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How Organisms Obtain Energy All living organisms use energy to carry out all biological processes. Real-World Reading Link New York City is sometimes called the city that never sleeps. Much like the nonstop movement of a big city, living cells are sites of constant activity. Transformation of Energy Many chemical reactions and processes in your cells are ongoing, even when you might not think that you are using any energy. Macromolecules are assembled and broken down, substances are transported across cell membranes, and genetic instructions are transmitted. All of these cellular activities require energy the ability to do work. Figure 1 shows some of the major advancements in the study of cellular energy. Thermodynamics is the study of the flow and transformation of energy in the universe. Reading Preview Essential Questions What are the two laws of thermodynamics? What is the difference between an anabolic pathway and a catabolic pathway? How does ATP work in a cell? Review Vocabulary trophic level: each step in a food chain or a food web New Vocabulary energy thermodynamics metabolism photosynthesis cellular respiration adenosine triphosphate (ATP) (t)/ism/phototake; (r)/ism/phototake Figure 1 Understand Cellular Energy Scientific discoveries have led to a greater understanding of photosynthesis and cellular respiration. Laws of thermodynamics The first law of thermodynamics is the law of conservation of energy, which states that energy can be converted from one form to another, but it cannot be created nor destroyed. For example, the stored energy in food is converted to chemical energy when you eat and to mechanical energy when you run or kick a ball. The second law of thermodynamics states that energy cannot be converted without the loss of usable energy. The energy that is lost is generally converted to thermal energy. Entropy (EN truh pee) is the measure of disorder, or unusable energy, in a system. Therefore, the second law of thermodynamics can also be stated as entropy increases. One example of the second law of thermodynamics is evident in food chains. Recall that in a food chain, the amount of usable energy that is available to the next trophic level decreases. Autotrophs and heterotrophs All organisms need energy to live. Directly or indirectly, nearly all the energy for life comes from the Sun. Some organisms make their own food, while others must obtain it from other organisms. Autotrophs are organisms that make their own food. Some autotrophs, called chemoautotrophs, use inorganic substances such as hydrogen sulfide as a source of energy. Other autotrophs, such as the plant in Figure 2, convert light energy from the Sun into chemical energy. Autotrophs that convert energy from the Sun are called photo-autotrophs. Heterotrophs, such as the aphid and the ladybug in Figure 2, are organisms that need to ingest food to obtain energy.

Figure 2 Almost all the energy in living organisms originates from the Sun, and energy flows from autotrophs to heterotrophs. Relate the laws of thermodynamics to the organisms in the figure. VOCABULARY WORD ORIGIN Autotroph comes from the Greek word autotrophos, meaning supplying one s own food Metabolism All of the chemical reactions in a cell are referred to as the cell s metabolism. A series of chemical reactions in which the product of one reaction is the substrate for the next reaction is called a metabolic pathway. Metabolic pathways include two broad types: catabolic (ka tuh BAH lik) pathways and anabolic (a nuh BAH lik) pathways. Catabolic pathways release energy by breaking down larger molecules into smaller molecules. Anabolic pathways use the energy released by catabolic pathways to build larger molecules from smaller molecules. The relationship of anabolic and catabolic pathways results in the continual flow of energy within an organism. Energy continually flows between the metabolic reactions of organisms in an ecosystem. Photosynthesis is the anabolic pathway in which light energy from the Sun is converted to chemical energy for use by the cell. In this reaction, autotrophs use light energy, carbon dioxide, and water to form glucose and oxygen. The energy stored in the glucose produced by photosynthesis can be transferred to other organisms when the molecules are consumed as food. Cellular respiration is the catabolic pathway in which organic molecules are broken down to release energy for use by a cell. In cellular respiration, oxygen is used to break down organic molecules, resulting in the production of carbon dioxide and water. Notice the cyclical nature of these processes in Figure 3, where the products of one reaction are the reactants for the other reaction.

