculina) vonderhochti SMF sp. nov., paratype; a: right valve, interior; b: right valve, exterior; x 3. Coll. Fig. 8. Cocculina dittmeri (Anderson, 1964); apical view; x 5. Coll. MGUH Fig. 9. Lepetella helgae sp. nov., holotype; a: lateral view; b: apical view; x 20. Coll. MGUH Fig. 10. Fig. 11. Fig. 12. Fig. 13. Fig. 14. Lepetella jytteae sp. nov., holotype; a: lateral view; b: apical view; x 2. Coll. MGUH Astraea (Lithopoma) pustulosa (von Münster, 1844); apertural view; x 10. Coll. MGUH Homalopoma (Boutillieria) simplex (Philippi, 1843); apertural view; x 10. Coll. MGUH Homalopoma (? Leptothyropsis) a: sp.; apertural view; b: umbilical view; c: apical view; x 5. Coll. ENÅ. holotype; apertural view; x 30. Coll. MGUH Tubiola subangulata sp. nov., Fig. 6 from KrefeldLinn, F.R.G. (water well). Fig. 7 from Ahnetal, F.R.G. All other specimens from Mogenstrup (coastal cliff), Denmark.

39 PLATE 1 77\1

40 Danm. Danm. Danm. Palaont. Handbuch Bull. Abhandl. 78 Ravn, J. P. J Tertiaer. In : Oversigt over Danmarks TafelErklarungenvon A. von Koenen. geol. geologi (ed. V. Madsen) geol. Unders., 5(4), Spez.Karte Preussen, 4(4): ixii, 31 pis. 208 pp., 2 pis. Tembrock, M.L., Muriciden aus dem Mittel und Schneder, K.I., & C. Beyer, A Late Oligocene (Chat Oberoligocan und den Vierlandschichten des Nord tian B) mollusc fauna from the claypit of Galten Brick seebeckens. Abhandl., 1: , 3 tabs, 10 works at Norre Vissing, Jylland, Denmark. Meded. pis. Werkgr. Tert. Kwart. Geol., 24(3): , 3 figs, 1 Ulleberg, K., Foraminiferal zonation of the Danish tab., 2 pis. Oligocene sediments. geol. Soc. Denmark, 36: Sorgenfrei, T., Marint NedreMiocaen i Klintingeho ved paa Als. geol. Unders., 2(65), 143 pp., 8 Wenz, W., Gastropoda. Allgemeiner Teil und pis. Prosobranchia. der Palaozoologie, 6: xii + Sorgenfrei, T., Molluscan assemblages from the 1639 pp., 4211 figs. marine Middle Miocene of South Jutland and their environments, pis. geol. Unders. (2) 79: 503 pp., Speyer, O., & A. von Koenen, Die Bivalven der Casseler TertiarBildungen, mit einem Vorwort und Manuscript received 23 October 1989, revised version accepted 3 August PLATE 2 Fig. 1. Tubiola sp.; apertural view; x 30. Coll. MGUH Fig. 2. Collonia (Collonia) troelsi sp. nov., holotype; MGUH a: apertural view; b: umbilical view; c: apical view; x 20. Coll. Fig. 3.? Skenea sp.; a: apical view; b: umbilical view; x 30. Coll. MGUH Fig. 4. Neojanacusplanatus (Speyer, 1864); apertural view; x 3. Coll. LJL. Fig. 5. Triforis (Trituba) sorgenfreii sp. nov., holotype; apertural view; x 30. Coll. MGUH Fig. 6. Triforis (Trituba) sorgenfreii sp. nov., paratype; apertural view; x 11. Coll. MGUH Fig. 7. Triforis (Trituba) sorgenfreii sp. nov., paratype; apertural view; x 4. Coll. MGUH Fig. 8. Laiocochlis (Laiocochlis) supraoligocaenica sp. nov., holotype; apertural view; x 20. Coll. MGUH Fig. 9. Cerithiopsis (s. lat.) antonjansei sp. nov., holotype; apertural view; x 10. Coll. MGUH Fig. 10. Fig. 11. Fig. 12. Fig. 13. Cirsotrema (Opaliopsis) subglabrum sp. nov., paratype; apertural view; x 10. Coll. MGUH Cirsotrema (? Opaliopsis) aff. koeneni A.W. Janssen, 1967; apertural view, x 20. Coll. JHR. Cirsotrema sp. 1; apertural view; x 6. Coll. MGUH (Opaliopsis) Cirsotrema (Opaliopsis) sp. 2; apertural view; x 30. Coll. MGUH Fig. 14. Trophonopsis (Pagodula) angustevaricata (Gripp, 1912); apertural view; a: b: dorsal view; x 5. Coll. MGUH Fig. 15. Coralliophila (Hirtomurex) kochi (Beyrich, 1854); a: apertural view; b: dorsal view; x 5. Coll. MGUH Fig. 16. Searlesia ravni sp. nov., paratype; dorsal view; x 1. Coll. MGUH Fig. 16. Searlesia ravni sp. nov., paratype; a apertural view; b: lateral view; x 1. Coll. MGUH Fig. 7 from Kirstinebjerg Skov (beach exposure), Denmark. All other specimens from Mogenstrup (coastal cliff), Denmark.

41 PLATE 2 79\2

42 80 PLATE 3 Fig. 1. Fig. 2. Scalaspira (Scalaspira) Latirus (Pseudolatirus) sp.; apertural view; x 9. Coll. MGUH sp.; apertural view; x 11. Coll. JHR. Fig. 3. Gibberula brevis (von Koenen, 1890); apertural view; x 8. Coll. MGUH Fig. 4. R. Janssen, 1978; a: apertural view; b: dorsal view; x Apiocypraea (Apiocypraea) cf. humbergi 10. Coll. MGUH Fig. 5. Clavatula mogenstrupensis sp. nov., holotype; apertural view; x 11. Coll. MGUH Fig. 6. Babylonella pusilla (Philippi, 1843); apertural view; x 8. Coll. MGUH Fig. Microdrillia sp. nov., holotype; a: apertural view; b: dorsal view; x 8.5. Coll. MGUH ingerae Fig. 8. Microdrillia speyeri (Koch & Wiechmann, 1872); apertural view; x 8. Coll. LJL. Fig. 9. Microdrillia (Andersondrillia) brejningensis sp. nov., holotype; a: apertural view; b: dorsal view; x 8. Coll. MGUH Fig. 10. Fig. 11. Microdrillia (Andersondrillia) brejningensis sp. nov., paratype; apertural view; x 8. Coll. MGUH Pleurotomella (Pleurotomella) rasmusseni sp. nov., holotype; a: apertural view, x 8 ½; b: dorsal view, x 8. Coll. MGUH a: 12. Fig. R.Janssen, 1978; apertural view; x (Pleurotomella) margaritata 8. Coll. JHR. Fig. 13. Rimosodaphnella lappanni sp. nov., holotype; apertural view; b: dorsal view; x 5. Coll. MGUH Fig. 14.? Fig. 15.? Actaeopyramis (s. lat.) pseudopunctata sp. nov., paratype; apertural view; x 34. Coll. MGUH sp. nov., holotype; apertural view, x 23. Coll. MGUH Actaeopyramis (s. lat.) pseudopunctata Fig. 9 from Brejning (beach exposure), Denmark; Fig. 13 from Fakkegrav Badehotel (coastal cliff), Denmark; Fig. 15 from KrefeldLinn (water well), F.R.G. All other specimens from Mogenstrup (coastal cliff), Denmark.

43 PLATE 3 81\3