Cloud modeling and Lotka-Voltera Spence Lunderman December 9, 2017 University of Arizona Department of Mathematics

Similar documents
8 Ecosystem stability

Dynamics on a General Stage Structured n Parallel Food Chains

Final Project Descriptions Introduction to Mathematical Biology Professor: Paul J. Atzberger. Project I: Predator-Prey Equations

Observational constraints on mixed-phase clouds imply higher climate sensitivity

Introduction to Biomathematics. Problem sheet

3.5 Competition Models: Principle of Competitive Exclusion

2D-Volterra-Lotka Modeling For 2 Species

Gerardo Zavala. Math 388. Predator-Prey Models

2 The model with Kaldor-Pasinetti saving

NUMERICAL SIMULATION DYNAMICAL MODEL OF THREE-SPECIES FOOD CHAIN WITH LOTKA-VOLTERRA LINEAR FUNCTIONAL RESPONSE

Stability Analysis And Maximum Profit Of Logistic Population Model With Time Delay And Constant Effort Of Harvesting

Stability and Hopf bifurcation analysis of the Mackey-Glass and Lasota equations

Theory of Ordinary Differential Equations. Stability and Bifurcation I. John A. Burns

Lab 5: Nonlinear Systems

Computing 3D Bifurcation Diagrams

M469, Fall 2010, Practice Problems for the Final

The Dynamic Behaviour of the Competing Species with Linear and Holling Type II Functional Responses by the Second Competitor

Dynamical Systems in Neuroscience: Elementary Bifurcations

A Producer-Consumer Model With Stoichiometry

Models Involving Interactions between Predator and Prey Populations

ROLE OF TIME-DELAY IN AN ECOTOXICOLOGICAL PROBLEM

DETC DELAY DIFFERENTIAL EQUATIONS IN THE DYNAMICS OF GENE COPYING

Delayed Dynamics in Heterogeneous Competition with Product Differentiation

Dynamical Systems and Chaos Part II: Biology Applications. Lecture 6: Population dynamics. Ilya Potapov Mathematics Department, TUT Room TD325

Continuous Threshold Policy Harvesting in Predator-Prey Models

Delayed Dynamics in Heterogeneous Competition with Product Differentiation

Math53: Ordinary Differential Equations Autumn 2004

Analysis of One Model of the Immune System

Math 215/255 Final Exam (Dec 2005)

Physics: spring-mass system, planet motion, pendulum. Biology: ecology problem, neural conduction, epidemics

Linearization of Differential Equation Models

A Discrete Model of Three Species Prey- Predator System

Stability and bifurcation of a simple neural network with multiple time delays.

Ordinary Differential Equations

The diffusive Lotka-Volterra predator-prey system with delay

SPIRAL WAVE GENERATION IN A DIFFUSIVE PREDATOR-PREY MODEL WITH TWO TIME DELAYS

STABILITY ANALYSIS OF DELAY DIFFERENTIAL EQUATIONS WITH TWO DISCRETE DELAYS

THETA-LOGISTIC PREDATOR PREY

Stability and bifurcation in a two species predator-prey model with quintic interactions

Copyright (c) 2006 Warren Weckesser

4.9 Free Mechanical Vibrations

Local Stability Analysis of a Mathematical Model of the Interaction of Two Populations of Differential Equations (Host-Parasitoid)

Predator-Prey Model with Ratio-dependent Food

Vectors, matrices, eigenvalues and eigenvectors

Introduction. Effect of aerosols on precipitation: - challenging problem - no agreement between the results (quantitative and qualitative)

ESAIM: M2AN Modélisation Mathématique et Analyse Numérique M2AN, Vol. 37, N o 2, 2003, pp DOI: /m2an:

Mathematical Modelling in Biology Lecture Notes

CHALMERS, GÖTEBORGS UNIVERSITET. EXAM for COMPUTATIONAL BIOLOGY A. COURSE CODES: FFR 110, FIM740GU, PhD

arxiv: v1 [nlin.ao] 19 May 2017

Mathematical models in life sciences. Lecture notes of a course held by Marco Di Francesco based on the book by J. D. Murray

Introduction to Dynamical Systems Basic Concepts of Dynamics

AN AK TIME TO BUILD GROWTH MODEL

5 Alfred Lotka, Vito Volterra, and Population Cycles

MA 266 Review Topics - Exam # 2 (updated)

Chapter 7. Nonlinear Systems. 7.1 Introduction

Lotka Volterra Predator-Prey Model with a Predating Scavenger

Oscillatory Turing Patterns in a Simple Reaction-Diffusion System

2 One-dimensional models in discrete time

Math 312 Lecture Notes Linear Two-dimensional Systems of Differential Equations

Key words and phrases. Bifurcation, Difference Equations, Fixed Points, Predator - Prey System, Stability.

