Pseudoscorpions of the Czech Republic and Slovakia: An annotated and revised checklist (Arachnida: Pseudoscorpiones)

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1 NORTH-WESTERN JOURNAL OF ZOOLOGY 8 (1): 1-21 NwjZ, Oradea, Romania, 2012 Article No.: Pseudoscorpions of the Czech Republic and Slovakia: An annotated and revised checklist (Arachnida: Pseudoscorpiones) Jana CHRISTOPHORYOVÁ 1*, František ŠŤÁHLAVSKÝ 2, Miroslav KRUMPÁL 1 and Peter FEDOR 3 1. Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynská dolina B 1, SK Bratislava, Slovakia. 2. Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ Prague 2, Czech Republic. 3. Department of Ecosozology (Conservation Biology), Faculty of Natural Sciences, Comenius University, Mlynská dolina B 1, SK Bratislava, Slovakia. * Corresponding author: J. Christophoryová, christophoryova@gmail.com Received: 03. April 2011 / Accepted: 27. August 2011 / Available online: 18. September 2011 / Printed: June 2012 Abstract. An annotated and revised checklist is presented comprising 58 species (8 families) of the arachnid order Pseudoscorpiones from the Czech Republic (38 species) and Slovakia (51 species), formerly Czechoslovakia. According to the latest detailed analyses and revisions Chthonius carinthiacus Beier, 1951, Neobisium validum (L. Koch, 1873), Roncus alpinus L. Koch, 1873 and Pselaphochernes dubius (O. P.-Cambridge, 1892) are excluded from the checklist from Slovakia. Formerly recorded Chthonius diophthalmus Daday, 1888, C. submontanus Beier, 1963, C. kewi Gabbutt, 1966, Neobisium spelaeum (Schiödte, 1847), N. jugorum (L. Koch, 1873), Roncus alpinus and R. transsilvanicus Beier, 1928 have not been included in the list from the Czech Republic. Some generally rare species are discussed. Keywords: Pseudoscorpiones, checklist, Central Europe, Czechoslovakia, species diversity. Introduction Pseudoscorpions are the fourth largest arachnid order (Harvey 2007) with more than 3380 species described in 25 families (Harvey 2009). Europe is a major centre of diversity, and in a review of the major geographical regions of the world (Africa, Asia, Europe, North America, Australia and Oceania, South America), Europe was found to have the highest richness with 760 species (Harvey 2007). But even here our knowledge on the distribution of many species appears rather sporadic and questionable, especially when older insufficient data have been taken into account. Moreover, in the Czech Republic and Slovakia, located in Central Europe, the knowledge on diversity and consequent progress have been limited since the division of the former Czechoslovakia in The first data on pseudoscorpions from the Czech Republic date back to the end of the 18th century (Preyssler 1790), with the description of Chthonius tetrachelatus. Almost hundred years later, Stecker (1874) introduced more significant records on 19 species, but some of them were later synonymized, including the newly described Chelifer serratus and Chernes bohemicus (see Harvey 2009). Some of the faunistic data published before the World War I (e.g. Stecker 1875, Simon 1879), which did not distinguish regions in the Austrian Hungarian Empire, have thus become confusing, with the exception of Chernes bohemicus (later synonymized with Lamprochernes nodosus, see Beier 1932b), endemic species in the Czech Republic. At the end of the 19th century there were only 14 pseudoscorpion species (belonging to five families) known from this region, but soon the first revising and summarizing checklist was published by Nosek (1901) and later updated (Nosek 1902). In Slovakia the first records on pseudoscorpions refer to several Hungarian authors of the late 19th century (e.g. Tömösváry 1882, Daday 1888a, 1888b, 1918). Nevertheless, most of our understanding of the pseudoscorpion fauna of Slovakia dates from the 1950s with the major contributions made by Verner (e.g. Verner 1958a, 1958b, 1960) and later by Krumpál (Krumpál 1979, 1980, Krumpál & Kiefer 1981, Krumpál & Cyprich 1987, 1988). The pseudoscorpions of the Czech Republic and Slovakia have been studied more intensively in the last decades of the 20th century, following the publication of the first Czechoslovak determination key (Verner 1971), with a lot of new and unpublished information, but without distinguishing national diversity. Before the dissolution of former Czechoslovakia in 1993 many rather general faunistic records were published, usually with no detailed information on the exact locality. The world pseudoscorpion catalogue from that period (Harvey 1991) contains several data on national diversity for Czechoslovakia, but difficult to separate for the two countries. Just three species (Neo-

2 2 bisium slovacum, N. beieri and Rhacochelifer quadrimaculatus) can be specified for Slovakia and five for the Czech Republic (Chthonius tetrachelatus, Chelifer cancroides and Lamprochernes nodosus, Chthonius diophthalmus and Neobisium sylvaticum inaculeatum) (Harvey 1991). Since the split in 1993, the pseudoscorpion fauna of the Czech Republic has been studied very intensively, resulting in plenty of first species records (Ducháč 1993a, 1995a, 1998a, Schmarda 1995, Šťáhlavský & Ducháč 2001, Ducháč & Mlejnek 2004, Ducháč et al. 2007, Laška et al. 2008, Christophoryová et al. 2011b) and the publication of faunistic data from numerous sites (Ducháč 1993b, 1997a, Šťáhlavský 2001, 2006a, 2006b, Šťáhlavský & Krásný 2007, Šťáhlavský & Tuf 2009). Apart from plenty of faunistic and ecological papers on pseudoscorpions in Slovakia in the 1990s (Krištofík et al. 1993, 1995, 1996, Krumpálová & Krumpál 1993, Ducháč 1996a, Fenďa et al. 1998), there has been an intensive research recently, analysing the diversity of pseudoscorpions in caves (Krumpál 2000, Mock et al. 2003, 2004, 2005, Christophoryová & Krumpál 2006c, Kováč et al. 2007, Papáč et al. 2007a, 2007b), bird nests (Cyprich et al. 2000, Krumpál et al , Krištofík et al. 2002, 2005, 2007, 2009, Christophoryová et al. 2007a, Christophoryová & Krištofík 2008, Christophoryová & Krumpál 2009, Christophoryová & Krumpálová 2010) as well as in numerous sites throughout the whole country (Krumpál & Krumpálová 2003, Christophoryová & Krumpál 2005, 2007b, Christophoryová et al. 2007b, Krumpál & Christophoryová 2007, Christophoryová 2009a, 2009b, Christophoryová & Mock 2009, Papáč et al. 2009, Christophoryová & Krumpál 2010). The identification key of pseudoscorpions from the Czech Republic and Slovakia was published recently (Christophoryová et al. 2011b). For the Czech Republic, the checklist with 28 species of seven families was published by Ducháč (1999b), revised soon after by Blick et al. (2004), presenting 33 species, and by Harvey (2009), with 5 more species. In Slovakia there is only one checklist available, completed by Harvey (2009) and offering information on 37 species (six families). The aim of our study is to revise species diversity of pseudoscorpions in Slovakia and the Czech Republic, formerly Czechoslovakia, taking the latest knowledge as well as national fauna specificity into account. The checklist is intended to provide a renewed foundation for future studies and discus- sions on this matter. Materials and Methods Christophoryová, J. et al. The current checklist has been compiled primarily using all available published records (determination keys, catalogues, conference proceedings), supplemented by faunistic information from many theses, deposited in university libraries. Analysing the century old works by Hungarian authors (Tömösváry 