Upper Albian agglutinated foraminifera from two wells in Northeast Germany

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1 445 Upper Albian agglutinated foraminifera from two wells in Northeast Germany RENATA SZAREK, BOGUMlEA KL OSOWSKA 2, ANDREAS PROKOpd, WOLFGANG KUIfNiI and THOMAS WAGNER 4 1. Institut fur Geowissenschaften der Christian-Albrechts UniversWit, Olshausenstr. 40, Kiel, Germany. 2. Marine Aardwetenschappen Centre, Vrije Universiteit, De Boelelaan 1085, 1081 HV Amsterdam, The Netherlands 3. Department of Geology, University of Ottawa, P.O.Box 450, Stn. A, Ottawa, Ontario, KIN 6N5, Canada. 4. Fachbereich 5, Geowissenschaften, Universitat Bremen, Klagenfurter Str. 5, Bremen, Germany. ABSTRACT We examined agglutinated foraminifera in Upper Albian sediments from two cored wells in distinctly different palaeogeographic settings in NE Germany. The southern well E Daehre 4/83 was influenced by high rates of terrigenous flux from the Bohemian Massif. The benthic foraminiferal assemblages from this well are characterised by high percentages of agglutinated foraminifera (average of about 30%), low faunal density (maximum 300 individual/ g sediment), and higher abundance of organically cemented forms such as Rhizammina, Ammodiscus, Glomospira, Haplophragmoides and Trochammina. The northern well Gt Waren 1/81 was situated at the flank of a pelagic high and was largely sheltered from terrigenous flux. Agglutinated foraminifera make up only approximately 15% of the benthic assemblages, the faunal density reaches 1000 individuals/ gram sediment and calcareous cemented forms including Spiroplectinata, Falsogaudryinella, Tritaxia, Dorothia, Arenobulimina and Eggerellina dominate the agglutinated assemblages. Since the palaeobathymetric positions of the two wells were not significantly different, the main factors causing the differences in the assemblage composition of agglutinated foraminifera may be differences in primary productivity and terrigenous flux. INTRODUCTION Agglutinated foraminiferal assemblages of mid Cretaceous Boreal shelf seas often contain unusually high numbers of "primitive" organically cemented forms including ammodiscids, astrorhizids, lituolids and trochamminids. These assemblages have no real analogues, since modern shelf assemblages are completely dominated by calcareous benthic foraminifera (i.e., Elphidium, Ammonia), Lower Cretaceous assemblages have high numbers of lagenids and epistominids, and Upper Cretaceous benthic assemblages of the Boreal chalk seas almost completely lack "primitive" agglutinated forms. Several environmental constraints were proposed to explain these unusual agglutinated assemblages, which superficially resemble deep-water faunas: (1) influence of cold arctic water masses (Kemper, 1987), (2) oxygen deficiency under warm, saline water masses (Kuhnt & Wiedmann, 1995), (3) high fluxes of terrigenous sediment (Prokoph et al., 1999). The material available from two equivalent stratigraphic sections of two wells in distinct palaeogeographic and palaeoceanographic positions offers the opportunity to test some aspects of these hypotheses. The Waren site, situated at the southern margin of the Pompeckj High, was sheltered from terrigenous supply and influenced by intrabasinal upwelling along the submarine high. The Daehre site, situated closer to the southern border of the North German Basin, was influenced by terrigenous supply from the Bohemian Massif and probably occasionally experienced restricted circulation, which may have resulted in a reduced primary productivity. The main palaeoenvironmental differences between the two studied wells were: (1) a significantly higher terrigenous flux in the southern well E Daehre 4/83 (palaeogeographic position close to the coastline of the Bohemian Massif, higher sedimentation rates); (2) a higher marine palaeoproductivity in the northern well Gt Waren 1/81 (palaeogeographic position close to the Pompeckj High, which probably caused intrabasinal upwelling, higher accumulation rates of planktic carbonate producers and primary consumers, presence of calcareous benthic foraminifera indicating enhanced carbon flux (Prokoph et al., 1999). The purpose of this study is to test how far the quantitative assemblage composition of agglutinated foraminifera reflects these palaeoenvironmental differences, and under which environmental conditions "primitive" organically cemented agglutinated foraminifera preferably occurred. In: Hart, M.B., Kaminski, M.A., & Smart, C.W. (eds) Proceedings of the Fifth International Workshop on Agglutinated Foraminifera. Grzybowski Foundation Special Publication, 7,

2 446 Szarek, Klosowska, Prokoph, Kuhnt & Wagner MATERIAL AND METHODS Upper Albian to Lower Cenomanian sedimentary successions were recovered during drilling in Northeast Germany within a depth range of m below surface. The locations and co-ordinates of wells are shown in Figure 1. The Upper Albian sediments were deposited within the eastern part of the epicontinental North German Basin, bordered by the Rhenish-Bohemian Massif to the south and connected with the North Sea Basin to the north (Best et al., 1989). The submarine Pompeckj High bordering the basin to the North may have somewhat limited the open marine circulation into the basin (Diener, 1966). The North German Basin was superimposed on the late Carboniferous Variscan foredeep basin (Betz et al., 1987). Main transgressions into the basin occurred from the western Lower Saxony Basin in the Middle Albian, Late Albian and Early Cenomanian, leading to a continuous pelagic connection from England to the Russian Platform in the Cenomanian. A short regressive phase is documented at the Albian-Cenomanian boundary (Frieg & Kemper, 1989). sample splitter. Assemblages of -300 specimens per sample were picked, then sorted in slides, determined and counted. Scanning electron micrographs were made on a Camscan 44 at the GPI, University of Kiel, and on a Cambridge SEM at the University of Tiibingen. C <lj U -.:;.;!! "... 0:: -.:; :0 kl... <Jl.8.!S SW NE E Daehre 4/83 Gt Waren 1/ F Hamburg lid Schwerin Waren 8 Figure 1. Locations of investigated wells against a simplified background of Albian palaeogeography. (position: E Daehre 4/83: 52 48'N; 'E and Gt Waren 1/81: 53 32'N; 11 42'E). The two studied wells (E Daehre 4/83 and Gt Waren 1/81) consist of alternating multicoloured marlstones and gray marly claystones. Lithology, chronostratigraphy and occurrences of stratigraphically important macrofauna are shown in Figure 2. A detailed lithologic description of the studied interval was given by Prokoph (1994). The sample material examined in the study comprises 49 samples from the Upper Albian sequences of both wells. Samples (approximately 100 cc) were completely disintegrated in diluted buffered hydrogen peroxide or anionic tensides, washed over a 63 /lm sieve and dried at 50 C. Residues were sieved (> 125 /lm, >630 /lm) and randomly divided with a b:3 sandstone calcareous I'I. silty marl Irina Lissa/ - clayey marl Inoceramus anglicus.. Biros E3 marl (j NeohiboLiles minimus Py Pyrite C5 Aucellina gryphaeoides Fe Limonite Q plant remains p Phosph.orite concretions )0 fish remains GL Glauconite I, coal bearing A-H - "bivalve maxima" A conglomera tes Figure 2. Lithology and chronostratigraphy of the Daehre and Waren wells after Prokoph (1997) A-H - "maxima of bivalves". The age assignment was based on calcareous nannofossils from the Eiffellithlls turriseiffelii zone, which indicate a late Albian age according to Sissingh (1977). The biostratigraphic correlation of Albian sediments in Northeast Germany was based on inoce-