Figure 3 In an ecosystem, photosynthesis and cellular respiration form a cycle. Identify the anabolic and catabolic pathways in the figure. MiniLab: Relate Photosynthesis to Cellular Respiration How do photosynthesis and cellular respiration work together in an ecosystem? Use a chemical indicator to examine how carbon dioxide is transferred in photosynthesis and cellular respiration. Procedure 1. Read and complete the lab safety form. 2. Prepare a data table to record the contents, treatment, initial color, and final color for two experimental test tubes. 3. Pour 100 ml of bromothymol blue (BTB) solution into a beaker. Using a straw, exhale gently into the solution until it just turns yellow. WARNING: Do not exhale so much that the solution bubbles over or that you get a headache. Do not suck on the straw. 4. Fill two large test tubes three-quarters full with the yellow BTB solution. 5. Cover one test tube entirely with aluminum foil. Place a 6-cm sprig of an aquatic plant into both of the tubes, tightly insert the stoppers into the tubes, and place them in a rack in bright light overnight. 6. Record your observations in your data table. Analysis 1. Describe the purpose of the tube covered with aluminum foil. 2. Explain how your results demonstrate that photosynthesis and cellular respiration depend on one another. ATP: The Unit of Cellular Energy Energy exists in many forms, including light energy, mechanical energy, thermal energy, and chemical energy. In living organisms, chemical energy is stored in biological molecules and can be converted to other forms of energy when needed. For example, the chemical energy in biological molecules is converted to mechanical energy when muscles contract. Adenosine triphosphate (uh DEN uh seen tri FAHS fayt) ATP is the most important biological molecule that provides chemical energy. ATP structure

ATP is a multipurpose storehouse of chemical energy that can be used by cells in a variety of reactions. Although other carrier molecules transport energy within cells, ATP is the most abundant energy-carrier molecule in cells and is found in all types of organisms. As shown in Figure 4, ATP is a nucleotide made of an adenine base, a ribose sugar, and three phosphate groups. Figure 4 The breakdown of ATP releases energy for powering cellular activities in organisms. ATP function ATP releases energy when the bond between the second and third phosphate groups is broken, forming a molecule called adenosine diphosphate (ADP) and a free phosphate group, as shown in Figure 4. Energy is stored in the phosphate bond formed when ADP receives a phosphate group and becomes ATP. As shown in Figure 4, ATP and ADP can be interchanged by the addition or removal of a phosphate group. Sometimes ADP becomes adenosine monophosphate (AMP) by losing an additional phosphate group. There is less energy released in this reaction, so most of the energy reactions in the cell involve ATP and ADP. Review Lesson Summary The laws of thermodynamics control the flow and transformation of energy in organisms. Some organisms produce their own food, whereas others obtain energy from the food they ingest. Cells store and release energy through coupled anabolic and catabolic reactions. The energy released from the breakdown of ATP drives cellular activities. Vocabulary Review Each of the following sentences is false. Make each sentence true by replacing the italicized word with a vocabulary term. 1. Metabolism is the energy currency of the cell. 2. The study of the flow and transformation of energy is called energy. 3. Bioenergetics can exist in many forms. 4. Chemical reactions that convert energy within a cell are refered to as autotrophs. 5. Light energy is converted into chemical energy during the process of sunlight.

Understand Main Ideas 6. Identify the major source of energy for living organisms. 7. Describe an example of the first law of thermodynamics. 8. Compare and contrast anabolic and catabolic pathways. 9. Explain how ATP stores and releases energy. 10. Which is not a characteristic of energy? A. cannot be created nor destroyed B. is the capacity to do work C. exists in forms such as chemical, light, and mechanical D. changes spontaneously from disorder to order 11. Which organism depends on an external source of organic compounds? A. autotroph B. heterotroph C. chemoautotroph D. photoautotroph Use the figure below to answer question 12. 12. Which part of this food chain provides energy to just one other part? A. the chemoautotroph B. the heterotroph C. the Sun D. the photoautotroph 13. What do cells store and release as the main source of chemical energy? A. ATP B. ADP C. NADP + D. NADPH Constructed Response 14. How do autotrophs and heterotrophs differ in the way in which they obtain energy? 15. Open Ended Use an analogy to describe the role of ATP in living organisms. Think Critically 16. Describe how energy is released from ATP. 17. Relate anabolic and catabolic reactions. Create an analogy for the relationship between photosynthesis and cellular respiration. Biology 18. Write an essay describing the laws of thermodynamics. Use examples related to biology to support your ideas. 19. Create an analogy to describe the relationship between photosynthesis and cellular

respiration.