1882, Daday 1888a, 1888b, 1918) has generated many questions, with their specific linguistic and geopolitical problems. The present paper operates only with exact and clear data. For example, for the town of Slovenské Nové Mesto (Slovakia), established in 1918 after the split of Hungarian Sátoraljaújhely, the previously published records were revised to avoid confusion. On the other hand, some records omitted by Harvey (1991, 2009) and positively connected with the area of Slovakia have been taken into account. The data for the Czech Republic (CR) are followed by information for Slovakia (SR). All taxa include their relevant references chronologically listed, accompanied by authors, year and faunistic information (relevant pages in literature). The nomenclature for taxa derived from Harvey (2009). In specific cases the faunistic data set may express identification problems (misidentified or partly misidentified species). Lapsus calami in names of taxa in certain papers are expressed by [sic]. Results This checklist includes 58 species in eight families of the order Pseudoscorpiones from the Czech Republic (38 species) and Slovakia (51 species), formerly Czechoslovakia (Table 1). Chthonius diophthalmus, C. submontanus, C. kewi, Neobisium spelaeum, N. jugorum, Roncus alpinus, and R. transsilvanicus have been recently excluded from the fauna of the Czech Republic, and Chthonius carinthiacus, Neobisium validum, Roncus alpinus and Pselaphochernes dubius from the fauna of Slovakia. Order PSEUDOSCORPIONES de Geer, 1778 Suborder EPIOCHEIRATA Harvey, 1992 Superfamily CHTHONIOIDEA Daday, 1888 Family CHTHONIIDAE Daday, 1888 Subfamily Chthoniinae Daday, 1888 Tribe Chthoniini Daday, 1888 Genus Chthonius C. L. Koch, 1843 Subgenus Chthonius (Chthonius) C. L. Koch, 1932 Chthonius (Chthonius) carinthiacus Beier, 1951 CR: Chthonius pygmaeus carinthiacus Beier: Růžička

3 Checklist of pseudoscorpions from the Czech Republic and Slovakia 3 Table 1. List of pseudoscorpion species of the Czech Republic and Slovakia. Abbreviations: CZ Czech Republic, SK Slovakia. Taxa Country CZ SK Chthoniidae Chthonius (Chthonius) carinthiacus Beier, Chthonius (C.) diophthalmus Daday, Chthonius (C.) heterodactylus Tömösváry, Chthonius (C.) hungaricus Mahnert, Chthonius (C.) ischnocheles (Hermann, 1804) + + Chthonius (C.) orthodactylus (Leach, 1817) + + Chthonius (C.) pygmaeus Beier, Chthonius (C.) ressli Beier, Chthonius (C.) subterraneus Beier, Chthonius (C.) tenuis L. Koch, Chthonius (Ephippiochthonius) boldorii Beier, Chthonius (E.) fuscimanus Simon, Chthonius (E.) tetrachelatus (Preyssler, 1790) + + Mundochthonius carpaticus Rafalski, Mundochthonius styriacus Beier, Neobisiidae Microbisium brevifemoratum (Ellingsen, 1903) + Microbisium suecicum Lohmander, Neobisium (Blothrus) slovacum Gulička, Neobisium (Neobisium) beieri Verner, Neobisium (N.) brevidigitatum (Beier, 1928) + Neobisium (N.) carcinoides (Hermann, 1804) + + Neobisium (N.) carpaticum Beier, Neobisium (N.) crassifemoratum (Beier, 1928) + + Neobisium (N.) erythrodactylum (L. Koch, 1873) + + Neobisium (N.) fuscimanum (C. L. Koch, 1843) + + Neobisium (N.) inaequale Chamberlin, Neobisium (N.) jugorum (L. Koch, 1873) + Neobisium (N.) macrodactylum (Daday, 1888) + Neobisium (N.) polonicum Rafalski, Neobisium (N.) simile (L. Koch, 1873) + Neobisium (N.) sylvaticum (C. L. Koch, 1835) + + Roncus lubricus L. Koch, Roncus transsilvanicus Beier, Syarinidae Syarinus strandi (Ellingsen, 1901) + Larcidae Larca lata (Hansen, 1884) + + Cheiridiidae Apocheiridium (Apocheiridium) ferum (Simon, 1879) + Cheiridium museorum (Leach, 1817) + + Atemnidae Atemnus politus (Simon, 1878) + Diplotemnus insolitus Chamberlin, Cheliferidae Chelifer cancroides (Linnaeus, 1758) + + Dactylochelifer latreillii (Leach, 1817) + + Mesochelifer ressli Mahnert, Rhacochelifer peculiaris (L. Koch, 1873) + Rhacochelifer quadrimaculatus (Tömösváry, 1882) + Chernetidae Chernes cimicoides (Fabricius, 1793) + + Chernes hahnii (C. L. Koch, 1839) + +

4 4 Christophoryová, J. et al. Table 1. (continued) Taxa Country CZ SK Chernes nigrimanus Ellingsen, Chernes similis (Beier, 1932) + + Chernes vicinus (Beier, 1932) + + Dendrochernes cyrneus (L. Koch, 1873) + + Dinocheirus panzeri (C. L. Koch, 1837) + + Allochernes peregrinus Lohmander, Allochernes wideri (C. L. Koch, 1843) + + Anthrenochernes stellae Lohmander, Lamprochernes chyzeri (Tömösváry, 1882) + + Lamprochernes nodosus (Schrank, 1803) + + Lasiochernes pilosus (Ellingsen, 1910) + Pselaphochernes scorpioides (Hermann, 1804) + + Total number of species et al., 1995: 47; Ducháč, 2002c: 68. Chthonius (Chthonius) pygmaeus carinthiacus Beier: Šťáhlavský, 1996: 4. Chthonius (Chthonius) carinthiacus Beier: Harvey, 2009: ; Šťáhlavský & Tuf, 2009: 98; Christophoryová et al., 2011b: 36. Chthonius cf. pygmaeus carinthiacus Beier: Ducháč, 1997b: ; Ducháč, 2002c: 124, ; Christophoryová, 2006b: 9. Chthonius cf. submontanus Beier: Ducháč, 2002c: 102 (misidentification). Chthonius (Chthonius) tenuis L. Koch, 1873: Šťáhlavský, 2000: 17; Šťáhlavský, 2001: 80; Šťáhlavský & Král, 2004: 50; Šťáhlavský, 2006a: 169; Šťáhlavský, 2006c: 37, 79 (all misidentification). SR (note 1): Chthonius (Chthonius) pygmaeus carinthiacus Beier: Šťáhlavský, 1996: 4. Chthonius (Chthonius) diophthalmus Daday, 1888 CR (note 2): Chthonius (Chthonius) diophthalmus Daday: Harvey, 2009:. SR: Chthonius diophthalmus Daday: Christophoryová, 2006b: 8; Christophoryová, 2008a: 132, Christophoryová & Mock, 2009: 17; Mock et al., 2009: 268; Papáč et al., 2009: 37. Chthonius (Chthonius) diophthalmus Daday: Krumpál & Kiefer, 1981: ; Christophoryová, 2009a: 256; Christophoryová et al., 2011b: 36. Chthonius ksenemanni Hadži, 1939: Krumpál & Krumpálová, 2003: 119; Christophoryová, 2006b: 8 (synonymized by Schawaller, 1989: 6). Chthonius (Chthonius) ksenemanni Hadži: Krumpál & Kiefer, 1981: 128. Chthonius (Chthonius) heterodactylus Tömösváry, 1882 CR: Chthonius heterodactylus Tömösváry: Ducháč & Mlejnek, 2004: 37 38; Mikula, 2006: 18; Laška et al., 2008: 171. Chthonius (Chthonius) heterodactylus Tömösváry: Ducháč et al., 2007: 31 33; Harvey, 2009: ; Christophoryová et al., 2011b: 36. SR: Chthonius heterodactylus Tömösváry: Tömösváry, 1882: ; Krumpál & Krumpálová, 2003: ; Ducháč & Mlejnek, 2004: 37; Christophoryová, 2006b: 7, 8; Kárpáthegyi, 2006: 116; Mikula, 2006: 38; Ducháč et al., 2007: 32; Christophoryová & Mock, 2009: 17. Chthonius (Chthonius) heterodactylus Tömösváry: Szent-Ivány, 1941: 86; Rafalski, 1967: 6; Krumpál & Kiefer, 1981: 128; Ducháč, 1989: 180; Ducháč, 2002c: 31; Harvey, 2009: ; Christophoryová et al., 2011b: 36. Chthonius (Chthonius) hungaricus Mahnert, 1980 SR: Chthonius (Chthonius) hungaricus Mahnert: Christophoryová, 2009a: 256; Papáč et al., 2009: 37; Christophoryová et al., 2011a: 2 3; Christophoryová et al., 2011b: 36. Chthonius hungaricus Mahnert: Christophoryová, 2009b: 128; Christophoryová & Krumpál, 2009: 14. Chthonius (Chthonius) ischnocheles (Hermann, 1804) CR: Chthonius Rayi L. Koch, 1873: Stecker, 1874: 238; Simon, 1879: 74 75; Nosek, 1901: 21; Nosek, 1902: 60; Caporiacco, 1926: 75; Ducháč, 2002c: 11 12, 13 (synonymized by Chamberlin, 1929: 72). Chthonius ischnocheles (Hermann): Ducháč, 1999b: 14; Ducháč, 2002c: 20, 68, 82, 136, 138; Blick et al., 2004:. Chthonius (Chthonius) ischnocheles (Hermann): Šťáhlavský, 1996: 4; Jagoš, 2001: 33 34; Harvey, 2009: ; Christophoryová et al., 2011b: 36. SR: Chthonius ischnocheles (Hermann): Krumpál, 2000: 96; Krumpál & Krumpálová, 2003: 119; Christophoryová, 2009a: 254.