3 Upper Albian agglutinated foraminifera, NE Germany 447 ramids: Inoceramus anglicus and Birostrina sllicata (Troeger, 1981), planktic foraminifera: Hedbergella infracretacea, Globigerinelloides bentonensis and Schackoina cenomana (Prokoph, 1994), mass occurrence of aucellines: Aucellina gryphaeoides ("bivalve maxima A-H") and mass occurrence of Neohibolites minimus ("belemnite maximum"). The first occurrence of Eiffellithus turriseiffelii and Globigerinelloides bentonensis are used as markers for the boundary between the early Late and the latest Albian in the North German Basin (Prokoph, 1997). RESULTS Although calcareous benthic foraminifera generally dominate the assemblages, significant numbers of agglutinated foraminifera (up to 50%) are present in both sections (Figure 3). Tests are generally large ( /Lm). Specimens of Spiroplectinata bettenstaedti are generally more than 1500 /Lm in length, and specimens of Dorothia gradata and Spiroplectinata annectens are more than 1000 /Lm in length. Forty-six taxa of agglutinated foraminifera were recognized in the Daehre (45) and Waren (37) samples. The abundance data of common taxa are given in Appendix 1. The average number of agglutinated foraminifera is 22 tests per gram of sediment in the Daehre samples and 26 tests per gram of sediment in the Waren samples. Concurrent fluctuations in abundance were observed in both wells for the following taxa: Falsogaudryinella moesiana, Rhizammina indivisa, Heterantyx cretosa, Tritaxia tricarinata and Textularia spp. Distribution and abundance plots of selected taxa are presented in Appendix 2. Assemblages Five agglutinated assemblages were distinguished within the foraminiferal successions, based on the occurrences of stratigraphically important taxa (e.g., Heterantyx cretosa) and on changes in the abundances, of dominant taxa (e.g., Falsogaudryinella moesiana). The most common taxa are listed in Table 1 in decreasing order. 1. Glomospira charoides Spiroplectinata bettenstaedti Samples: F29-F18 & WF20-WF15 The highest occurrence of this assemblage is marked by the last occurrence of Spiroplectinata bettenstaedti. The abundance of agglutinated foraminifera in this interval is almost constant (representing approximately 35% of the assemblage). At the boundary between the early Late and the latest Albian, the only observed change within the agglutinated assemblage is the FO of Heterantyx cretosa (F19 & WF16), a characteristic form of the Upper Albian. This assemblage is dominated by organically cemented agglutinated foraminifera, which represent epifaunal suspension feeders such as Rhizammina and Rhabdammina, epifaunal deposit feeders such as Ammodiscus semi- infaunal detritivores such as Haplophragmoides, Arenobulimina and Recurvoides (Nagy et al., 1995). 2. Falsogaudryinella moesiana Samples: F17-Fll & WF14-WF12 This assemblage is characterised by a significant increase in the abundance of calcareous cemented species, especially Falsogaudryinella moesiana, which reaches up to 60 specimens in one gram of sediment in the Waren samples. In the Daehre samples the peak of abundance of presumed infaunal species (e.g., Falsogaudryinella moesiana, Heterantyx cretosa) is combined with a decrease in the abundance of tubular, erected, epifaunal morphotypes including Rhizammina and Rhabdammina. Tubular, coiled forms such as Glomospira, Ammodiscus and Spiroplectinata disappear within this interval. 3. Heterantyx cretosa Samples: F10-F9 & WFll-WF8 The base of the assemblage is marked by a bivalve maximum and the top by the last occurrence of Heterantyx cretosa. Foraminiferal abundance significantly increases at the base of this interval, especially for planktonic foraminifera, which show abundance values several times higher than average. Some organically cemented agglutinated genera disappear within this interval in the Waren section (e.g., Haplophragmoides, Ammodiscus, Glomospira) but the abundance of these genera increases at the Daehre site. These changes in foraminiferal distribution may indicate a change to more open marine conditions in the Waren area. 4. Arenobulimina macfadyeni Samples: F8-F5 & WF7-WF4 This assemblage consists of agglutinated organically cemented foraminifera. Arenobulimina macfadyeni, the most common species within this interval, has very constant abundance values, while other species show fluctuations in abundance. Agglutinated foraminiferal abundance reaches 50% of the benthic foraminiferal assemblage in the Daehre section, but at the top of the section the total number of foraminifera decreases. The number of agglutinated specimens per gram of sediment in Waren is generally very low. The genera with calcareous cement are almost absent, and only Falsogaudryinella moesiana and F. alta occur in low numbers. 5. Ammodiscus cretaceus Samples: F4-F1 & WF3-WF1 This assemblage is characterised by the common occurrence of Ammodiscus cretaceus, although the assemblages from the Daehre and Waren sections show significant differences. The Daehre samples are almost barren of benthic foraminifera. Only agglutinated foraminifera with organic cement are present in low numbers within this impoverished assemblage (e.g., Rhizammina, Glomospira, Trochammina). The position of the Waren site seems to provide more

4 448 Szarek, Klosowska, Prokoph, Kuhnt & Wagner E Daehre 4/83 TOC (%) Assemblages 5. Ammodiscus crrillaus 4. Arenobulimina macfodyeni -AUrssonella orycona cre/osa 2. Falsaga.dryinel1a moesiana I. Glomospira cnaroides Spiroplectinatll bettens",edli Gt Waren 1181 TOC(%) Number specimens per 1 gram sediment Plankton &: Benthos Benthos Assemblages 5. Ammodiscus mille- 4. Armobulimina mocfodyeni -Marssonella nrycona 3. Heterantyx c,..tosa -Tritllxia tricarinlltlj 2. Falsaga.dryinel1a moesillna I. Glomospira cluroi.u. -Spiroplectin.'" bettenstjjedli Planktcn o %Planktan o Benthos o 'Y. Calcareous benthos Agglutinated benthos % Agglutinated benthos Figure 3. Abundance and distribution patterns of the five agglutinated foraminiferal assemblages recognized in the samples of the Daehre and Waren wells. favorable conditions for benthic foraminifera. The occurrence of Rhizammina indivisa, Spiroplectinata complanata and S. annectens in the upper part of this site is significant, although the abundance of agglutinated foraminifera decreases towards the top of the interval within the Waren section. DISCUSSION Assemblages from the lower part of the Daehre section are diverse and contain a significant proportion of agglutinated foraminifera. Infaunal and semiinfaunal morphotypes including Pseudobolivina, Verneuilinoides, Bulbobaculites, Spiroplectammina and Recurvoides indicate fairly well-oxygenated conditions with a redox boundary situated well below the sediment water interface (Kuhnt, 1995; Nagy et ai., 1995). Epifaunal suspension and detritus feeders including Ammodiscus, Rhizammina, Rhabdammina and Glomospira are also common. Assemblages from the upper part of the section probably experienced periodically dysoxic conditions, as shown by the dominance of opportunistic forms including Scherochorella, Psammosphaera, Textularia, Ammodiscus and Haplophragmoides, which probably indicate oxygendeficient conditions at the sea floor (Kuhnt, 1995;

5 >-l C "0 g: "0 Daehre assemblages Agglutinated foraminifera Agglutinated foraminifera Waren assemblages Agglutinated foraminifera Agglutinated foraminifera t-' ;l> with organic cement with calcareous cement with organic cement with calcareous cement r' c= I't) I't)... Pl Pl = 5. Ammodiscus cretaceus RIlizammina indivisa Arenoblllimina macfadyeni 5. Ammodiscus cretaceus Ammodiscus cretaceus Spiroplectinata complanata g ::l samples: FI-F4 Ammodiscus eretaceus Eggerellina mariae samples: WFI-WF3 Rhizamnzilla imldivisa Spiroplectinata allllectens :> depth (m): 1377, Haploplzragnzoides sp. depth (m): Rlzabdalllmina sp. Arenobulimina macfadyeni (JQ 0:: (JQ Benthos: 15-35% Reopllax minuta Benthos: 8-15% Eggerellina mariae Qj' - Agg. Benthos: 30-50% Pseudobolivina sp. Agg. Benthos: 10-30% ::l... Haplophragmoides spp. (JQ ::s (JQ... Q.. ;:t. ::l 0 '"" g 0- S I't) 4. Arenobulimina macfadyeni RIlizammina indivisa Falsogaudryinella moesiana 4. Arenobulimina macfadyeni Haplophragmoides sp. Tritaxia tricarinata 2" - Marssonella oxycona Rllabdammina sp. Textularia spp. - Marssollella oxycona Glomospirella gault ina Marssonella oxycolla samples: F5-F8 Ammodiscus eretaceus Arenobulimina macfadyeni samples: WF4-WF7 Pseudosigmoilina antiqua Marssonella turris Pl... depth (m): 1381,7-1385,7 Ammodiscus infimus Dorothia gradata depth (m): Arenobulimina macfadyeni Benthos: 20-50% Recurvoides spp. Marssonella oxycona Benthos: 15-30% Eggerellina mariae... "" 0 ::s Agg. Benthos: 35-50% Psammospharea fusca Falsogaudryinella alta Agg. Benthos: 15-30% Dorothia gradata Pl I't) Pseudosigmoilina alltiqua '"" S... Glomospira charoides -P' Haploplzragmoides spp. "" Z TroclUImmina ro spp.... rrj CJ 3. Heterantyx cretosa Rhizammina indivisa Falsogaudryinel/a moesiana 3. Heterantyx cretosa Rhizammina illdivisa Tritaxia tricarillata Pl I't) ff> - Tritaxia tricarinata Trochammina spp. Arenobulimina macfadyeni - Tritaxia tricarinata Ammodiscus eretaceus Textularia c1zapmani ff> ro 3 samples: F9-FIO Glomospira charoides Eggerellina mariae samples: WF8-WFll Falsogaudryinel/a moesiana g. ::l depth (m): 1385,7-1387,7 Ammodiscus cretaceus Spiroplectinata annectens depth (m): Arenobulimina macfadyeni '< Ben thos: 15% Haploplzragmoides spp. Tritaxia tricarinata Benthos: 15-20% Spiroplectinata complanata PJ (JQ Agg. Benthos: 25-40% Heterantyx cretosa Agg. Benthos: 10-35% Eggere/lina mariae ro Textularia chapmani Dorothia gradata Spiroplectinata annectens Dorotlda gradata Heteralltyx eretosa 2. Falsogaudryinella moesiana Rlzizammina illdivisa Falsogaudryinella moesiana 2. Falsogaudryillella moesiana Rhizammina indivisa Falsogaudryinel/a moesiana samples: Fll-FI7 Glomospira cllaroides Arenobulimina macfadyeni samples: WFI2-WF14 Glomospira charoides Textularia spp. depth (m): 1387,7-1394,7 Ammodiscus cretaceus Eggerel/ina mariae depth (m): Haplophragmoides spp. Spiroplectinata comp/anata Benthos: -50%, max. 60%, min. 38% Haploplzragmoides spp. Heterantyx eretosa Benthos: 20-35% Ammodiscus eretaceus Tritaxia tricarinata Agg. Benthos: -30%, max. 50%, TroclUImmina spp. Marssonella oxycona Agg. Benthos: 12-30% Arenobulimina macfadyeni min. 20% Eggere/lilla mariae Spiroplectinata anllectens Heteralltyx eretosa 1. Glomospira charoides Rhizammina indivisa Falsogaudryinella moesiana 1. Glomospira charoides Glomospira c1laroides Arellobu/imina macfadyeni - Spiroplectinata bettenstaedti RlUIbdammina sp. Tritaxia tricarinata - Spiroplectinata bettenstaedti Rhizanlmina indivisa Falsogaudryinella moesiana samples: F18-F29 Glomospira charoides Textularia clzapmani samples: WFI5-WF20 Rhabdammina sp. Falsogaudryinel/a alta depth (m): 1394,7-1406,7 Rluzbdammina sp. Spiroplectinata complanata depth (m): Haplophragmoides spp. Spiroplectilzata comp/anata Benthos: -30%, max. 40%, min. 20% Glomospira cllaroides Arenobulimina macfadyeni Benthos: 30-35% Ammodiscus eretaceus Dorothia gradata Agg. Benthos: -35%, max. 40%, Dorot/lia filiformis Spiroplectinata annectmns Agg. Benthos: 25-30% Spiropiectillata allnectens min. 30% Haplophragmoides spp. Spiroplectinata bettenstaedti Heterantyx eretosa Trocllammina spp. Marssonella oxycona Saccammina placenta Marssonella turris \0