Weak Allee effect in a predator-prey system involving distributed delays

IN real world, the study of population dynamics including

1 Simple one-dimensional dynamical systems birth/death and migration processes, logistic

(8.51) ẋ = A(λ)x + F(x, λ), where λ lr, the matrix A(λ) and function F(x, λ) are C k -functions with k 1,

PTYS 214 Spring Announcements. Midterm 3 next Thursday!

Research Article Stability Switches and Hopf Bifurcations in a Second-Order Complex Delay Equation

Predator-swarm interactions

as Hopf Bifurcations in Time-Delay Systems with Band-limited Feedback

Chapter 14 Three Ways of Treating a Linear Delay Differential Equation

Math Ordinary Differential Equations

GATE EE Topic wise Questions SIGNALS & SYSTEMS

Andronov Hopf and Bautin bifurcation in a tritrophic food chain model with Holling functional response types IV and II

Lesson 9: Predator-Prey and ode45

dt = A x, x(0) = C. (1) where A is an n n matrix of constants, with n distinct eigenvalues. The solution formula is

1.Introduction: 2. The Model. Key words: Prey, Predator, Seasonality, Stability, Bifurcations, Chaos.

MA 138: Calculus II for the Life Sciences

Time Response Analysis (Part II)

DYNAMICS IN 3-SPECIES PREDATOR-PREY MODELS WITH TIME DELAYS. Wei Feng

Satellite-based estimate of global aerosol-cloud radiative forcing by marine warm clouds

Hopf Bifurcation Analysis of a Dynamical Heart Model with Time Delay

Is chaos possible in 1d? - yes - no - I don t know. What is the long term behavior for the following system if x(0) = π/2?

Bifurcation and Stability Analysis of a Prey-predator System with a Reserved Area

Queues with Choice via Delay Differential Equations

A Predator-Prey Model with a Time Lag in the Migration

Three ways of treating a linear delay differential equation

Observed Southern Ocean Cloud Properties and Shortwave Reflection

Stability and Hopf bifurcation of a ratio-dependent predator prey model with time delay and stage structure

Figure 5: Bifurcation diagram for equation 4 as a function of K. n(t) << 1 then substituting into f(n) we get (using Taylor s theorem)

Non-Linear Models. Non-Linear Models Cont d

MATH 1700 FINAL SPRING MOON

Complex Behavior in Coupled Nonlinear Waveguides. Roy Goodman, New Jersey Institute of Technology

Analysis of the Takens-Bogdanov bifurcation on m parameterized vector fields

Modelling biological oscillations

Nonlinear Autonomous Dynamical systems of two dimensions. Part A

second order Runge-Kutta time scheme is a good compromise for solving ODEs unstable for oscillators

Report submitted to Prof. P. Shipman for Math 540, Fall 2009

BIFURCATION PHENOMENA Lecture 1: Qualitative theory of planar ODEs

MATH 251 Final Examination December 16, 2015 FORM A. Name: Student Number: Section:

HOMEWORK 3: Phase portraits for Mathmatical Models, Analysis and Simulation, Fall 2010 Report due Mon Oct 4, Maximum score 7.0 pts.

4 Second-Order Systems

Transcription:

1 Low dimensional cloud modeling Cloud modeling and Lotka-Voltera Spence Lunderman December 9, 17 University of Arizona Department of Mathematics Cloud modeling is a very difficult task: one must understand the cloud s microphysics, albedo or ability to reflect sunlight, mixture of liquid and ice, etc. These difficulties result in wide variations of how clouds are represented in global climate models []. For example, certain cloud models that covert water to ice at relatively low temperatures lead to a global climate model with a higher mean-state cloud fraction [3]. In an attempt to tackle some of these known difficulties, researchers are looking to qualitatively describe clouds with low dimensional models. An example of this comes from Koren and Fiengold (11): they use the Lotka-Volterra predator prey equations to model the interaction between the rain rate of a cloud and the cloud s liquid water path (column integrated liquid water content). It would seem that these two observables have some predator prey interactions. As water droplets form, they consume the water vapor within the cloud, potentially leading to the destruction of the cloud. In the absence of rain, the amount of water vapor can grow until reaching a critical point. Figure 1 shows a cloud system over the Atlantic ocean. In the figure, there are areas of dense clouds structures and areas of open cells which show the dark ocean below. A goal of this work is to capture qualitative details of these cloud types using these predator prey insights. As a cloud presents rain, the liquid water path (LWP) and height of the cloud (H) decrease. However, instead of setting up the predator-prey model for the interactions between LWP and R, the authors opt to design the system around H and the cloud drop concentration N. H relates to LWP by LW P c 1 H (1) where c 1 is a function of cloud based temperature and pressure and to first order the rain rate R can be expressed as R α H3 N for appropriate α. The balance equations for cloud depth H and cloud drop concentration N are () H o H(t) Ḣ(t T ) and dn τ1 N o N(t) τ Ṅ(t T ). (3) In the above equations H o and N o are the full environmental potential for cloud development; this means the first term in both equations represent the system s approach to their full potential with characteristic time constants τ 1 and τ. Ḣ and Ṅ are the loss of H and N after some delayed time T. Based on theoretical and empirical results, the authors derive Ḣ and Ṅ to yield the final set of equations H o H(t) αh (t T ) τ1 c 1 N(t T ) and dn N o N(t) τ αc H 3 (t T ). (4) For the specified parameters, figure 1 illustrates the interactions between the cloud depth and drop concentration as well as the derived interactions between LWP and rainrate. Our goal now is to find the system s steady states and understand their stabilities. To make life simpler, we perform the following transformations to yield dimensionless solutions which we can later transform back: Let t t τ 1, H.1 c H, and N c 1E11ατ 1 N (5) and define c c 1E9c 1, τ τ 1 τ. This leaves us with the following delay differential equations H o H(t) ch (t T ) N(t T ) and dn τ(n o N(t)) H 3 (t T ). (6)

Figure 1: Left: Cloud depth H (blue) and cloud drop concentration N (brown). Middle: Cloud depth plotted against cloud drop concentration. Right: Rainrate (blue) and liquid water path (brown); these values are calculated by () and (1) respectively. For all three figures, the parameters are H o 65m, N o 469E6 m 3, α E 3 m 5 /d, c 1 E 9 m 1, c.3 m 1, τ 1 8 min, and τ 84 min, T min. A steady state solution to the above delay differential equation is independent of the delay T. We solve for the steady state solutions in terms of H o and N o keeping in mind our larger parameter estimation goals. Solving the equations and dn and only keeping positive solutions we get H N + N + 4H o N and N N o H3 τ. (7) Given that we are treating H o and N o as free parameters, the contour plots for the steady state solutions are a surface in the space (H o ; N) for the cloud depth H and (N o ; H) for the drop concentration N. The contour plots for the steady state solutions can be seen in figure. Figure : Steady state contour plots for: cloud depth, m, (left) and drop concentration, m 3, N (right). The parameters and initial conditions are the same as figure 1. For the cloud depth contour plot in figure, there appear to be two regimes: for higher drop concentration the steady states are mainly dictated by H o and lower drop concentration seems to control the steady solution with little influence from H o. As for the drop concentration contour plot, the steady state solutions are seemly dictated by the cloud depth, influenced little by N o. Studying stability of steady states and doing bifurcation analysis for delay differential equations is a difficult problem. For example, determining linear stability for a differential equations requires finding the eigenvalues of the linearized system. For delay differential equations, the characteristic equation for the linearized system can have infinitely many roots, often making it a difficult task to determine stability. For these reasons, we step away from the derived cloud models and focus our attention towards a slightly easier problem. We will now study the steady states and their stability for the delayed Lotka-Voltera equations. It is our hopes that this work can direct future work with these cloud models. Predator Prey The Lotka-Voltera equations are a standard mathematical model to illustrate the interactions between a predator and its prey. Independently derived by Alfred J. Lotka and Vito Volterra over a century ago, these equations and many variations have been studied extensively in biological and ecological sciences. The equations are given by dc C(α βr) and dr R(γ δc)

where C and R represent the predator and prey population respectively and α, β, γ, and δ are positive model parameters. Complexity can be adding to these models by taking into account the amount of time it between birth and maturity of the predator or prey. Adding this into the model yields the equations dc dr C[α βr(t T )] and R[γ δc(t T )] (8) where T represent some time delay for the predator and prey. These are the equations that we will work with to determine the steady states and their linear stability, in an effort to guide future work with the cloud models. This system has two biologically relevant steady states: one at (C, R) (, ) and the nontrivial one at (C, R) (γ/δ, α/β) which can easily derived. For the trivial steady state, we can easily determine the linear stability. C(t T ) C C α (9) + R(t T ) R γ R The characteristic matrix for this linearizion about (, ) is λ α T λ e (λ) λ γ λα. λ+γ Then the characteristic equation is given by P (λ) det (λ) (λ α)(λ + γ) (1) and the roots to (11) are the desired eigenvalues: α and γ. Given that all model parameters are positive, we know that we have a saddle point at the fixed point (, ). For the second relevant steady state (γ/δ, α/β), the linearized of (8) about the steady state is! βγ C C C(t T ) δ + δα (11) R R(t T ) R β yielding the characteristic matirx λ (λ) δα λ β! βγ T λ λ δ e. δα T λ βe λ βγ δ! et λ (1) (13)