5 Checklist of pseudoscorpions from the Czech Republic and Slovakia 5 Chthonius (Chthonius) ischnocheles (Hermann): Krumpál, 1980: 23; Krumpál & Kiefer, 1981: 128; Šťáhlavský, 1996: 4; Jagoš, 2001: 33 34; Harvey, 2009: ; Christophoryová et al., 2011b: 36. Chthonius (Chthonius) ischnocheles ischnocheles (Hermann, 1804) CR: Chthonius (Chthonius) ischnocheles ischnocheles (Hermann): Harvey, 2009:. SR: Chthonius (Chthonius) ischnocheles ischnocheles (Hermann): Harvey, 2009:. Chthonius (Chthonius) orthodactylus (Leach, 1817) CR: Chthonius orthodactylus (Leach): Krásný, 1993: 37; Ducháč, 1998b: ; Ducháč, 1999b: 14; Ducháč, 2002c: 17, 18, 19, 68, 82, 138; Blick et al., 2004: ; Christophoryová, 2006b: 85; Mikula, 2006: 18. Chthonius (Chthonius) orthodactylus (Leach): Studničková, 1968: 25; Krumpál & Kiefer, 1981: 128; Ducháč, 1989: 180; Ducháč, 1994a: 140; Šťáhlavský, 1996: 5 6; Šťáhlavský, 2000: 17; Šťáhlavský, 2001: 80; Ducháč, 2002c: 31, 43; Šťáhlavský & Král, 2004: 50; Šťáhlavský, 2006c: 37, 79; Šťáhlavský & Krásný, 2007: 430; Harvey, 2009: ; Christophoryová et al., 2011b: 36. SR: Chthonius (Chthonius) orthodactylus (Leach): Krumpál, 1980: 23; Krumpál & Kiefer, 1981: 128; Šťáhlavský, 1996: 5; Harvey, 2009: ; Christophoryová et al., 2011b: 36. Chthonius (Chthonius) pygmaeus Beier, 1934 SR: Chthonius (Neochthonius) pygmaeus Beier: Krumpál & Kiefer, 1981: 127; Kárpáthegyi, 2007b: 92. Chthonius pygmaeus Beier: Ducháč, 2002c: 139. Chthonius (Neochthonius) pygmaeus pygmaeus Beier: Beier, 1963: 23. Chthonius (Chthonius) pygmaeus Beier: Kárpáthegyi, 2007a: 82 83; Christophoryová et al., 2011b: 36. Chthonius aff. pygmaeus Beier: Krumpál & Krumpálová, 2003: 120. Chthonius (Chthonius) ressli Beier, 1956 CR (note 3): Chthonius (Chthonius) ressli Beier: Šťáhlavský & Ducháč, 2005: 83; Šťáhlavský, 2006a: 169; Harvey, 2009: ; Christophoryová et al., 2011b: 36. SR (note 3): Chthonius (Chthonius) ressli Beier: Christophoryová et al., 2011b: 36. Chthonius ressli Beier: Christophoryová, 2008a: 132. Chthonius aff. ressli Beier: Mock et al., 2004: 38. Chthonius cf. ressli Beier: Mock et al., 2005: 100; Christophoryová, 2009a: 254. Chthonius (Chthonius) submontanus Beier, 1963 CR (note 4): not Chthonius cf. submontanus Beier: Ducháč, 2002c: 102 (misidentification, see Chthonius carinthiacus). Chthonius (Chthonius) subterraneus Beier, 1931 SR (note 5): Chthonius (Chthonius) subterraneus Beier: Christophoryová et al., 2011b: 36. Chthonius subterraneus Beier: Krumpál & Krumpálová, 2003: 119. Chthonius (Chthonius) tenuis L. Koch, 1873 CR (note 6): Chthonius tenuis L. Koch: Stecker, 1874: ; Simon, 1879: 72 73; Nosek, 1901: 21; Nosek, 1902: 60; Caporiacco, 1926: 76; Krásný, 1993: 38; Ducháč, 1998b: ; Ducháč, 1999b: 14; Ducháč, 2002c: 12, 13, 19, 68, 82, 141; Blick et al., 2004:. Chthonius (Chthonius) tenuis L. Koch: Šťáhlavský, 1996: 6; Šťáhlavský & Krásný, 2007: 430; Harvey, 2009: ; Christophoryová et al., 2011b: 36. not Chthonius (Chthonius) tenuis L. Koch: Šťáhlavský, 2000: 17; Šťáhlavský, 2001: 80; Šťáhlavský & Král, 2004: 50; Šťáhlavský, 2006a: 169; Šťáhlavský, 2006c: 37, 79 (all misidentification, see Chthonius carinthiacus). SR: Chthonius tenuis L. Koch: Krumpál & Krumpálová, 2003: 119; Mock et al., 2004: 38; Mock et al., 2005: 100; Christophoryová, 2009a: 254. Chthonius (Chthonius) tenuis L. Koch: Šťáhlavský, 1996: 6; Christophoryová et al., 2011b: 36. Subgenus Chthonius (Ephippiochthonius) Beier, 1930 Chthonius (Ephippiochthonius) boldorii Beier, 1934 SR: Chthonius (Ephippiochthonius) boldorii Beier: Christophoryová & Krumpál, 2005: 79; Christophoryová, 2006a: 63; Christophoryová, 2006b: 24; Christophoryová & Krumpál, 2006a: 43; Christophoryová & Krumpál, 2007a: 14; Christophoryová & Krumpál, 2007b: 29; Christophoryová et al., 2007b: 20; Christophoryová, 2008b: 22; Christophoryová & Krištofík, 2008: 22; Christophoryová et al., 2011b: 36. Chthonius boldorii Beier: Christophoryová & Krumpál, 2006b: 4; Christophoryová, 2008a: 133; Christophoryová & Krumpál, 2009: 15. Chthonius (Ephippiochthonius) fuscimanus Simon, 1900 CR: Chthonius austriacus Beier, 1931: Ducháč, 1998b: ; Ducháč, 1999b: 14; Ducháč, 2002c: 16, 82, 136 (synonymized by Gardini, 1980: 261). Chthonius (Ephippiochthonius) austriacus austria-

6 6 cus Beier, 1931: Rafalski, 1967: 7. Chthonius (Ephippiochthonius) fuscimanus Simon: Šťáhlavský, 1996: 9; Šťáhlavský, 2001: 80 81; Muster et al., 2004: 306; Šťáhlavský & Král, 2004: 50; Šťáhlavský, 2006a: 169; Šťáhlavský, 2006c: 37, 80; Harvey, 2009: ; Šťáhlavský & Tuf, 2009: 98; Christophoryová et al., 2011b: 36. Chthonius fuscimanus Simon: Ducháč, 1995c: ; Ducháč, 1997b: ; Ducháč, 2002c: 67, 68, 124; Blick et al., 2004: ; Christophoryová, 2006b: 9, 85; Laška, 2006: 19. SR: Chthonius (Ephippiochthonius) austriacus austriacus Beier: Krumpál & Kiefer, 1981: 127 (synonymized by Gardini, 1980: 261). Chthonius (Ephippiochthonius) fuscimanus Simon: Šťáhlavský, 1996: 9; Christophoryová & Krumpál, 2005: 79; Christophoryová, 2006a: 63; Christophoryová, 2006b: 24; Christophoryová & Krumpál, 2006a: 43; Christophoryová & Krumpál, 2007a: 14; Christophoryová & Krumpál, 2007b: 29; Christophoryová et al., 2007b: 20. Chthonius fuscimanus Simon: Christophoryová, 2006b: 8, 84; Christophoryová & Krumpál, 2006b: 4; Christophoryová, 2008a: 133; Christophoryová et al., 2011b: 36. Chthonius (Ephippiochthonius) kewi Gabbutt, 1966 CR (note 7): Chthonius (Ephippiochthonius) kewi Gabbutt: Harvey, 2009:. Chthonius cf. kewi Gabbutt: Ducháč, 1996c: 81 84; Ducháč, 1997b: ; Ducháč, 2002c: 69 72, 127, 138; Christophoryová, 2006b: 9. Chthonius (Ephippiochthonius) tetrachelatus (Preyssler, 1790) CR: Scorpio tetrachelatus Preyssler: Preyssler, 1790: 56; Ducháč, 2002c: 7. Chthonius trombidioides (Latreille, 1804): Stecker, 1874: 238; Ducháč, 2002c: 12 (misidentification). Chthonius tetrachelatus (Preyssler): Simon, 1879: 70 71; Nosek, 1901: 21; Nosek, 1902: 60 61; Ducháč, 1994b: ; Ducháč, 1995a: 52; Ducháč, 1995d: 130; Růžička et al., 1995: 47; Ducháč, 1996c: 82; Šťáhlavský, 1998: ; Ducháč, 1999b: 14; Ducháč, 2002c: 13, 19, 65, 68, 82, 102, , ; Blick et al., 2004: ; Christophoryová, 2006b: 9, 85; Laška, 2006: 19; Mikula, 2006: 18; Šťáhlavský, 2006b: 161; Gardini, 2009: 24; Christophoryová, 2010: 8. Chthonius (Ephippiochthonius) tetrachelatus (Preyssler): Ducháč, 1988: 105; Ducháč, 1989: ; Harvey, 1991: ; Krásný, 1993: 39 