6 450 Szarek, Klosowska, Prokoph, Kuhnt & Wagner Kaminski et al., 1995). Intervals with high numbers of epifaunal suspension feeders including Rhizammina and Rhabdammina may indicate better oxygenated environments. Agglutinated foraminifera compose approximately 35-50% of the benthic assemblages. This comparatively high abundance may be related to the low organic carbon flux and high terrigenous flux at this site. Agglutinated assemblages from the Waren section exhibit low diversity in some samples, dominance by infaunal morphotypes and fluctuations in abundance, but no remarkable decrease in faunal density. The presence of Ammodiscus cretacells, Glomospira charoides and G. gordialis, in some intervals may indicate oxygen deficiency (Kuhnt, 1995). Intervals of increased diversity and abundance may indicate periods of better oxygenated conditions at the sea-floor. These intervals are also characterized by the reddish colour of the sediments. Agglutinated foraminifera constitute only 10-35% of the benthic assemblage and benthic foraminifera constitute less than 35% of the total foraminiferal assemblage, indicating open marine conditions with "normal" or increased primary productivity. Common occurrences of open marine calcareous species including Gyroidinoides, Berthelina and nodosariids imply pelagic environments (outer shelf to upper bathyal) (Moullade, 1984). CONCLUSIONS The agglutinated foraminiferal assemblages of the Daehre site generally indicate a well oxygenated, oligotrophic shelf environment with significant terrigenous flux. Occasional changes in abundance were probably caused by fluctuating terrigenous input from the Bohemian Massif. Agglutinated foraminiferal assemblages include a large portion of "primitive" forms with organic cement (Rhizammina, Ammodiscus, Glomospira, Haplophragmoides and Trochammina). Oxygen-depleted conditions at the sea floor only occurred during the deposition of the upper part of the section and were probably linked to a sea level rise. The composition of the agglutinated foraminiferal assemblages of the Waren site implies open marine pelagic conditions. "Primitive" organically cemented are generally rare and mainly consist of epifaunal detritus feeding forms, that may have taken advantage of increased organic particle flux during periods of enhanced surface productivity. Agglutinated foraminifera with calcareous cement make up the largest part of the agglutinated assemblages and include suspension feeders (Spiroplectinata) and infaunal detritus feeders (Falsogaudryinella, Tritaxia, Dorothia, Arenobulimina and Eggerellina). ACKNOWLEDGMENTS Dr. Schneider (Geothermie Neubrandenburg GmbH) and Dr. Schretzenmayr (Erdoel/Erdgas Commern Gmbh) are thanked for permission to sample the core material. Dr. Ann Holbourn (Kiel) is thanked for correcting our manuscript. Mrs. Schuldt (Kiel) and Mr. Hiittemann (Tiibingen) are thanked for photographic and SEM work. SEM photographs were taken on the Camscan 44 at the CPI Kiel and on a Cambridge SEM at the University of Tiibingen. TAXONOMIC NOTES The agglutinated foraminifera classification used here follows the taxonomic classification of Loeblich & Tappan (1987) with numerous modifications based on a revision proposed by Meyn & Vespermann (1994) and Ragl (1995). Amlllobaculites subcretaceus Cushman & Alexander, 1930 PI. 2, Fig. 5 Ammobaculites subcretacells Cushman & Alexander, 1930, p. 6, pi. 2, figs Loeblich & Tappan, 1949, p. 251, pi. 46, figs Magniez-Jannin 1975, p. 42, pi. 1, figs Weldlch, 1990, p. 92, pi. 2, figs Coarse grained, agglutinated test with large, initial planispiral stage, followed by a short uniserial part with compressed chambers. Ammodiscus cretaceus (Reuss, 1845) Plate 1, Fig. 5 Operculina cretacea Reuss, 1845, p. 35, pi. 13, figs 64, 65a-b. Ammodiscus cretacells (Reuss).- Cushman, 1946, p. 17, pi. 1, fig Hemleben & Traster, 1984, p. 517, pi. 1, fig. 17. Weidic, 1990, p. 82, pi. 1, fig. 5.-.Reicherter et ai., 19?4, pi. 7a, fig. a.- I-folbourn & KamInski, 1997, p. 35, pi. 4, figs 7-8. Tubular test with relatively large proloculus; broad whorls in a regular planispiral coil, overlapping the previous ones; siliceous, homogeneous or finely agglutinated wall. Ammodiscus infimus Franke, 1936 Plate 1, Figs 6-7 Almodiscus infimus (Strickland).- Franke, 136, p: 15, pl..1, fig. 14a-b.- Ceroch & Nowak, 1984, pi. 1, fig. 11, pi. 5, fig Weidich, 1990, p. 82, pi. 34, figs Reicherter et at., 1994, pi. 7a, figs g, h.- Holbourn & Kaminski, 1997, p. 35, pi. 4, figs Tubular, pi ani spiral test with tendency to uncoil. Ammodiscus peruvianus Berry, 1928 Plate 1, Fig. 8 AmmodisclIs peruvial!us Berry, 1928 p. 342, pi Ragl, 1976, p. 35 pi. 2, fig Gradstem & Berggren, 1981, p. 242, pi. 2, figs Arenobulimina chapmani Cushman, 1936 Plate 3, Fig. 17 Bulimina preslii Reuss.- Chapman, 1892, p. 755, pi. 12, fig. 4. Arenobullmina chapmani Cus!,man, 1936,,P. 26, pi: 4, fig. 7. Neagu, 1965, p. 10, pi. 2, fig. 9.- Magmez-Jannm, 1975, p. 79, pl. 7, figs Carter & Hart, 1977,,P. 15, pi. 1, fg. 4. Barnard & Banner, 1981, p. 404, pi. 4, fig. 1, pi. 7, flg. l. Frieg & Price, 1982, p. 59, pi. 2.2, figs h, i, j, m. Trochospiral test, final whorl has five chambers and comprises more than half the length of the test, sutures between chambers depressed, arch-shaped

7 Upper Albian agglutinated foraminifera, NE Germany 451 aperture, surrounded by a narrow lip; surface of the test rough, built of quartz grains with large amount of calcareous cement. Arenobulimina frankei Cushman, 1936 Plate 3, Fig. 14 Arenobulimina frankei Cushman, 1936, p. 27, pi. 4, fig. 5a-b. Gawor-Biedowa, 1969, p. 84, pi. 5, figs 4-5; pi. 7, figs 6, 7ab, 8a-b; text-figs Price, 1977, p. 508, pi. 59, figs 5-6, 9. Large, subconical, trochospiral test; last whorl increasing rapidly in width; chambers inflated, separated by slightly depressed sutures; interiomarginal arch-shaped aperture; coarse agglutinated wall with large amount of carbonate material. Arenobulimina ex gr. macfadyeni Cushman, 1936 Plate 3, Fig. 16 Arenobulimina macfadyeni Cushma.n, 1936, p. 26, p!. 4, fig. ab.- Neagu, 1965, p. 10, pi. 2, figs Magmez-Jannm, 1975, p. 78, pi. 7, figs Barnard & Banner, 1981, p. 402, pi. 2,Jig. 4-7, pi. 6, fg Frieg & Price, 198, p. 61: pi. 2.2, figs k, I, n.- Weldlch, 1990, p. 108, pi. 37, figs 1-2, pi. 38, figs In this group we included forms having subconical, trochospiral test, gradually increasing in width, inflated chambers, separated by slightly depressed sutures; interiomarginal arch-shaped aperture, smooth, fine-grained wall, mainly agglutinated quartz grains. Arenobulimina d. obliqua Price, 1977 Plate 3, Fig. 15 Arenobulimina d. obliqua (d'orbigny).- Price, 1977, p. 510, pi. 59, fig Frieg & Price, 1982, p. 53, pi. 2.1, fig. h. Small, trochospiral test; last whorl comprises twothirds of the length of the test, elongated chambers, usually indistinct sutures; interiomarginal archshaped aperture, smooth, fine grained wall with large amount of carbonate material. Bigenerina sp. Plate 2, Fig. 18 Biserial slightly twisted initial part, composed of five sets of chambers, followed by two sets of larger, globular chambers and finally short semi-uniserial part; fine-grained wall. Bulbobaculites problematicus (Neagu, 1962) Plate 2, Fig. 6 Ammobaculites agglutinans (d'orbigny) ssp. problematiclls Neagu, 1962, p. 61, pi. 2, figs Ammdbaculites problematicus Neagu.- Geroch & Nowak, 1984, pi. 1, figs 17-18; pi. 6, fig. 23. HaplopJiragmium lueckei Cushman & Hedberg.- Moullade et al., 1988, p. 363, pi. 3, figs 1-6. Bulbobaculztes problematicus (Neagu).- Kuhnt & Kaminski, 1990, pp , text-figs 5,5a; pi. 4, fig. a-h.- Bubfk, 1995, p. 80, pi. 12, figs 2, 17. Caudammina ovulum (Grzybowski, 1896) Reophax ovulum Grzy:bowski, 1896, p. 276, pi. 8, figs Hormosina ovulum (Grzybowski).- GerocFt, 1960, pi. 2, fig Bubfk, 1995, p. 82. Caudammina crass a (Geroch, 1966) Hormosina ovul"! crassa Geroch, 1966, p. 463, 6/19, 21-26; 7/ WeldICh, 1990, p. 84, pi. 33, fig. 25. Hormosina crassa Geroch.-.Kuhnt, 1?90, p. 31, pi. 2, fig. 5. Bubfk, 1995, p. 82, pi. 2, figs 12-13, d. p1. 9, fig. 1. Caudammina crassa (Geroch).- Holbourn & Kaminski, 1997, p. 38, pi. 7, figs 4-9. Caudammina praecaudata (Hanzlikova, 1973) Pelosina praecaudata Hanzlfkova, 1973, p. 137, pi. 2, fig. 9. Hormosina praecaudata (Hanzlfkova).-Weidich, 1990, p. 84, pi. 1, fig. 13; pi. 33, figs Dorothia filiformis (Berthelin, 1880) Plate 4, Fig. 1 Gaudryina filiform is Berthelin, 1880, p. 25, pi. 1, fig. 8a-d. Geroch, 1966, fig. 12 (11,12).- Haig, 1980, p. 120, pi. 8, figs 1-5; pi. 11, fig. 2. Dorotliia filiformis (Berthelin).- Cushman, 1937, p. 73, pi. 8, fgs M.agniez-Jannin, 1975, p. 83 pi. 8, figs 1-2; texthg Weldich, 1990, p. 109, pi. 36, fig. 22. Gaudryinopsis filiformis (Berthelin).- Haig & Lynch, 1993, p Elongated, narrow test, triserial at first, becoming biserial; last chambers usually flattened; interiomarginal apertural slit on the last chamber; medium to fine-grained wall. Dorothia gradata (Berthelin, 1880) Plate 4, Figs 2-5 Gaud,ryina gradata Bertheln, 1880,y. 24, pi. 1, fig. 6a-c. Halg, 1980, p. 121, pi. 8, figs 17-18, pi. 11, fig. 1. Dorotliia gradata (Berth.elin).- Gawor-Biedowa, 1,972, p. 29, pi. 2, fig. 7a-b.- Dalley, 1973, p. 48, pi. 3, flj:;. 13a-b. Magniez-Jannin, 1975, p. 86, pi. 8, figs 3-6; text-fig. 38. Gaudryinopsis gradata (Berthelm).- Kaig & Lynch, 1993, p. 346, pi. 1, fig. 21. Elongated finely agglutinated test with initial triserial stage, triangular in section, then becoming biserial with inflated chambers; the aperture is an arch at the base of last chamber. Most of the specimens from site M2-19 have compressed, flattened or curved tests such as in Plate 4, Figs 4 & 5. Eggerellina mariae Ten Dam, 1950 Plate 4, Figs Eggerellina mariae Ten Dam, 1950, p. 15, pi. 1, fig. 17a-e. Magniez-Jannin, 1975, p. 94, pi. 6, figs Weidich, 1990, p Falsogaudryinella ex gr. moesiana (Neagu, 1965) Plate 3, Figs 9-13 In this group we lumped elongated, cone-shaped forms with tricarinate outline, unusually large last three chambers and a terminal, elongated, slit-like aperture. We used a very broad concept for these forms, which include representatives of Falsogaudryinella moesiana and F. alta. Specimens closely resembling FalsogaudryineUa subrotunda (Ten Dam, 1950) from the Upper Albian of the Netherlands are also included in the variability of our population. Glomospira charoides Gones & Parker, 1860)