and characteristic equation P (λ) λ + αγe T λ. (14) There are infinitely many roots to (14). To begin understand the stability of the steady state, we first consider two special cases. The simplest case to consider is if λ. In this case P () αγ. Given that α and γ are positive, P () does not have any roots and we no longer need to consider this case. A more interesting case is if the eigenvalue is purely imaginary, i.e. λ iω for ω >. Then (14) becomes trying to solve Expanding (15) and splitting into real and imaginary parts, we get (iω) + αγe T ωi. (15) αγ cos ( ωt ) ω (16) and αγ sin ( ωt ). (17) Given that α and γ are positive, (17) gives that ω π k for k Z. Substituting ω π/t into (16), T αγ cos ( π ) T π αγ ( π ) (18) T (19) w αγ () Thus, λ ( αγ k)i for k Z\{} is a root of (14) provided the delay T π αγ Let us define a function λ(t ) : µ(t ) + iω(t ) around T π αγ such that µ(t ) and ω(t ) αγ. With hopes to satisfy the Hopf bifurcation theorem conditions [4] [5], we want to show that dre(λ) >. (1) λλ(t ) If we can show (1), it would be one step towards showing that (8) has a Hopf bifurcation at (C, R) (γ/δ, α/β). We implicitly differentiate (1) equated to zero Evaluating (3) at λ λ(t ) and T T yields λ dλ αγt et λ dλ () dλ (λ αγt et λ ) (3) The right factor is nonzero implying that dλ dλ (4 αγi π αγ). (4) resulting in dre(λ). λλ(t ) Thus, we can not use the Hopf bifurcation theorem to gain understanding of our dynamical system. We focus our attention towards understanding, in the most general case, the linear stability of the steady state solution (C, R) (γ/δ, α/β). We need to understand the roots of (14). Taking small bites, we assume that αγ 1 and T 1/ leaving us to find the solutions to λ + e λ (5)

With an eye towards determining stability, we focus on the sign of the real parts of solutions to (5) which are of the form Re(λ) Re W k, i )] (6) Where W ( k, i ) is the k th solution of the Lambert W function which is defined as the inverse function to f(z) ze z. A few evaluations gives us all the information we need: Re W, i )].166 and Re W 1, i )] 1.867 which shows that this steady state is linearly unstable. 3 Conclusions To gain insight into cloud modeling, a notoriously difficult task, we considered a simplified model that captures qualitative cloud behavior. The Lotka-Volterra predator prey models provide a well studied system to derive more complex dynamical systems that present oscillatory behavior. Although this work is not complete, studying the stability of steady states in the delayed Lotka-Volterra models provide useful insight. This instability of the fixed point is not physically realistic in that a predator prey system cannot have unbounded oscillations. Hopefully future work will find stable steady states in the Koren-Fiengold cloud models that match observable cloud patterns. References [1] Koren I., Feingold G., (11) Aerosol cloud precipitation system as a predator-prey problem. Proceedings of the National Academy of Sciences. [] Yan, W. (16), Modeling the effects of clouds on climate, Eos, 97, doi:1.19/16eo49999 [3] McCoy, Daniel T. and Tan, Ivy and Hartmann, Dennis L. and Zelinka, Mark D. and Storelvmo, Trude (16) On the relationships among cloud cover, mixed-phase partitioning, and planetary albedo in GCMs. Journal of Advances in Modeling Earth Systems [4] Hale, Jack K., and Huseyin Kocak. Dynamics and Bifurcations. New York, NY: Springer New York, 1991. Print. [5] Guckenheimer, J. and Holmes, P. (). Nonlinear oscillations, dynamical systems, and bifurcations of vector fields. New York: Springer. [6] Koen E., Tatyana L., Dirk R. () Numerical bifurcation analysis of delay differential equations. Journal of Computational and Applied Mathematics. [7] Forde J. (5) Delay Differential Equation Models in Mathematical Biology. [] Kalmar-Nagy T. (9) Stability analysis of delay-differential equations by the method of steps and inverse Laplace transform. Differential Equations and Dynamical Systems