40; Christophoryová, J. et al. Ducháč, 1994a: ; Šťáhlavský, 1996: 7 8; Šťáhlavský, 2000: 18; Jagoš, 2001: 34 35, 48; Šťáhlavský, 2001: 81 82; Ducháč, 2002c: 26, 30 31, 43 44; Šťáhlavský & Král, 2004: 50; Šťáhlavský, 2006a: ; Šťáhlavský, 2006c: 37, 80; Christophoryová et al., 2007b: 19; Šťáhlavský & Krásný, 2007: 430; Harvey, 2009: ; Šťáhlavský & Tuf, 2009: 98; Christophoryová, 2010: 4; Christophoryová et al., 2011b: 36. SR: Chthonius tetrachelatus (Preyssler): Daday, 1888a: 134; Daday, 1888b: ; Daday, 1918: 2; Krumpál & Cyprich, 1988: 42; Ducháč, 1994a: 146; Fenďa et al., 1998: 27; Ducháč, 2002c: 49; Krumpál & Krumpálová, 2003: ; Christophoryová, 2006b: 8, 84; Christophoryová & Krumpál, 2006b: 4; Christophoryová, 2008a: 133; Christophoryová & Mock, 2009: 16. Chthonius (Ephippiochthonius) tetrachelatus (Preyssler): Szent-Ivány, 1941: (as Chthonius (Epiphiochthonius) [sic] tetrachelatus); Studničková, 1968: 24 25; Krumpál & Kiefer, 1981: 127; Ducháč, 1989: ; Šťáhlavský, 1996: 7; Jagoš, 2001: 34 35; Ducháč, 2002c: 30 31; Christophoryová, 2006a: 63; Christophoryová, 2006b: 24; Christophoryová & Krumpál, 2007a: 14; Christophoryová & Krumpál, 2007b: 29; Harvey, 2009: ; Christophoryová, 2010: 4; Christophoryová et al., 2011b: 36. Chthonius (Ephippiochthonius) cf. tetrachelatus (Preyssler): Christophoryová & Krumpál, 2006a: 43. Genus Mundochthonius Chamberlin, 1929 Mundochthonius carpaticus Rafalski, 1948 CR: Mundochthonius carpaticus Rafalski: Christophoryová et al., 2011b: 36, 38. SR: Mundochthonius carpaticus Rafalski: Krumpál & Krumpálová, 2003: 118; Christophoryová, 2008a: 133; Christophoryová, 2009a: 256; Christophoryová, 2009b: 129; Christophoryová & Mock, 2009: 16; Mock et al., 2009: 268; Papáč et al., 2009: 37; Christophoryová, 2010: 4; Christophoryová et al., 2011b: 36. Mundochthonius styriacus Beier, 1971 CR: Mundochthonius styriacus Beier: Ducháč, 1998b: ; Šťáhlavský, 1998: ; Ducháč, 1999b: 14; Šťáhlavský, 2000: 19; Šťáhlavský, 2001: 83; Šťáhlavský & Ducháč, 2001: 47; Ducháč, 2002c: 19, 82, ; Blick et al., 2004: ; Drogla & Lippold, 2004: 18; Šťáhlavský & Král, 2004: 50; Šťáhlavský & Ducháč, 2005: 83; Šťáhlavský, 2006c: 37, 80; Petrov & Šťáhlavský, 2007: 16; Harvey, 2009: ; Christophoryová, 2010: 8; Christophoryová et al., 2011b: 36.

7 Checklist of pseudoscorpions from the Czech Republic and Slovakia 7 Suborder IOCHEIRATA Harvey, 1992 Superfamily NEOBISIOIDEA Chamberlin, 1930 Family NEOBISIIDAE Chamberlin, 1930 Subfamily Neobisiinae Chamberlin, 1930 Genus Microbisium Chamberlin, 1930 Microbisium brevifemoratum (Ellingsen, 1903) CR: Microbisium brevifemoratum (Ellingsen): Christophoryová et al., 2011b: 36, 38. Microbisium suecicum Lohmander, 1945 CR: Microbisium suecicum Lohmander: Ducháč, 1998b: ; Šťáhlavský & Ducháč, 2001: 47; Ducháč, 2002c: 151; Blick et al., 2004: ; Drogla & Lippold, 2004: 18; Šťáhlavský & Ducháč, 2005: 83; Christophoryová, 2006b: 9; Šťáhlavský, 2006a: 170; Harvey, 2009: ; Christophoryová et al., 2011b: 36. SR: Microbisium suecicum Lohmander: Christophoryová, 2009b: 129; Christophoryová & Krumpál, 2009: 14; Christophoryová et al., 2011b: 36, 38. Genus Neobisium Chamberlin, 1930 Subgenus Neobisium (Blothrus) Schiödte, 1847 Neobisium (Blothrus) slovacum Gulička, 1977 SR: Neobisium (Blothrus) slovacum Gulička: Gulička, 1977a: 6 8; Gulička, 1977b: 27 28; Gulička, 1985: 111; Harvey, 1991: 383; Ducháč, 1996a: ; Ducháč, 1998c: 112; Ducháč, 1999c: ; Ducháč, 2000: 9 10; Ducháč & Mlejnek, 2000a: 15 20; Ducháč & Mlejnek, 2000b: 48 49; Krumpál, 2000: 95; Ducháč, 2001a: 8; Ducháč, 2001b: 1 142; Ducháč, 2002d: 18; Mlejnek & Ducháč, 2002: ; Ducháč, 2004: 52 53; Christophoryová, 2006b: 7; Christophoryová & Krumpál, 2006c: 20; Christophoryová et al., 2007c: 11; Harvey, 2009: ; Christophoryová, 2009a: 254, 256; Christophoryová et al., 2011b: 36. Neobisium slovacum Gulička: Gulička, 1982: 174; Ducháč, 1997c: 10 11; Vlk et al., 2001: 68 (as Neobissium [sic] slovacum); Papáč et al., 2007a: 52; Christophoryová, 2008a: 133. Neobisium (Blothrus) cf. slovacum Gulička: Ducháč, 1996b: Neobisium cf. slovacum Gulička: Ducháč, 1997c: Neobisium (Blothrus) spelaeum (Schiödte, 1847) CR (note 8): Blothrus spelaeus Schiödte: Stecker, 1875: Subgenus Neobisium (Neobisium) Chamberlin, 1930 Neobisium (Neobisium) beieri Verner, 1958 SR: Neobisium (Neobisium) beieri Verner: Verner, 1958b: ; Beier, 1963: 101; Studničková, 1968: 20; Verner, 1971: 25; Harvey, 1991: 347; Harvey, 2009: ; Christophoryová et al., 2011b: 36. Neobisium beieri Verner: Krumpál, 1980: 23; Christophoryová, 2006b: 7; Christophoryová et al., 2007c: 11. Neobisium (Neobisium) brevidigitatum (Beier, 1928) SR: Neobisium (Neobisium) brevidigitatum (Beier): Harvey, 2009: ; Christophoryová et al., 2011b: 36. Neobisium brevidigitatum (Beier): Krumpál, 1980: 23; Krumpál & Krumpálová, 2003: 120; Christophoryová, 2006b: 8. Neobisium (Neobisium) carcinoides (Hermann, 1804) CR: Obisium muscorum Leach, 1817: Stecker, 1874: 241; Nosek, 1901: 21; Nosek, 1902: 66, 70; Nosek, 1957: 59; Ducháč, 2002c: 12, 13, 14 (synonymized by Beier, 1963: 115). Obisium carcinoides (Hermann): Stecker, 1874: 240; Nosek, 1901: 21; Nosek, 1902: 64; Ducháč, 2002c: 12, 13. Neobisium muscorum (Leach): Verner, 1957: 88; Verner, 1959b: 379; Ducháč, 1999a: 105; Ducháč, 2002c: 16; Christophoryová, 2006b: 9, 85; Christophoryová et al., 2007b: 19. Neobisium (Neobisium) muscorum (Leach): Verner, 1971: 25. Neobisium (Neobisium) carcinoides (Hermann): Studničková, 1968: 16 18; Ducháč, 1988: 105; Ducháč, 1989: ; Ducháč, 1994a: ; Šťáhlavský, 1996:14 17; Šumpich et al., 1999: 163; Ducháč, 2002c: 26, 31 32, 44 45; Harvey, 2009: ; Šťáhlavský & Tuf, 2009: 98 99; Christophoryová, 2010: 5; Christophoryová et al., 2011b: 36. Neobisium carcinoides (Hermann): Ducháč, 1993b: 66; Krásný, 1993: 42 45; Ducháč, 1994b: ; Ducháč, 1995a: 52; Ducháč, 1995d: 130; Růžička et al., 1995: 47; Ducháč, 1997a: 117; Ducháč, 1999a: ; Ducháč, 1999b: 14; Šťáhlavský, 2000: 20; Jagoš, 2001: 29 30; Šťáhlavský, 2001: 84 85; Ducháč, 2002a: 58 59; Ducháč, 2002b: 24; Ducháč, 2002c: 17, 18, 35, 65, 68, 76, 82, 83 91, 93, 102, 103, 107, , , , 118, , , , 130, 131, 132, 134, ; Ducháč, 2003: 3; Blick et al., 2004: ; Laška, 2006: 19; Mikula, 2006: 18; Šťáhlavský, 2006a: ; Šťáhlavský, 2006b: 163; Šťáhlavský, 2006c: 70, 80; Šťáhlavský, 2007: 60; Šťáhlavský & Krásný, 2007: ; Božanić, 2008: 6; Christophoryová, 2010: 8.