8 452 Szarek, Klosowska, Prokoph, Kuhnt & Wagner Plate 1, Figs 9-10 Trochammina squamilta var. charoides Jones & Parker, 1860, p Glo11lospira charoides (Jones & Parker).- Berggren & Kaminski, 1990, pi. 1, fig. 2 (lectotype).- Reicherter et ill., 1994, pi. 7a, fig. 0.- Holoourn & Kaminski, 1997, p. 36, pi. 5, figs Glol11ospiril cllilroides corollil Cushman & Jarvis.- Weidich, 1990, p. 83, pi. 34, figs 8-9. Glomospira gordialis Gones & Parker, 1860) Plate 1, Fig. 11 TroclwIl111lillil squilmiltil var. gordiillis Jones & Parker, 1860, p Glo11lospira ordiillis Oones & Parker).- Bartenstein & Brand, 1951, pi. I, figs Berggren & Kaminski, 1990, pi. 1, fig. 1 (lectotype).- Reicherter et al., 1994, pi. 7a, fig. c. Holbourn & Kaminski, 1997, p. 36, pi. 5, fig. 5. Glomospira irregularis (Grzybowski, 1898) Ammodiscus irregulilris Grzybowski, 1898, p. 285, pi. 11, figs 2-3. Glomospiril irregularis (Grzybowski).- Geroch, 1966, pi. 8, figs Bartens.tein, 1974, pi. 2, figs eicherter et ill., 1994, pi. 7a, fig. d.- Holbourn & Kammski, 1997, p. 36, pi. 6, figs 1-2. Glomospirella gaultina (Berthelin, 1880) Plate 1, Fig. 12 Ammodiscus gilultillus Berthelin, 1880, p. 19, pi. 1, fig. 3a-b. Bartenstein et ill., 1966, p. 140, pi. 1, fig. 29. GlO11lospirellil gilultilza (Berthelin).- Magniez-Jannin, 1975, p. 26, pi. 1, figs Krasheninnikov & Pflaumann, 1978, p. 568, pi. 2, fig. 5.- Haig & Lynch, 1993, p. 346, pi. 1, fig. 6. Holoourn & Kaminski, 1997, p. 37, pi. 6, figs 4-6. Test glomospirally coiled in the initial stage, later planispiral; finely agglutinated. Haplophragmium aequale (Roemer, 1841) Plate 2, Figs 7-8 Spirolilla ileqllillis Roemer, 1841, p. 98, pi. 15, fig. 27. Hilplophragmilll.ll ileqllille Chapman, 18.92, p. 32?, fl. 6,.figs Bartenstem, 1952, p. 325, pi. 1, figs 2, 11, P. 2, figs 17-26; pi. 3, figs 1-6; pi. 6, figs 6-8; pi. 7, figs Meyn & Vespermann, 1994, p. 72, pi. 1, figs 9-13; pi. 2, figs 1-7. Elongated, robust test with small, initial planispiral coil; long uniserial part with two to four inflated chambers; last chamber has short neck with simple aperture. Haplophragmoides aff. bulloides (Beissel, 1891) Plate 2, Figs 1-2 Haplophragmium bulloides Beissel, 1891, pi. 2, figs 1-3; pi. 4, figs Haplophrilgmoides blilloides (Beissel).- Neagu, 1970, pi. 3, figs 3-4. Hilplophragmoides aff. blilloides (Beissel).- Holbourn & Kaminski, 1997, p. 40, pi. 9, figs Planispiral, tightly involute and inflated test; slightly depressed sutures; slit like aperture at the base of the last chamber. Our specimens are very similar to Haplophragmoides aff. bulloides (Beissel, 1891) as figured by Holbourn & Kaminski (1997), to Haplophragmoides suborbiclilaris (Grzybowski, 1896) as figured by Geroch (1960, pi. 5, fig. 1). In most of specimens, the aperture is indeterminate, but better pre- served forms appear to have a long, slit like aperture, indicating that the species belongs to the genus Haplophragmoides. Haplophragmoides concavlis (Chapman, 1892) Plate 1, Fig. 16 Troc/w11lmi1lil COIlCilVil Chapman, 1892, p. 327, pi. 6, fig. 14a-b. Haplophragmoides concilvus (Chapman).- Bartenstein & Brand, 1951, pi. 1, figs Weidich, 1990, p. 87, pi. 7, figs 2, 11, 19; pi. 36, figs Reicherter et al., 1994, pl 7b, figs a-d.- Holbourn & Kaminski, 1997, p. 40, pi. 9, figs 3, 7. Slightly evolute and planispiral test; five to six usually compressed chambers; straight depressed sutures; medium to finely agglutinated wall; aperture usually indeterminate. Haplophragmoides nonioninoides (Reuss, 1863) Plate 1, Fig. 15 HaplophragmillHl nonioninoides Reuss, 1863, p. 30, pi. 1, fig. 8a-b. HaploplJrac;:moides nonioninoides (Reuss).- Bartenstein & BolIi 1986, p. 952, pi. 2, figs Reicherter et ill., 1994, pi. 7a, fig. I, m, n.- Meyn & Vespermann, 1994, p. 70, pi. 1, figs 1-8. Planispiral, involute and inflated test, slightly depressed sutures; aperture is a long slit at the base of the last chamber; finely agglutinated wall. Heterantyx cretosa (Ten Dam, 1947) Plate 2, Figs Spiroplectilmmina rectilngularis Ten Dam, 1947, p. 25, pi. 1, fig.2a-b. Hetermltyx cretosa (Ten Dam).- Haig, 1992, pi. 1, figs Holbourn & Kaminski, 1997, p. 45, pi. 15, figs 1-5. Test elongated and flattened; initially planispirally coiled; adult stage biserial, composed of ten to twelve pairs of chambers, rectangular in cross-section, separated by elevated, sutures. Lagenammina sp. Plate 1, Fig. 3 Unilocular form, laterally compressed with broad, elongated neck; medium grained agglutinated wall. This form resembles the species Lagenammina lagenoides (Crespin, 1953) in Holbourn & Kaminski (1997, p. 33, pi. 3, figs 5,7), but differs in the broader base of neck. Lituotllba lituiformis (Brady, 1879) Plate 2, Fig. 9 Trochmmninil Iituiformis Brady, 1879, p. 59, pi. 5, fig. 16. Liluotllb lituiformis (Brady).- Kaminski et ill:, 1988, p. 190, pi. 4, figs Kuhnt, 1990, p. 318, pi. 1, figs Marssonella oxycona (Reuss, 1860) Plate 4, Figs 6-7 Gillldryinil OXyCOllil Reuss, 1860, p. 229, pi. 12, fig. 3a-c. Milrssollellil OXYCOl1il (Reuss).- Cushman, 1933, p. 36, pi. 4, fig Belford, 1960, p. 16, pi. 4, fig. 1.- Desai & Banner, 1987, p. 20, pi. 3, figs a-i (lectotype).- Haig & Lynch, 1993, p. 346, pi. 1, fig. 20.