8 8 Neobisium muscorum germanicum Beier, 1931: Verner, 1957: 88. SR: Obisium muscorum Leach: Daday, 1888a: 132; Daday, 1888b: 188; Ellingsen, 1910: 400; Nosek, 1957: 84 (synonymized by Beier, 1963: 115). Obisium carcinoides (Hermann): Tömösváry, 1882: ; Daday, 1888a: 131; Daday, 1888b: 187; Daday, 1918: 2. Obisium (Obisium) muscorum Leach: Beier, 1928: Neobisium muscorum (Leach): Verner, 1958b: 198; Krumpál & Cyprich, 1988: 42; Fenďa et al., 1998: 24; Mock et al., 2003: 37; Christophoryová & Krumpál, 2004: 10; Christophoryová, 2006b: 8; Christophoryová & Krumpál, 2006b: 4; Kováč et al., 2007: 49. Neobisium (Neobisium) muscorum (Leach): Verner, 1971: 25; Christophoryová & Krumpál, 2005: 79; Christophoryová, 2006a: 63; Christophoryová, 2006b: 24; Christophoryová & Krumpál, 2006a: 43; Christophoryová & Krumpál, 2006c: 20; Christophoryová & Krumpál, 2007b: 29; Christophoryová et al., 2007b: 20. Neobisium (Neobisium) carcinoides (Hermann): Studničková, 1968: 16 18; Ducháč, 1989: ; Šťáhlavský, 1996: 14; Jagoš, 2001: 29 30; Ducháč, 2002c: 31 32; Christophoryová, 2008b: 22; Christophoryová & Krištofík, 2008: 22; Christophoryová, 2009a: 256; Harvey, 2009: ; Christophoryová, 2010: 5; Christophoryová & Krumpál, 2010: 107; Christophoryová et al., 2011b: 36. Neobisium carcinoides (Hermann): Krumpál, 1980: 23 24; Fenďa et al., 1998: 27; Krumpál, 2000: 96; Krumpál et al., : 52; Ducháč, 2002c: ; Krumpál & Krumpálová, 2003: 120; Farkač et al., 2004: 221; Šťáhlavský, 2006c: 80; Christophoryová, 2008a: 133; Benová & Svatoň, 2009: 153; Christophoryová, 2009b: ; Christophoryová & Mock, 2009: 16; Mock et al., 2009: 268; Papáč et al., 2009: 37; Christophoryová & Krumpálová, 2010: 108. Neobisium cf. muscorum (Leach): Papáč et al., 2007b: 154. Neobisium (Neobisium) carcinoides carcinoides (Hermann, 1804) CR: Neobisium (Neobisium) muscorum muscorum (Leach, 1817): Hadži, 1939: Neobisium (Neobisium) carcinoides carcinoides (Hermann): Harvey, 2009:. SR: Neobisium (Neobisium) carcinoides carcinoides (Hermann): Harvey, 2009:. Christophoryová, J. et al. Neobisium (Neobisium) carpaticum Beier, 1935 SR: Neobisium (Neobisium) carpaticum Beier: Christophoryová, 2006a: 63; Christophoryová, 2006b: 24; Christophoryová & Krumpál, 2007b: 29; Harvey, 2009: ; Christophoryová, 2010: 5; Christophoryová et al., 2011b: 36. Neobisium carpaticum Beier: Krumpál, 1980: 24; Krumpál, 2000: 96; Krumpál & Krumpálová, 2003: 120; Christophoryová, 2006b: 8; Christophoryová & Krumpál, 2006b: 4; Christophoryová, 2009b: 130; Christophoryová & Mock, 2009: 17. Neobisium (Neobisium) erythrodactylum (L. Koch): Christophoryová & Krumpál, 2005: 79; Christophoryová & Krumpál, 2006a: 43; Christophoryová et al., 2007b: 20; Christophoryová, 2008b: 22 (all misidentification). Neobisium (Neobisium) crassifemoratum (Beier, 1928) CR: Neobisium (Neobisium) crassifemoratum (Beier): Christophoryová et al., 2011b: 36. Neobisium crassifemoratum (Beier): Ducháč, 2002b: 24; Ducháč, 2002c: 93, ; Ducháč, 2003: 3 4; Blick et al., 2004: ; Laška, 2006: Neobisium cf. crassifemoratum (Beier): Ducháč, 2002c: 132. SR: Neobisium (Neobisium) crassifemoratum (Beier): Verner, 1958a: ; Verner, 1960: 167; Beier, 1963: ; Rafalski, 1967: 8; Harvey, 2009: ; Christophoryová et al., 2011b: 36. Neobisium crassifemoratum (Beier): Krumpál, 1980: 24; Krumpál & Cyprich, 1988: 42; Fenďa et al., 1998: 27; Ducháč, 2003: 3; Krumpál & Krumpálová, 2003: 121; Farkač et al., 2004: 221; Christophoryová, 2006b: 7; Benová & Svatoň, 2009: 153. Neobisium (Neobisium) erythrodactylum (L. Koch, 1873) CR: Obisium erythrodactylum L. Koch: Stecker, 1874: ; Nosek, 1901: 21; Nosek, 1902: 65; Ducháč, 2002c: 12, 13. Neobisium (Neobisium) erythrodactylum (L. Koch): Studničková, 1968: 21; Ducháč, 1989: 181; Ducháč, 1994a: 143; Šťáhlavský, 1996: 11 12; Ducháč, 2002c: 32, 46; Harvey, 2009: ; Šťáhlavský & Tuf, 2009: ; Christophoryová et al., 2011b: 36. Neobisium erythrodactylum (L. Koch): Verner, 1957: 88; Verner, 1959b: 379; Ducháč, 1997a: 118; Ducháč, 1999b: 14; Šťáhlavský, 2000: 21; Jagoš, 2001: 31, 46; Šťáhlavský, 2001: 85 86; Ducháč, 2002b: 24; Ducháč, 2002c: 16, 17, 68, 77, 82, 93, 148; Ducháč, 2003: 3, 4; Blick et al., 2004: ; Christophoryová, 2006b: 9, 85; Mikula, 2006: 18; Šťáh-

9 Checklist of pseudoscorpions from the Czech Republic and Slovakia 9 lavský, 2006a: 172; Šťáhlavský, 2006b: 163; Šťáhlavský, 2006c: 81; Christophoryová et al., 2007b: 19. SR: Obisium erythrodactylum L. Koch: Tömösváry, 1882: ; Daday, 1888a: 132; Daday, 1888b: 188; Ellingsen, 1910: 397; Daday, 1918: 2. Obisium (Obisium) erythrodactylum L. Koch: Beier, 1928: Neobisium (Neobisium) erythrodactylum (L. Koch): Šťáhlavský, 1996: 11; Jagoš, 2001: 31; Harvey, 2009: ; Christophoryová et al., 2011b: 36. Neobisium erythrodactylum (L. Koch): Krumpál, 1980: 24; Krumpál & Krumpálová, 2003: 121; Krumpál & Christophoryová, 2007: 95. Not Neobisium (Neobisium) erythrodactylum (L. Koch): Christophoryová & Krumpál, 2005: 79; Christophoryová & Krumpál, 2006a: 43; Christophoryová et al., 2007b: 20; Christophoryová, 2008b: 22 (all misidentification, see Neobisium (Neobisium) carpaticum). Neobisium (Neobisium) fuscimanum (C. L. Koch, 1843) CR: Obisium fuscimanum C. L. Koch: Stecker, 1874: 240; Nosek, 1901: 21; Nosek, 1902: 65; Ducháč, 2002c: 12, 13. Obisium (Obisium) fuscimanum C. L. Koch: Beier, 1928: 295. Neobisium (Neobisium) fuscimanum (C. L. Koch): Beier, 1932a: 92 93; Rafalski, 1967: 12; Studničková, 1968: 19 20; Ducháč, 1989: 181; Ducháč, 1994a: ; Šťáhlavský, 1996: 12 13; Ducháč, 2002c: 32, 45 46; Harvey, 2009: ; Christophoryová Neobisium fuscimanum (C. L. Koch): Verner, 1957: 87; Beier, 1963: ; Ducháč, 1995b: ; Ducháč, 1999b: 14; Šťáhlavský, 2000: 21; Jagoš, 2001: 32 33; Šťáhlavský, 2001: 86; Ducháč, 2002b: 24; Ducháč, 2002c: 17, 57 63, 68, 82, 93, 107, 111, 117, 130, ; Ducháč, 2003: 3, 4; Blick et al., 2004: ; Kárpáthegyi & Kontschán, 2005: 5; Christophoryová, 2006b: 85; Laška, 2006: 20; Šťáhlavský, 2006a: 172; Šťáhlavský, 2006b: 163; Šťáhlavský, 2006c: 81. Neobisium fuscimanum ponticum Beier: Ducháč, 2002c: 16. Neobisium (Neobisium) fuscimanum ponticum Beier, 1963: Rafalski, 1967: 12. Neobisium fuscimanum fuscimanum (C. L. Koch): Ducháč, 2002c: 