9 Upper Albian agglutinated foraminifera, NE Germany 453 Dorotlli oxycona (Reuss).- Trujilo, 1960, p. 309, pi. 44, fig. 5.- Shter, 1968, p. 50, pi. 3, flg Geroch & NowaK, 1984, pi. 4, fig. 14. Marssonella tllrris (d'orbigny, 1839) Plate 4, Figs 8-9 Textularia turris d'orbigny, 1839, p. 46, pi. 4, figs Marssonella tllris (d'orbigny).- Barnard, 1963, p. 42, text-fig. 2a-h. Dorotllia tllrris (d'orbigny).- Hanzlfkova, 1973, p. 161, pi. 6, figs ? Marssonella ozawai Cushman, 1936, p. 43, pi. 4, fig. loa-b.- Hart et al., 1989, p. 176, pi. 7.2, figs 8-9. Strongly elongated, slender, conical test; initial trochospirally coiled part, composed of three to five chambers; biserial part composed of seven to eight sets of wide chambers; apertural surface flat or slightly depressed; elongated slit aperture, situated at the base of last chamber; fine-grained wall. This form is very similar to Marssonella ozawai Cushman (1936), until an examination of the holotype is possible, we regard M. ozawai as a junior synonym of M. tllrris (d'orbigny, 1839). Psammosplzaera fusca Schulze, 1875 Psammospllaera fllsca Schulze, 1875, p. 113, pi. 2, fig. 8a-f. Psammospllaera fusca Schulze.- Weldich, 1990, p. 81, pi. 1, fig. 11, pi. 33, fig Bender, 1995, p. 48, pi. 1, fig. 14. Kolbourn & Kaminski, 1997, p. 32, pi. 2, figs 4-5. Unilocular, spherical test; coarse to medium grained agglutinated wall; apertures rarely visible. Pselldobolivina spp. Plate 2, Fig. 17 We included forms with distinct initial biserial part, followed by flattened and rounded chambers in a loose biserial series, gradually increasing in size; finely agglutinated wall. Recllrvoides imperfectus Hanzlfkova, 1953 Plate 2, Fig. 3 ReCllrvoides imperfectlls Hanzlikova, 1953, pi. 9, fig. 1a-d. Geroch & Nowak, 1984, pi. 2, fig. 19; pi. 6, fig. 1.- Bubfk, 1995, p. 85, l?1. 4, fig: 8, d. 9; pi. 1.3, fig. 12:- Weidich, 1990, p. 89, pi. 5, figs 1-15, pi. 6, fig. 24, pi. 35, figs Recllrvoides sp. Plate 2, Fig. 4 These forms are generally rare and diagenetically strongly compressed, some with radially elongated chambers; finely agglutinated wall; usually indeterminate aperture. Rlzabdammina spp. Plate 1, Fig. 1 In this group we included all large, fairly straight, unbranched, tubular forms with a thick wall. Rlzizammina indivisa Brady, 1884 Plate 1, Fig. 2 Rhizammina indivisa B.rady, 1884, p.. 277, pi. 29, figs 5, 7. Geroch, 1966, pi. 1, figs Weldlch, 1990, p. 79, pi. 33, figs 4-6. Test unbranching with usually thin, flattened and distorted wall. Saccammina alexanderi (Loeblich & Tappan, 1950) Proteonina alexanderi Loeblich & Tappan, 1950, p. 5, pi. 1, fig. la-b. SaCCaml7lilla alexander; (Loeblich & Tappan).- Eicher & Worstell, 1970, p. 280, pi. 1, fig. 7.- Weidich, 1990, p. 81, pi. 2, fig. 7; pi. 34, figs Saccammina placenta (Grzybowski, 1898) Reophax placenta Grzybowski, 1898, p. 276, pi. 10, figs Saccammina placenta (Grzybowski).- Geroch, 1960, p. 37, pi. 2, figs -?- Gradstein & Berggren, 1981, p. 241, pi. 2, fig. 3.- Weldlch, 1990, p. 81. Saccammina splzaerica Sars, 1872 Plate 1, Fig. 4 Saccammina sphaerica Sars, 1872, p Moullade et al., 1988, p. 366, pi. 2, fig Kuhnt, 1990, p. 324, pi. 2, figs 8-9. Scheroclzorella minuta (Tappan, 1940) Plate 1, Fig. 13 Haplosticlze D1 Hecht, 1938, pi. 4a, figs 4-8. Reoplzax l11;nlla Tappan, 1940, p. 94, pi. 14, fig. 4a-b; 1943, p. 480, pi. 77, fig. 4a-b. Reop17ax millutus (Tappan).- Ten Dam, 1950, p. 6, pi. 1, fig. 3.- Bartenstein & Bettenstaedt, 1962, p. 282, pi. 39, fig. 16. Weidich, 1990, p. 85, pi. 34, fig. 19. Sc1leroc1lOrella minllta (Tappan, 1940).- Geroch & Kaminski, 1995, p. 119, pi. 1, fig. 3, text-fig. 4. Spiroplectammina spp. We included in this group all varieties of Spiroplectammina, which have elongated and flattened biserial test with small initial, planispiral part, no dentate outline and finely agglutinated wall. Spiroplectinata annectens (Parker & Jones, 1863) Plate 3, Figs 1-3 Textularia mll1ectells Parker & Jones, 1863, p. 92, text-fig. 1. Spiroplectinata unnectells (Parker & Jones).- Ten Dam, 1950, p. 13, pi. 1, fig Bartenstein & Bettenstaedt, 1962, p. 293, text-fig Weidich, 1990, p. 104, pi. 11, figs 9-10; pi. 36, fig Haig & Lynch, 1993, p. 347, pi. 2, figs 2-5. Spiroplectinata bettenstaedti Grabert, 1959 Plate 3, Fig. 5 Spiroplectinata bettcllstuedti Grabert, 1959, p. 15, EI. 1, figs 14-15; pi. 2, figs 42-45; pi. 3, figs Haig & ynch, 1993, p. 347, pi. 2, figs 6, Large, very elongated, narrow test; agglutinated material contains large amount of pyritic particles. Probably a phylogenetic endmember of the Spiroplectinata annectens - S. complanata - S. bettenstaedti lineage. Spiroplectinata c011lplanata (Reuss, 1860) Plate 3, Fig. 4 Proroporus complmzatlls Reuss, 1860, p. 231, pi. 12, fig. 5. Spiroplectinutu complallata (Reuss).- Grabert, 1959, p. 14, pi. 1, fig. 13; pi. 2, figs 39-41; pi. 3, figs Bartenstein & Bettenstaedt, 1962, p. 293, text-fig. 23; pi. 39, fig Haig & Lynch, 1993, p. 347, pi. 2, figs 7, 13.

10 454 Szarek, Klosowska, Prokoph, Kuhnt & Wagner We follow the concept of Grabert (1959) and Bartenstein & Bettenstaedt (1962) who regard S. annectens and S. complanata as different species. This is in contrast to Ten Dam (1950), who regarded S. complanata as the megalospheric generation of S. annectens. Subreophax scalaris (Grzybowski, 1896) Plate 1, Fig. 14 Reophax guttifera Brady var. scala ria, Grzybowski, 1896, p. 277, pi. 8, fig. 26. Subreophax scalaria (Grzybowski).- Kaminski et ai., 1988, p. 187, pi. 2, figs Subreophax scalaris (Grzybowski).- Kuhnt, 1990, p. 326, pi. 3, figs 4-5. Textularia chapmani Lalicker, 1935 Plate 2, Figs Textularia chapmani Lalicker, 1935, p. 13, pi. 2, figs Ten I?am, 1950, p. 1, pi. 1, Jig Neagu, 1965, P: 5, pi. 1, fig Magmez-Jannm, 1975, p. 54, pi. 3, figs 1-9. Weidich, 1990, p. 97, pi. 12, figs 15-17; pi. 37, figs 17, Textularia spp. Plate 2, Fig. 16 We included under this name all typical textulariid forms. Textulariopsis rioensis (Carsey, 1926) Plate 2, Fig. 15 Textularia rioe.nsis Carsey, 1?6, p. 24, pi. 7, fig. 2.-.Haig, 1981, pi. 1, figs WeldIch, 1990, p. 98, p[ 12, figs 18-19,21; pi. 37, figs Textulariopsis rioensis (Carsey).- Loeblich & Tappan, 1982, p. 67, pi. 2, figs Textu7ariopsls sp. B.- Haig & Lynch, 1993, p. 347, pi. 2, fig. 17. Tritaxia pyramidata Reuss, 1863 Plate 3, Figs 6-7 Tritaxia pyramidata Reuss, 1863, p. 32, pi. 1, fig. 9a-c. Magniez-Jannin, 1975, p. 71, pi. 5, figs (with synonymy).- Weidich, 1990, p. 10.5, pi. 12, fig Meyn & Vespermann, 1994, p. 76, pl 4, figs 3-8. Tritaxia tricarinata (Reuss, 1845) Plate 3, Fig. 8 Textularia tricarinata Reuss, 1845, p. 39, pi. 8, fig. 60. Tritaxia tricarinata (Reuss).- Risch, 1971, p. 34, pi. 1, figs Weidich, 1990, p. 106, pi. 12, fig. 26. Trochammina spp. Plate 2, Fig. 10 We included in this group all compressed forms with trochospiral, planoconvex test; with five to eight chambers in the last whorl, separated by slightly depressed sutures. Verneuilinoides neocomiensis (Mjatliuk, 1939) Plate 2, Fig. 19 Verneuilina neocomiensis Mjatliuk, 1939, pi. 1, figs Verneuilinoides subfiliformis Bartenstein,.1952, pi. 12a figs I ll.- Geroch & Nowak, 1984, pi. 2, fig. 5, pi. 7, fig. 15. Weidich, 1990, p. 108, pi. 32, fig. 9. Verneuilinoides neocomiensis (Mjatliuk).- Kuhnt, 1995, pi. 4, figs Holbourn & Kaminski, 1997, p. 53, pi. 23, figs 7-9. Elongated triserial arrangement of chambers throughout, about seven sets of chambers; interiomarginal arched aperture; finely agglutinated wall. According to Mjatliuk (1939), Verneuilinoides neocomiensis has seven to ten sets of chambers. Vernellilinoides subfiliformis described by Bartenstein (1952), has twelve sets of chambers. According to Maync (1973), both species V. neocomiensis and V. subfiliformis are synonymous. REFERENCES Best, G., Elstner, F. & Kemper, E Jilngere Unterkreide und Cenoman in der sildlichen Nordsee und in Schleswig-Holstein. In: Kemper, E. (ed.), Alb und Untercenoman in Mittel- und Nordeuropa. Geologisches Jahrbuch, A 113, Betz, D., Filhrer, F., Greiner, G. & Plein, E Evolution of the Lower Saxony Basin. In: Ziegler, A. (ed.), Compressional Intra-Plate Deformations in the Alpine Foreland. Tectonophysics, 137, Diener, I Stratigraphisches Korrelationsschema filr die Kreide der Deutschen Demokratischen Republik und angrenzender Gebiete. Abhandlungen des zentralen geologischen Instituts der DDR, 5, Frie, C. & Kemper, E Mikropalaontologische Ghederung una Abgrenzung von Oberalb und Untercenoman in Nordwestdeutschland. In: Kemper, E. (ed.), Alb und Untercenoman in Mittel- und Nordeuropa. Geologisches Jahrbuch, A 113, Moullade, M Interet des petits foraminiferes benthiques profonds pour la stratigraphie et l'analyse des paleoenvironnements oceaniques mesozoiques. In: Oertli, H.J. (ed.), Benthos '83, 2nd International Symposium on Benthic Foraminifera (Pall, April 1983), Nagy, J., Gradstein, F.M., Kaminski, M.A. & Holbourn, AE.L Foraminiferal morphogroups, paleoenvironments and new taxa from Jurassic and Cretaceous strata of Thakkhola, Nepal. In: M.A Kaminski, S. Geroch, & M.A Gasinski (eds), Proceedings of the Fourth International Workshop on Agglutinated Foraminifera. Grzybowski Foundation Specia1 Publication 3, Kaminski, M.A, Boersma, A, Tyszka, J. & Holbourn, AE.L Response of deep-water agglutinated foraminifera to dysoxic conitios in the California Borderl.and.Basins. In: M.A. Kammskl, S. Geroch, & M.A Gasmski (eds), Proceedings of the Fourth InternationaL Workshop on Agglutinated Foraminifera. Grzybowski Foundation Special Publication 3, Kemper, E Das Klima der Kreide-Zeit. GeoLogisches Jahrbuch, A 96, Kuhnt, W. & Wiedmann, J Cenomanian-Turonian Source Rocks: Paleobiogeographic and Paleoenvironmental Aspects. In: Huc, A.Y. (ed.), Pale08eography, PaloeC/imate, and Source Rocks. AAPG Studies m Geology 40, Prokoph, A Zyklische Sedimentation im Oberalb des Norddeutschen Beckens. Tiibinger Geowissenschaftliche Arbeiten, A 19, Prokoph, A Paleoenvironment and stratigral?hy of late Albian - early Cenomanian planktonic forammifera from NE-Germany. Freiberger Forschungshefte, C 468, Prokoph, A, Szarek, R., Klosowska, B. & Kuhnt, W Albian Benthic Foraminiferal Biofacies and Paleogeography of Northeast Germany. Neues Jahrbuch fiir Geologie IIndPaliiont%gie, Abhandillngen, 212, Sis singh, W Biostratigraphy of Cretaceous calcareous nannoplankton. Geologij en Mijnbouw, 56 (1), Troeger, K.A German Democratic Republic. In: Reyment, R.A. & Bengtson, P. (eds), Aspects of Mid Cretaceous Regional GeoLogy, 1-28.

11 Upper Albian agglutinated foraminifera, NE Germany 455 TAXONOMY REFERENCES Barnard, T The morphology and development of species of Marssonella, and Pseudotextulariella from the chalk of England. Palaeontology, 6, Barnard, T. & Banner, F.T The Ataxophragmiidae of England: Part I, Albian-Cenomanian Arenobulimina and Crenaverneuilina. Revista Espanola de Micropaleontolog{a, 12 (3), Bartenstein, H Taxonomische Bemerkungen zu den Ammobaculites, Hae1ophragmium, Lituola and verwand ten Gattungen (for.). Senckenbergiana,33 (4-6), Bartenstein, H Upper Jurassic-Lower Cretaceous primitive arenaceous foraminifera from DSDP Sites 259 and 261, Eastern Indian Ocean. In: Veevers, J.J., Heirtzler, J.R et ai., Initial Reports of the Deep Sea Drilling Project, 27, (Washington, D.C.), Bartenstein, H. & Bettenstaedt, F Marine Unterkreide (Boreal and Tethys). In: Simon, W. & Bartenstein, H. (eds), Leitfossilen der Mikropaliiontologie. Gebruder Borntraeger, Berlin, Bartenstein, H. & Bolli, H.M The foraminifera in the Lower Cretaceous of Trinidad, W.l Part 5: Maridale Formation. Upper Part: Hedbergella rohri zone. Eclogae Geologicae Helvetiae, 79 (3), Bartenstein, H. & Brand, E Mikropalaeontologische Untersuchungen zur Stratigraphie des nordwestdeutschen Vafendis. Abhandlungen der Senckenbergischen Natllrforschenden Gesellschaft, 485, Bartenstein, H., Bettenstaedt, F. & Bolli, H.M Die Foraminiferen der Unterkreide von Trinidad, B.W.1. Part 2. Maridale Formation (Typlokalitat). Eclogae Geologicae Helvetiae, 59 (1), Beissel, I Die Foraminiferen der Aachener Kreide. Abhandlungen der Koniglichen Preussischen Geologischen Landesanstalt, Neue Folge, 3,1-78. Belford, D.J Upper Cretaceous foraminifera from the Toolona Calcilutite and Gingin Chalk, Western Australia. Bureau of Mineral Resources, Geology and Geophysics, Bulletin, 57, Bender, H Test structure and classification in agglutinated foraminifera. In: M.A. Kaminski, S. Geroch, & M.A Gasinski (eds), Proceedings of the Fourth International Workshop on Asglutinated Foraminifera. Grzybowski Foundation SpeCIal Publication 3, Berggren, W.A. & Kaminski, M.A Abyssal agglutinates: back to basics. In: Hemleben, c., Kaminski, M.A, Kuhnt, W. & Scott, D.B. (eds), Paleoecology, Biostratigraphy, Paleoceanography and Taxonomy of Agglutinated Foraminifera. NATO ASI Series, C 327, Kluwer Academic Publishers, Berry, E.W The smaller foraminifera of the middle Lobitos shales of northwestern Peru. Eclogae Geologicae Helvetiae, 21, Berthelin, G Memoire sur les foraminiferes fossiles de l'etage Albien de Montcley (Doubs). Memoires de la Societe Geologique de France, 3 (1), Brady, H.B Notes on some of the reticularian Rhizopoda of the "Challenger" Expedition: Part 1. On new or little known arenaceous types. Quarterly Journal of Microscopical Sciences 19, Brady, H.B Report on the foraminifera dredged by HMS Challenger, during the years , Rept. Scientific Results Explor. Voyage HMS Challenger. Zoology, 9, Bubik, M Cretaceous to Paleogene agglutinated foraminifera o.f the Bile Karpatyyni.t (West Carpathians, Czech Republic). In: M.A. Kammskl, S. Geroch, & M.A Gasinski (eds), Proceedings of the Fourth International Workshop on Agglutinatea Foraminifera. Grzybowski Foundation Special Publication 3, Carsey, D.O Foraminifera of the Cretaceous of Central Texas. Texas University Bulletin, 2612, Carter, D.J. & Hart, M.B Aspects of mid-cretaceous micropaleontology. Bulletin of the British Museum of Natural History, (Geology), 29 (1), Chapman, F The Foraminifera of the Gault of Folkstone. Journal of the Royal Microscopical Society, 2, ; 3, Crespin, I Lower Cretaceous foraminifera from the Great Artesian Basin, Australia. Contributions from the Cushman Foundation for Foraminiferal Research, 4 (1), Cushman, J.A Some new foraminiferal genera. Contributions from the Cushman Laboratory for Foraminiferal Research, 9 (2), Cushman, J.A New genera and species of the families Verneuilinidae and Valvulinidae and of the subfamily Virgulininae. Cushman Laboratory for Foraminiferal Research, Special Contribution 6, Cushman, J.A A Monograph of the foraminiferal Family Verneuilinidae. Cushman Laboratory for Foramzniferal Research, Special Contribution 7, Cushman, J.A Upper Cretaceous Foraminifera of the Gulf Coastal Region of the United States and adjacent areas. Professional Paper United States Geological Survey, 206, Cushman, J.A & Alexander, c Some Vaginulinas and other foraminifera from the Lower Cretaceous of Texas. Contributions from the Cushman Laboratory for Foraminiferal Research, 6 (1),1-10. Dailey, D.H Early Cretaceous foraminifera from the Budden Canyon Formation, northwestern Sacramento Valley, California. University of California Publications in Geological Sciences, 106, Desai, D. & Banner, F.T The evolution of Early Cretaceous Dorothiinae (Foraminiferida). Journal of Micropalaeontology, 6 (2), Eicher, D.L. & Worstell, J? Cenomanian and Turonian foraminifera from the Great Plains, United States. Micropaleontology, 16 (3), Franke, A Die Foraminiferen des deutschen Lias. Abhandlungen der Preussischen Geologischen Landesanstalt, Neue Folge, 169, Frieg, C. & Price, RJ The subgeneric classification of Arenobulimina. In: Banner, F.T. & Lord, AR (eds), Aspects of Micropalaeontology, Gawor-Biedowa, E The genus Arenobulimina Cushman from the Upper Albian and Cenomanian of the Polish lowlands. Rocznik Polskiego Towarzystwa Geologicznego, 39, Gawor-Biedowa, E The Albian, Cenomanian and Turonian foraminifers of Poland and their stratigraphic importance. Acta Palaeontologica Polonica, 17, Geroch, S Microfaunal assemblages from the Cretaceous and Paleogene Silesian unit in the Beskid Alaski Mts. (Silesian Carpathians). Biuletyn Instytutu Geologicznego, 153, Geroch, S Male otwornice dolnej kredy serii Slqskiej w Polskich Karpatach (Lower Cretaceous Small Foraminifera of the Silesian Series, Polish Carpathians). Rocznik Polskiego Towarzystwa Geologicznego, 36, Geroch, S. & Kaminski, M An emendation of some Cretaceous species of "Reophax" (Foraminiferida) from northwest Europe and Poland. In: M.A. Kaminski, S. Geroch, & M.A. Gasinski (eds), Proceedings of the Fourth International Workshop on Agglutinated Foraminifera. Grzybowski Foundation Special Publication 3, Geroch, S. & Nowak, W Proposal of zonation for the Late Tithonian-Late Eocene, based upon arenaceous foraminifera from the Outer Carpathians, Poland. In: H.J. OertH (ed.), Benthos '83, Proceedings of the 2nd International Symosium on Benthic Foraminfera, (Pau, April, 1983), Grabert, B Phylogenetische Untersuchungen an Gaudryina und Spiroplectznata (Foram.) besonders aus dem nordwestdeutschen Apt und Alb. Abhandlungen der Senckenbergischen NaturJorschenden Gesellschaft, 498,1-71. Gradstein, F.M. & Berggren, W.A Flysch-type agglutinated foraminiiera and the Maestnchtian to Paleogene history of the Labrador and North seas. Marine Micropaleontology, 6,