16. SR: Obisium (Obisium) fuscimanum C. L. Koch: Beier, 1928: 295. Neobisium (Neobisium) fuscimanum (C. L. Koch): Szent-Ivány, 1941: 87; Šťáhlavský, 1996: 12; Jagoš, 2001: 32 33; Harvey, 2009: ; Christophoryová et al., 2011b: 37. Neobisium fuscimanum (C. L. Koch): Krumpál, 1980: 24; Krumpál, 2000: 96; Krumpál & Krumpálová, 2003: 121; Kárpáthegyi & Kontschán, 2005: 5; Christophoryová, 2006b: 7; Christophoryová & Mock, 2009: 17; Mock et al., 2009: 268. Neobisium (Neobisium) inaequale Chamberlin, 1930 SR: Neobisium inaequale Chamberlin: Krumpál & Cyprich, 1988: 42; Fenďa et al., 1998: 27; Krištofík et al., 2005: 172 (as Neobium [sic] inaequale); Christophoryová, 2010: 2. Neobisium (Neobisium) inaequale Chamberlin: Harvey, 2009: ; Christophoryová Neobisium (Neobisium) jugorum (L. Koch, 1873) CR (note 9): Obisium jugorum L. Koch: Simon, 1879: 60 61; Caporiacco, 1926: 75. SR (note 9): Neobisium (Neobisium) jugorum (L. Koch): Verner, 1960: 167; Rafalski, 1967: 11; Verner, 1971: 25; Christophoryová Neobisium jugorum (L. Koch): Christophoryová, 2006b: 7. Neobisium (Neobisium) macrodactylum (Daday, 1888) SR (note 10): Neobisium (Neobisium) macrodactylum (Daday): Harvey, 2009: ; Christophoryová et al., 2011b: 37. Neobisium macrodactylum (Daday): Krumpál, 1980: 24; Christophoryová, 2006b: 8. Neobisium (Neobisium) macrodactylum macrodactylum (Daday, 1888) SR: Neobisium (Neobisium) macrodactylum macrodactylum (Daday): Harvey, 2009:. Neobisium (Neobisium) polonicum Rafalski, 1936 SR: Neobisium polonicum Rafalski: Ducháč, 1995e: 11 17; Ducháč, 1999d: ; Krumpál, 2000: 95; Krumpál & Krumpálová, 2003: ; Christophoryová, 2006b: 7; Šťáhlavský, 2006c: 81; Vařil, 2007: 15; Murienne et al., 2008: 174. Neobisium (Neobisium) polonicum Rafalski: Krumpál, 1979: ; Harvey, 2009: ; Christophoryová Neobisium (Neobisium) simile (L. Koch, 1873) SR (note 11): Obisium simile L. Koch: Tömösváry, 1882: Neobisium (Neobisium) simile (L. Koch): Christophoryová Neobisium (Neobisium) sylvaticum (C. L. Koch, 1835)

10 10 CR: Obisium sylvaticum C. L. Koch: Stecker, 1874: (as Obisium silvaticum [sic]); Ducháč, 2002c: 12 (as Obisium silvaticum [sic]). Obisium dumicola C. L. Koch: Nosek, 1901: 21; Nosek, 1902: 64, 70, Ducháč, 2002c: 13, 14 (synonymized by Beier, 1963: 83). Neobisium sylvaticum (C. L. Koch): Verner, 1957: 87; Verner, 1959b: 379; Krásný, 1993: 48; Ducháč, 1994b: ; Ducháč, 1995a: 52; Ducháč, 1997a: 118; Ducháč, 1999b: 14; Šťáhlavský, 2000: 21 22; Jagoš, 2001: 32; Šťáhlavský, 2001: 86; Ducháč, 2002b: 24; Ducháč, 2002c: 16, 17, 18 19, 65, 68, 77, 82, 93, 108, 114, , 120, , , 130, 132, 134, ; Ducháč, 2003: 3, 4; Blick et al., 2004: ; Christophoryová, 2006b: 9, 85; Mikula, 2006: 18; Šťáhlavský, 2006a: 172; Šťáhlavský, 2006b: 163; Christophoryová et al., 2007b: 19; Šťáhlavský & Krásný, 2007: 432; Božanić, 2008: 6. Neobisium (Neobisium) sylvaticum (C. L. Koch): Ducháč, 1988: 105; Harvey, 1991: ; Šťáhlavský, 1996: 10 11; Ducháč, 2002c: 26; Harvey, 2009: ; Šťáhlavský & Tuf, 2009: 100; Christophoryová, 2010: 5; Christophoryová et al., 2011b: 37. Neobisium (Neobisium) sylvaticum inaculeatum Hadži, 1939: Harvey, 1991: ; Harvey, 2009: (synonymized by Schawaller, 1989: 9). Neobisium (Neobisium) sylvaticum sylvaticum (C. L. Koch): Studničková, 1968: SR: Obisium sylvaticum C. L. Koch: Tömösváry, 1882: ; Daday, 1918: 2. Obisium dumicola C. L. Koch: Tömösváry, 1882: ; Daday, 1888a: 131; Daday, 1888b: 187 (synonymized by Beier, 1963: 83). Obisium (Obisium) dumicola nitidum Daday, 1988: Daday, 1888a: ; Daday, 1888b: (synonymized Beier, 1932a: 137). Neobisium sylvaticum (C. L. Koch): Ducháč, 1998c: ; Krumpál, 2000: 97; Krištofík et al., 2002: 605; Krumpál & Krumpálová, 2003: 122; Christophoryová, 2009b: 130; Christophoryová & Mock, 2009: 17; Papáč et al., 2009: 37; Christophoryová, 2010: 2. Neobisium (Neobisium) sylvaticum (C. L. Koch): Šťáhlavský, 1996: 10; Jagoš, 2001: 32; Christophoryová, 2008b: 22; Christophoryová, 2009a: 256; Harvey, 2009: ; Christophoryová, 2010: 5; Christophoryová Neobisium (Neobisium) validum (L. Koch, 1873) SR (note 12): Obisium validum L. Koch: Daday, 1918: 2. Christophoryová, J. et al. Obisium (Obisium) validum (L. Koch): Daday, 1888a: 130; Daday, 1888b: 186. Neobisium validum (L. Koch): Christophoryová, 2008a: 133. Genus Roncus L. Koch, 1873 Roncus alpinus L. Koch, 1873 CR (note 13): Roncus alpinus L. Koch: Cârdeiu, 1948: 4. Obisium alpinum (L. Koch): Simon, 1879: 65; Caporiacco, 1922: 67. SR (note 13): Roncus alpinus L. Koch: Tömösváry, 1882: ; Daday, 1918: 2; Krumpál & Krumpálová, 2003: 122; Christophoryová & Šťáhlavský, 2007: 12; Christophoryová, 2008a: 133. Roncus lubricus L. Koch, 1873 CR: Roncus lubricus L. Koch: Stecker, 1874: 237; Ducháč, 1998b: ; Šťáhlavský, 1998: ; Šťáhlavský, 2000: 22; Šťáhlavský, 2001: 86 87; Ducháč, 2002c: 11, 13; Harvey, 2009: ; Christophoryová et al., 2011b: 37. Obisium (Roncus) lubricum (L. Koch): Nosek, 1902: 65. Roncus lubricus lubricus L. Koch: Blick et al., 2004:. Roncus lubricus L. Koch: Šťáhlavský & Ducháč, 2001: Roncus cf. lubricus L. Koch: Ducháč, 2002c: 20, 151. SR: Roncus lubricus L. Koch: Tömösváry, 1882: ; Verner, 1971: 26; Ducháč, 2002c: 151; Krumpál & Krumpálová, 2003: ; Christophoryová & Krumpál, 2005: 79; Christophoryová, 2006a: 63; Christophoryová, 2006b: 8, 24; Christophoryová & Krumpál, 2006b: 4; Šťáhlavský, 2006c: 81; Christophoryová & Krumpál, 2007b: 29; Christophoryová et al., 2007b: 20; Christophoryová & Šťáhlavský, 2007: 12; Christophoryová, 2008a: 133; Christophoryová, 2008b: 22; Christophoryová, 2009b: ; Harvey, 2009: ; Papáč et al., 2009: 37; Christophoryová Obisium lubricum (L. Koch): Daday, 1888a: ; Daday, 1888b: Roncus (Roncus) lubricus L. Koch: Szent-Ivány, 1941: 88; Ducháč, 1994a: 143; Ducháč, 2002c: 46. Roncus (Roncus) lubricus lubricus L. Koch: Beier, 1963: 182. Roncus transsilvanicus Beier, 1928 CR (note 14): Roncus transsilvanicus Beier: Harvey, 2009:. SR (note 14): Roncus transsilvanicus Christophoryová & Šťáhlavský, 2007: 12; Christophory-