12 456 Szarek, Klosowska, Prokoph, Kuhnt & Wagner Grzybowski, J Otwornice czerwonych il6w z Wadowic. Rozprawy WydziaJu Matematyczllo Przyrodniczego, Akade11lla UmieftnoEci w Krakowie, 2 (30), Grzybowski, J Otwornice poklad6w naftonoenych okolicy Krosna. Rozprawy Wydzialu Matematyczllo Przyrodlliczego, Akademia UmieftnoEci w Krakowie, 2 (33), Haig, D.W Early Cretaceous Textulariine foraminiferids from Queensland. Palaeontographica, A, 70, 1-] 38. Haig, D.W Mid-Cretaceous foraminiferids from the Wahgi Valley, Central Highlands of Papua-New Guinea. Micropaleontology, 27 (4), Haig, D.W Aptian-Abian foraminifers from Site 766, Cuvier Abyssal Plain, and comparison with coeval faunas from Australian region. hi: Gradstein, F., Ludden, J.N. et al. Proceedings of the Ocean Drilling Program, Scientific Results, 123: College Station, TX (Ocean Drilling Program), Haig, D.W. & Lynch, D.A A late early Albian marine transgressive pulse over northeastern Australia, precursor to epeiric basin anoxia: Foraminiferal evidence. Marine Micropaleol1tology, 22, Hanzlfkova,? Mikropaleontologicko-strtigraficke z,hodnocem vrtby H. Zukov NP-15. Sbornfk Ustredlliho Ustavu geologickfllo, 20, paleont., Hanzlfkova, E Foraminifers of the variegated Godula member in Moravia (Cenomanian-Turoman). Sbornik Geologickycll Ved, Paleontologie, 15, Hart, M.B., Bailey, H.W., Crittenden, S., Fletcher, B.N., Price, RJ. & Swiecicki, A Cretaceous. In: Jenkins, D.G. & Murral', J.W. (eds), Stratigraphical atlas offossil foramillifera, 2nd edition. Ellis Horwood Ltd., Chichester, Hecht, F.E Standard-Gliederung des Nordwestdeutschen Unterkreide nach Foraminiferen. Abhandlungen der Senckellbergischell Naturforschel1den Gesellschaft, 443, Hemleben, C & Traster, J Campanian-Maestrichtian deep-water foraminifers from Hole 543A, Deep Sea Drilling Project. In: Bij-Duval, B., Moore, J.C et al. Initial Reports of the Deep Sea Drilling Project, 78 A, Holbourn, AE.L. & Kaminski, M.A Lower Cretaceous deep-water benthic foraminifera of the Indian Ocean. Grzybowski Fundation Special Pubfication no. 4, Jones, T.R & Parker, W.K On the Rhizopodal fauna of the Mediterranean, compared with that of the Italian and some older Tertiary deposits. Quarterly Journal of the Geological Society of London, 16, Kaminski, M.A., Gradstein, F.M., Bersgren, W.A, Geroch, S. & Beckmann, J.P Agglutmated foraminiferal assemblages from Trinidad: Taxonomy, Stratigraphy and Paleobathymetry. In: F.M. Gradstein & F. Ragl (eds), Proceedings of Second International Workshop on Agglutinated Foraminifera, Vienna Abhandlungen der geologischel1 Blmdesanstait, 41, King, C, Bailey, H.W., Burton, CA. & King, AD Cretaceous of the North Sea. In: Jenkins, D.G. & Murray, J.W. (eds), Stratigraphical atlas of fossil foraminifera, 2nd edition. Ellis Horwood Ltd., London, Krasheninnikov, V.A. & Pflaumann, U Cretaceous agglutinated foraminifera of the Atlantic Ocean off west Africa (Leg 41, Deep Sea Drilling Project). In: Lancelot, Y., Seibold, E. et al. [nitial Reports of the Deep Sea Drilling ProJect, 41, Kuhnt, W Agglutinated foraminifera of western Meditrranean pper Cretaceou.s pelagic limestones (Umbnan Apennmes, Italy, and betic Cordillera, Southern Spain). Micropaleontology, 36, Kuhnt, W Deep-water agglutinated foraminifera from the Lower Cretaceous (Neocomian)'complex a Aptychus' Formation (Corridor de Boyar, Betic Cordillera, southern Spain). Journal of Micropalaeontology, 14 (1), Kuhnt, W. & Kaminski, M.A Paleoecology of Late Cretaceous to Paleocene deep-water aggrutinated foraminifera from the North Atlantic and western Tethys. In: Hemleben, C, Kaminski,M.A., Kuhnt, W. & Scott, D.B. (eds), Paleoecology, Biostratigraphy, Paleoceanography and Taxonomy of Agglutinated Foraminifera. NATO AS! Series C 327, (Kluwer Academic Publishers), Kuhnt, W., Geroch, S., Kaminski, M.A., Moullade, M. & Neagu, T Upper Cretaceous abyssal claystones in the North Atlantic and Western Tethys: current status of biostratigraphical correlation using agglutinated foraminifers and palaeoceanographic events. Cretaceous Research, 13, Lalicker, CG New Cretaceous Textulariidae. Contriblltions from tlze Cushman Foundatioll for Foraminiferal Research, 11 (1), Loeblich, A.R & Tappan, H Foraminifera from the Walnut Formation (Lower Cretaceous) of northern Texas and southern Oklahoma. Journal of PaleOlltology, 23 (3), Loeblich, AR & Tappan, H Foraminifera of the type Kiowa Shale Lower Cretaceous of Kansas. Ulliversity of Kansas Paleontological Contributions, Protozoa, 3, Loeblich, A.R & Tappan, H A revision of Mid Cretaceous Textulanan foraminifers from Texas. JOllrnal of Micropalaeontology, 1, Loeblich, A.R. & Tappan H Foraminiferal Genera and their Classification. van Nostrand Reinhold Co., New York, 1-970, pi Magniez-Jannin, F Les Foraminifires de l'albien de I'A II be: Paleontologie, Stratigraphie, Ecologie. Cahiers de Paleontologie. Editions du CBRS, Paris, Maync, W Lower Cretaceous foraminiferal fauna from Gorringe Bank, eastern North Atlantic. In: Hollister, CD., Ewing, J.I., et al. (eds), Illitial Reports of the Deep Sea Drilling Project, 13 (2), Meyn, H. & Vespermann, J Taxonomische Revision von Foraminiferen der Unterkreide SE-Niedersachsens nach Roemer (1839, 1841, 1842), Koch (1851) lind Reuss (1863). Senckel1bergiana letizaea, 74, Mjatliuk, E.Y Foraminifera of the Upper Jurassic and Lower Cretaceous of the region of the Middle Volga River and the Great Syrte. Trudy VNIGRI, A 120, Moullade, M., Kuhnt, W. & Thurow, J Agglutinated benthic foraminifers from Upper Cretaceous variegated clays of the North Atlantic Ocean (DSDP Leg 93 and ODP Leg 103). In: G. Boillot, E.L. Winterer, et al. (eds), Proceedings of the Ocean Drilling Program, Scientific Results, 103, Neagu, T Studiul foraminiferelor aglutinante din argilele Cretacic superioare de pe Valea Sadovei (Cimpulung-Moldovenesc) si bazinul superior al vaii Buzaului. Stud. Cerc. Geol. Geophys. Geogr., Ser. Geol., 7, Neagu, T Albian foraminifera of the Rumanian Plain. Mlcropaleol1tology, 11 (1), Neagu, T Micropaleontological and stratigraphical study of the Upper Cretaceous deposits between the upper valleys of the Buzau and Riul rivers (Eastern Carpathians). Memorii, Institutul Geologic, 12, d'orbigny, A Memoire sur les Foraminiferes de la Craie Blanche. Memoires de la Societe Geologique de France, 1, Parker, W.K. & Jones, T.R On the nomenclature of the Foraminifera. VIII, Textularia. Annals and Magazine of Natural History, 3 (11), Price, R.J The evolutionary interpretation of the Foraminiferida Arenobuli11lina, Gavelinella and Hedbergella in the Albian of North-West Europe. Palaeontology, 20 (3), Reicherter, K., Pietsch, T., Kuhnt, W., Manthey, J., Homeier, G., Wiedmann, J. & Thurow, J MId-Cretaceous paleogeography and paleoceanography of the Betic Seaway (Betic Cordil1era, Spain). Palaeogeograpizy, Palaeoclimatology, Palaeoecology, 107, Reuss, A.E Die Versteinenmgen der bohmischen Kreideformation. Erste Abteilung. Stuttgart: E. Schweizerbart'sche Verlagsbuchhandlung und Druckerei, 1-58.