11 Checklist of pseudoscorpions from the Czech Republic and Slovakia 11 ová, 2008a: 133; Murienne et al., 2008: 174; Harvey, 2009: ; Christophoryová Roncus aff. transsilvanicus Beier: Krumpál & Krumpálová, 2003: 123. Family SYARINIDAE Chamberlin, 1930 Subfamily Syarininae Chamberlin, 1930 Genus Syarinus Chamberlin, 1925 Syarinus strandi (Ellingsen, 1901) CR: Syarinus strandi (Ellingsen): Ducháč, 1998a: ; Ducháč, 1998b: ; Ducháč, 1999b: 14; Šťáhlavský & Ducháč, 2001: 48; Ducháč, 2002c: 79 81, 82, 152, 159; Blick et al., 2004: ; Drogla & Lippold, 2004: 25; Šťáhlavský & Ducháč, 2005: 83; Harvey, 2009: ; Christophoryová Superfamily GARYPOIDEA Simon, 1879 Family LARCIDAE Harvey, 1992 Genus Larca Chamberlin, 1930 Larca lata (Hansen, 1884) CR: Larca lata (Hansen): Ducháč, 1993a: 36; Ducháč, 1993b: 66; Ducháč, 1995d: 130; Judson & Legg, 1996: 205; Šťáhlavský, 1996: 18; Ducháč, 1998b: ; Ducháč, 1999b: 14; Ducháč, 2002c: 35, 39, 68, 82, 152; Blick et al., 2004: ; Drogla & Lippold, 2004: 26; Šťáhlavský & Ducháč, 2005: 83; Christophoryová, 2006b: 9; Šťáhlavský, 2006c: 80; Christophoryová et al., 2007b: 19; Harvey & Volschenk, 2007: 368; Petrov & Šťáhlavský, 2007: 18; Christophoryová & Krištofík, 2008: 22; Murienne et al., 2008: 174; Harvey, 2009: ; Christophoryová et al., 2011b: 37. SR: Larca lata (Hansen): Christophoryová & Krištofík, 2008: 22; Christophoryová & Krumpál, 2009: 14; Christophoryová & Krumpálová, 2010: 108; Christophoryová et al., 2011a: 2 4; Christophoryová Superfamily CHEIRIDIOIDEA Hansen, 1894 Family CHEIRIDIIDAE Hansen, 1894 Subfamily Cheiridiinae Hansen, 1894 Genus Apocheiridium Chamberlin, 1924 Subgenus Apocheiridium (Apocheiridium) Chamberlin, 1924 Apocheiridium (Apocheiridium) ferum (Simon, 1879) CR: Apocheiridium ferum (Simon): Ducháč, 1997b: ; Ducháč, 1998b: ; Ducháč, 1999b: 14; Šťáhlavský & Ducháč, 2001: 48; Ducháč, 2002c: 82, 127, ; Blick et al., 2004: ; Šťáhlavský, 2006c: 78. Apocheiridium (Apocheiridium) ferum (Simon): Harvey, 2009: ; Christophoryová Genus Cheiridium Menge, 1855 Cheiridium museorum (Leach, 1817) CR: Cheiridium museorum (Leach): Stecker, 1874: 232; Nosek, 1901: 21; Nosek, 1902: 49, 70; Verner, 1957: 88; Studničková, 1968: 22; Verner, 1971: 27; Krásný, 1993: 33 34; Ducháč, 1994a: 143; Ducháč, 1995d: 130; Šťáhlavský, 1996: 19 20; Ducháč, 1999b: 14 (as Cheridium [sic] museorum); Šťáhlavský, 2000: 4 5; Šťáhlavský, 2001: 87 88; Ducháč, 2002c: 11 (as Chernes [sic] museorum), 13, 14, 18, 19, 46, 68, 82, 161; Blick et al., 2004: ; Šťáhlavský, 2006b: 163; Šťáhlavský, 2006c: 78; Šťáhlavský & Krásný, 2007: 432; Harvey, 2009: ; Christophoryová SR: Cheiridium museorum (Leach): Tömösváry, 1882: ; Nosek, 1957: 84; Verner, 1971: 27; Krumpál & Cyprich, 1988: 42; Ducháč, 1994a: 143; Krištofík et al., 1995: 490 (as Cheridium [sic] museorum); Šťáhlavský, 1996: 19; Fenďa et al., 1998: 24; Krumpál et al., : 52; Ducháč, 2002c: 46; Christophoryová, 2008a: 133; Christophoryová & Krištofík, 2008: 22; Harvey, 2009: ; Christophoryová, 2010: 1, 2, 5; Christophoryová & Krumpál, 2010: 107; Christophoryová & Krumpálová, 2010: 108; Christophoryová Superfamily CHELIFEROIDEA Risso, 1826 Family ATEMNIDAE Chamberlin, 1931 Subfamily Atemninae Chamberlin, 1931 Genus Atemnus Canestrini, 1884 Atemnus politus (Simon, 1878) SR: Atemnus politus (Simon): Krumpálová & Krumpál, 1993: 36; Christophoryová, 2006b: 8, 85; Šťáhlavský, 2006c: 78; Christophoryová, 2008a: 133; Harvey, 2009: ; Christophoryová et al., 2011b: 37. Subfamily Miratemninae Beier, 1932 Genus Diplotemnus Chamberlin, 1933 Diplotemnus insolitus Chamberlin, 1933 SR: Diplotemnus insolitus Chamberlin: Harvey, 2009: ; Christophoryová Diplotemnus piger (Simon, 1878): Verner, 1959a: 61 63; Beier, 1963: 248; Verner, 1971: 28 (synonymized by Dashdamirov & Schawaller, 1993: 7). Diplotemnus ophthalmicus Redikorzev, 1949: Christophoryová, 2006b: 7; Christophoryová, 2008a: 133 (synonymized by Dashdamirov & Schawaller, 1993: 7). Diplotemnus insolitus insolitus Chamberlin, 1933 SR: Diplotemnus insolitus insolitus Chamberlin: Harvey, 2009:.

12 12 Family CHELIFERIDAE Risso, 1826 Subfamily Cheliferinae Risso, 1826 Genus Chelifer Geoffroy, 1762 Chelifer cancroides (Linnaeus, 1758) CR: Chelifer cancroides (Linnaeus): Stecker, 1874: 236; Nosek, 1901: 21; Nosek, 1902: 51, 70; Verner, 1957: 88; Studničková, 1968: 23; Verner, 1971: 31; Ducháč, 1989: 181; Harvey, 1991: ; Krásný, 1993: 35; Ducháč, 1994a: 145; Ducháč, 1995d: 130; Sláma, 1995: 544; Šťáhlavský, 1996: 31 32; Ducháč, 1999b: 14; Šťáhlavský, 2000: 4, 23; Šťáhlavský, 2001: 88; Ducháč, 2002c: 11, 13, 13 14, 18, 19, 32, 48, 68, 82, 111, ; Krištofík et al., 2003: 232; Blick et al., 2004: ; Šťáhlavský, 2006c: 78; Šťáhlavský, 2007: 60; Šťáhlavský & Krásný, 2007: 432; Harvey, 2009: ; Christophoryová Chelifer ixoides C. W. Hahn, 1834: Stecker, 1874: 236; Ducháč, 2002c: 11 (synonymized by Simon, 1878: 155). Chelifer serratus Stecker, 1974: Stecker, 1874: ; Nosek, 1901: 21; Ducháč, 2002c: 11, 13 (synonymized by Beier, 1932b: 236). not Chelifer cancroides (Linnaeus): Verner, 1957: 88 (partly misidentification, partly see Mesochelifer ressli). SR: Chelifer cancroides (Linnaeus): Tömösváry, 1882: ; Daday, 1888a: 121; Daday, 1888b: 176; Nosek & Lichard, 1962: 35; Krumpál & Cyprich, 1988: 42; Cyprich et al., 1992: 78; Ducháč, 1994a: 145; Šťáhlavský, 1996: 31; Fenďa et al., 1998: 24; Cyprich et al., 2000: 190; Krumpál et al., : 52; Ducháč, 2002c: 48; Krumpál & Krumpálová, 2003: 123; Varga & Fedor, 2005: 45; Harvey, 2009: ; Christophoryová, 2010: 1, 2, 5; Christophoryová & Krumpál, 2010: 107; Christophoryová & Krumpálová, 2010: 108; Christophoryová et al., 2011b: 37. Chelifer cancroides cancroides (Linnaeus, 1758) CR: Chelifer cancroides cancroides (Linnaeus): Harvey, 2009:. SR: Chelifer cancroides cancroides (Linnaeus): Harvey, 2009:. Genus Dactylochelifer Beier, 1932 Dactylochelifer latreillii (Leach, 1817) CR: Chelifer latreillii Leach: Nosek, 1902: 54 (as Chelifer Latreille sub Schäfferi). Chelifer Schäfferi C. L. Koch, 1839: Stecker, 1874: 237; Nosek, 1901: 21; Ducháč, 2002c: 11, 13 (synonymized by Simon, 1879: 22). Dactylochelifer latreillei (Leach): Verner, 1957: 88; Šťáhlavský, 1996: 33; Ducháč, 1998b: ; Ducháč, Christophoryová, J. et al. 1999b: 14; Ducháč, 2002c: 68, 82, 162; Blick et al., 2004: ; Harvey, 2009:. Dactylochelifer latreillii (Leach): Christophoryová SR: Chelifer degeerii C. L. Koch, 1835: Tömösváry, 1882: (as Chelifer De Geeri) (synonymized by Pavesi, 1884: 455). Dactylochelifer latreillei (Leach): Krumpál & Cyprich, 1988: 42; Krištofík et al., 1993: 495; Ducháč, 1994a: 145; Šťáhlavský, 1996: 33; Fenďa et al., 1998: 24; Krumpál et al., : 52; Ducháč, 2002c: 48; Krištofík et al., 2002: 605 (as Dactilochelifer [sic] latreillei); Christophoryová, 2006b: 8; Christophoryová et al., 2007a: 16; Krumpál & Christophoryová, 2007: 97; Christophoryová, 2008a: 133; Christophoryová, 2008b: 22; Christophoryová & Krištofík, 2008: 22; Harvey, 2009: ; Christophoryová, 2009b: 131; Christophoryová, 2010: 1, 2, 5 6; Christophoryová & Krumpál, 2010: ; Christophoryová & Krumpálová, 2010: 108. Dactylochelifer latreillii (Leach): Christophoryová Dactylochelifer latreillii latreillii (Leach, 1817) CR: Dactylochelifer latreillii latreillii (Leach): Harvey, 2009:. SR: Dactylochelifer latreillii latreillii (Leach): Harvey, 2009:. Genus Mesochelifer Vachon, 1940 Mesochelifer ressli Mahnert, 1981 CR: Mesochelifer ressli Mahnert: Ducháč, 1995c: ; Šťáhlavský, 1996: 33; Ducháč, 1997b: ; Ducháč, 1999b: 14; Ducháč, 2002c: 67, 68, 82, 124, ; Blick et al., 2004: ; Christophoryová, 2006b: 9; Šťáhlavský, 2006a: ; Šťáhlavský, 2006c: 79; Harvey, 2009: ; Šťáhlavský & Tuf, 2009: 100; Christophoryová Chelifer cancroides (Linnaeus): Verner, 1957: 88 (partly misidentification). SR: Mesochelifer ressli Mahnert: Krumpálová & Krumpál, 1993: 37; Ducháč, 1995c: ; Ducháč, 2002c: 67; Drogla & Lippold, 2004: 29; Christophoryová, 2006b: 8; Harvey, 2009: ; Christophoryová, 2010: 2; Christophoryová et al., 2011b: 37. Genus Rhacochelifer Beier, 1932 Rhacochelifer peculiaris (L. Koch, 1873) SR (note 15): Chelifer peculiaris L. Koch: Tömösváry, 1882: ; Daday, 1888a: ; Daday, 1888b: ; Daday, 1918: 2. Rhacochelifer peculiaris (L. Koch): Verner, 1960:

13 Checklist of pseudoscorpions from the Czech Republic and Slovakia ; Verner, 1971: 31; Christophoryová, 2008a: 133; Christophoryová Rhacochelifer quadrimaculatus (Tömösváry, 1882) SR: Chelifer quadrimaculatus Tömösváry: Tömösváry, 1882: ; Daday, 1918: 2. Rhacochelifer quadrimaculatus (Tömösváry): Verner, 1960: 168; Verner, 1971: 31; Harvey, 1991: 530; Christophoryová, 2006b: 7; Christophoryová, 2008a: 133; Harvey, 2009: ; Christophoryová, 2010: 2; Christophoryová Family CHERNETIDAE Menge, 1855 Subfamily Chernetinae Menge, 1855 Tribe Chernetini Menge, 1855 Genus Chernes Menge, 1855 Chernes cimicoides (Fabricius, 1793) CR: Chelifer cimicoides (Fabricius): Nosek, 1901: 21; Nosek, 1902: 51, 70; Ducháč, 2002c: 13, 14. Chernes cimicoides (Fabricius): Stecker, 1874: 234; Verner, 1971: 30; Šťáhlavský, 1996: 25; Ducháč, 1999b: 14; Ducháč, 2002c: 11, 18, 68, 82, 186; Krištofík et al., 2003: 232; Blick et al., 2004: ; Šťáhlavský, 2006b: 163; Harvey, 2009: ; Christophoryová Chernes Mengei L. Koch, 1873: Stecker, 1874: 234; Ducháč, 2002c: 11 (synonymized by Beier, 1932b: 155). Chelifer Mengei L. Koch: Nosek, 1901: 21; Ducháč, 2002c: 13. not Chernes cimicoides (Fabricius): Verner, 1957: 88 (misidentification, see Chernes hahnii). SR: Chernes cimicoides (Fabricius): Tömösváry, 1882: ; Daday, 1888a: ; Daday, 1888b: 172; Verner, 1960: 168; Verner, 1971: 30; Krumpál & Cyprich, 1988: 42; Ducháč, 1994a: 144 (as Chernes cimocoides [sic]); Šťáhlavský, 1996: 25; Krumpál et al., : 52; Ducháč, 2002c: 47 (as Chernes cimocoides [sic]); Krištofík et al., 2002: 605; Christophoryová, 2006b: 8; Christophoryová, 2009b: 131; Harvey, 2009: ; Christophoryová, 2010: 2; Christophoryová & Krumpál, 2010: 108; Christophoryová Chernes hahnii (C. L. Koch, 1839) CR: Chernes hahnii (C. L. Koch): Ducháč, 1994a: 144 (as Chernes hahni); Šťáhlavský, 1996: 26 (as Chernes hahni); Ducháč, 1999b: 14 (as Chernes hahni); Šťáhlavský, 2000: 25; Šťáhlavský, 2001: 90 91; Ducháč, 2002c: 20, 47, 68, 82 (all as Chernes hahni), 187; Blick et al., 2004: ; Vítková et al., 2005: 147; Šťáhlavský, 2006a: 173; Šťáhlavský, 2006b: 163; Šťáhlavský, 2006c: 79; Šťáhlavský & Krásný, 2007: ; Božanić, 2008: 6; Harvey, 2009: ; Šťáhlavský & Tuf, 2009: 100; Christophoryová, 2010: 5; Christophoryová Chernes cimicoides (Fabricius): Verner, 1957: 88 (misidentification). SR: Chernes hahnii (C. L. Koch): Krumpál & Cyprich, 1988: 42 (as Chernes hahni); Ducháč, 1994a: 144 (as Chernes hahni); Krištofík et al., 1996: 560 (as Chernes hahni); Šťáhlavský, 1996: 26 (as Chernes hahni); Krumpál et al., : 52 (as Chernes hahni); Ducháč, 2002c: 47 (as Chernes hahni); Krumpál & Christophoryová, 2007: 96; Christophoryová & Mock, 2009: 17; Harvey, 2009: ; Christophoryová, 2010: 1, 6; Christophoryová & Krumpál, 2010: 108; Christophoryová & Krumpálová, 2010: 108; Christophoryová et al., 2011b: 37. Chernes similis (Beier): Christophoryová et al., 2007a: 16; Christophoryová & Krištofík, 2008: 22 (all misidentification). Chernes nigrimanus Ellingsen, 1897 CR: Chernes nigrimanus Ellingsen: Ducháč, 1995c: ; Schmarda, 1995: 61 65; Šťáhlavský, 1996: 26; Ducháč, 1998b: ; Ducháč, 1999b: 14; Šťáhlavský, 2001: 73; Ducháč, 2002c: 19, 67, 68, 82, 188; Blick et al., 2004: ; Drogla & Lippold, 2004: 33 34; Šťáhlavský & Ducháč, 2005: 83; Šťáhlavský, 2006c: 79; Harvey, 2009: ; Christophoryová Chernes similis (Beier, 1932) CR: Chernes similis (Beier): Šťáhlavský, 1996: 27; Ducháč, 1997b: ; Ducháč, 1998b: ; Ducháč, 1999b: 14; Šťáhlavský, 2000: 13; Ducháč, 2002c: 18, 20, 82, 127, 188; Blick et al., 2004: ; Šťáhlavský, 2006a: ; Šťáhlavský, 2006c: 79; Harvey, 2009: ; Christophoryová SR: Toxochernes similis (Beier): Krumpál, 1980: 24; Krumpál et al., : 52. Chernes similis (Beier): Krumpál & Cyprich, 1988: 42; Krumpálová & Krumpál, 1993: 37; Šťáhlavský, 1996: 27; Ducháč, 1997b: ; Ducháč, 2002c: 127; Christophoryová & Krumpál, 2005: 79; Christophoryová, 2006a: 63; Christophoryová, 2006b: 8, 24; Christophoryová & Krumpál, 2006b: 4; Christophoryová & Krumpál, 2007b: 29; Christophoryová et al., 2007b: 20; Harvey, 2009: ; Christophoryová not Chernes similis (Beier): Christophoryová et al., 2007a: 16; Christophoryová & Krištofík, 2008: 22 (all misidentification, see Chernes hahnii). Chernes vicinus (Beier, 1932) CR: Chernes vicinus (Beier): Šťáhlavský & Ducháč, 2001: 49; Ducháč, 2002c: 189; Blick et al., 2004: ;