13 Upper Albian agglutinated foraminifera, NE Germany 457 Reuss, A.E Die Foraminiferen der westphalischen Kreideformation. Sitzlmgberichte der Mathematisclle Natllrwissenschaftliche Klasse der Kayserliche Akademie der Wissensclzaftel1 in Wien, 40, Rellss, A.E Die Foraminiferen des norddeutschen Hils und Gault. SitZllngberichte der Mathematische Natllrwissellscllllftliche Klasse der Kayserliche Akademie der WissenschaftCll in Wien, 46, Risch, H Stratigraphie der huheren Unterkreide der bayerischen Kalkalpen mit Hilfe von Mikrofossilien. Palaeontographica, A 138 (1-4),1-80. Roemer, F.A Die Versteinenmgen des norddeutschen Kreidegebirges. Hannover, Ragl, F Late Cretaceolls to Pleistocene foraminifera from the southeast Pacific Basin, DSDP, Leg 35. In: Hollister, C. D., Craddock, c., et al. Initial Reports of the Deep Sen Drilling Project, 35, Ragl, F A Late Cretaceous Flysch-type Agglutinated Foraminiferal Fauna from the Troc1l1lmminoides proteus Type Locality (Wien-Hiltteldorf, Austria). In: M.A. Kaminski, S. Geroch, & M.A. Gasinski (eds), Proceedings of the FOllrth International Workshop on Aggilltinated Foraminifera. Grzybowski Foundation Special Publication 3, Sars, G.O Unders0gelser over Hardangerfjordens Fauna. Fordhandlllllger i Videnskabsselskabet I Kristiania, 1871, (Institute of Oceanographic Sciences Translation no. 179). Schulze, F.E Zoologische Ergebnisse der Nordseefahrt vom 21. Juli bis 9. September 1872, I, Rhizopoden, II. Jahresberichte Kommission zur Untersllchllng der Deutscher Meere in Kiel/iir die Jailr 1872, 1873, Sliter, W.V Upper Cretaceous foraminifera from southern California and northwestern Baja California, Mexico. Ulliversity of Kansas Paleontological Contri blltions, 49 (7), Tappan, H Foraminifera from the Grayson Formation of northern Texas. Journal of Paleontology, 14, Ten Dam, A Sur quelques especes nouvelles ou peu connues dans Ie Cretace inferieur (Albien) des Pays-Bas. Geologij ell Mijnbollw, 8, Ten Dam, A Les Foraminiferes de l'albien des Pays Bas. Memoires de la Societe Geologiqllc de France, Nouvelle Serie 63, 29, Trujillo, E.F Upper Cretaceous foraminifera from near Redding, Shasta County, California. JOllrnal of Paleontology, 34, Weidich, K Die kalkalpine Unterkreide und ihre Foraminiferenfauna. Zittefiana, Abhandlungen der Bayeriscilcll Staatssammillng fiir Paliiolltologie lind historische Geologie, 17, ,

14 '" S... k. KIOfOWJkl, Prokoph, Kuhnt" W.,nu

15 Upptr Albl.n.gglulln.IN foramlniftra, Nil Ctrmany PI J.l.II""""","8"" I,.If ""J/"M... (Bo_I), (I) \ mt (2) 1>12 2'1 (5U m). J. /to c, Ito ijftpttft<'''' ft.nd,i.. F1 (JOO rn).. Ron""""" w. M2 J.I (320 m). S. A...xr.;./".. Cuoh... n " AJe.. ndft. M2 (.j()6 mj, t. liop1""""'i"" "'" pwi...,"".. (NHZU), FJ (550 mj,,... liopiop1t"'8."'"'" atfiva1t (R;""... J. I7J 1>12 25 (4E101 m). (8) M2 25 {3iXI ml. Lill«llu"" /".i/onrojf (Brady). Fl (493 rn). 10. T_... p. 1'11'9\41 9 _rnl U U. IIrlmm'y' n.f. (Ten [).am). (U FI5 (75 mi. (IJ Ml 19 (:180 ""'I r",,,,,,,,,, rlwp""'"' ur"'i<et. (1 ) WFI6 IW rnj. (1 4) M2-I9 (473 ml. IS. ft.,...,,,,,,,,, """"... (C'1'XyI. WFlO (450/1 m). 16. r",.j.,.. "P" F5 {346 ml 1'.... JoNI'... w.. (433 rn). 1& 8W"""."".p.. FlO (480.om). u. V",... ii.".;j., _... (MjaWo k). 518 (4)1 m.

16

17 Upp<" Albian.sglutln.lOd ror.",i"lrr, NE Cornlony

18 Ammodiscus infimus Ammodist:us sp. CIQmospira spp. Pseudoslmoilina lmtiqun Reopho.x minutus HapJophram()ides spp. Recllrvoidt"$ spp. Ammobaeulites subcretacclis Haplophra,"illltt aequole Ht-teranlllx Crctosa Trocllammiua sp. Textularia spp. PselidoboJivina sp. S{tiropiedinato SJ)irop/ecfinata Spirop/ectinata E Daehre 4183 deplh(m) sample FI F2 F & futtis EgereJli"a nlarioe Rllflbdammina sp. Rlliznmm;na indivisa PsammosviJaera fusca Saccammina spp. Ammodiscus crttacc1is Ammodiscus iufimus Ammodiscus 51'. l'sf'udo$lrmoilina anliquo spp. spp. AmJdiles 5ubcretgcells H(lplopllraillnr aequale Hetmm!lIx erelosa Trocharnmina sp. Texlular;a spp. Pseudobolivina sp. SpirOf.,leclina/a Quneetens Sp;ropieclinatalJt'lIellslaedti SpiropJectinata complallat(j TrUaxia lricarinata FalsoRfwdrilinella ex gr. nlqfsialju Arenobulimina DOfOlhia E Daehre 4183 deplh (m) sample F26 F27 F , : F J F F F F F Gt Waren WFI WF2 WF F WF FlO Fll WFS WF Fl WF : F WF F FIS U Fl F F1S F F F F F F F2S WF9 WFlO WF11 WFl2 WF13 WF14 WFl5 WF16 WF17 WFl8 WF19 WF l l ? "ti IV Q.. >r i"" Q 6' : l a. III g' III [ :3 S' iil S' a [ Iii 8: ::i' a: e:.. '" 1,... ()I:j iil :3 '! V'l N III... rt) 1'" 1S o III Ii'\'.!» "1:1 ::r = ::r -:; R>

19 Upper Albian agglutinated foraminifera, NE Germany 463 Appendix 2. Comparison of the distribution and abundance of the most common taxa (percentage from total agglutinated assemblage). E Daehre 4/83 Ammodiscus Textularia Gt Waren V8t deplll Rhizammillil 1m) indivisa Ammodiscus cretaceus Glomospira charoides 1380':-::-::F=t-::-:CrF=t--FF:::::'=1=:=!- 1382t::=l==t--t:'::;';=-Pl=t=-=:t:= 1384.fh.. ; ; deplh Marssonella. Arenobulimillil Eggerellina Heterantyx 1m) orycona & turns sr. marille cretosa depth Spiroplectinata Falsogaudryine/Ia Tritaxia.: 1m) annectens S. complanata ex gr. moesiana triazrinata d 1378"--n'--.., Fl 1380Ito= : bivalve f6,- max. A -log : WF t...)....) c 1386 Ai. : : : WF 4..,.:g =p-!:'F=t-iF::-t=*:-:--F:F:=-t-:=t-j=i==, :...! WF S :;;: 1380 i.!! : <> 1 1! g<] 1072 '... L...! : y...,.. "..., 1: ;:i bw_ i.l... l: "',... { F15 i m,n. 0 WF ; FIG ;i: i WF 11 F17 f :WF 12 -ci398---tt:s.g,--l----,.::t::t-t:±t:--j::j::j-ic..:.. i'i:.....!! F23 : : :WF18 h-.. :.... : : :: J * -h-4,==-d : i o",i, k tk!> k k <f. :.. ti.:.::1::jel., < P.$ k k

20 <>,---