Crossword puzzles! Activity: stratification. zonation. climax community. succession. Match the following words to their definition:

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Activity: Match the following words to their definition: stratification zonation climax community succession changing community structure across a landscape changing community composition over time changes in community structure with height a mature community that is self maintaining Jun 27 10:12 a.m. Crossword puzzles! Feb 23 12:18 p.m. 1

Quiz! Mar 8 11:49 a.m. Epiphytes and Lianes Perching plants, or epiphytes, cling to the trunks of the canopy trees or grow in the leaf litter that accumulates between the branching limbs of large trees. Epiphytic species include many ferns and orchids; about half of the world s estimated 30 000 orchid species are epiphytic. Lianes are rooted in the ground, but clamber into the canopy where higher light levels enable them to develop extensive foliage. Staghorn fern Fern Orchid 2

Without looking back in your book make a sketch and use the labels to complete the diagram Ground Layer Tree Fern Epiphyte Shrub Emergent Subcanopy Canopy Ecological Succession Ecological succession is the process by which communities in a particular area change over time. Succession takes place as a result of complex interactions of biotic and abiotic factors. Past community Community composition changes Present community Future community Some species in the past community were out competed or did not tolerate altered abiotic conditions. The present community modifies such abiotic factors as: Light intensity and quality Wind speed and direction Air temperature and humidity Soil composition and water content Changing conditions in the present community will allow new species to become established. These will make up the future community. 3

Early Successional Communities A succession proceeds in several stages, until the formation of a climax community, which is stable until further disturbance. Early successional (or pioneer) communities are characterized by: Simple structure, with a small number of species interactions. Broad niches. Low species diversity. Primary Succession Primary succession refers to colonization of a region where there is no pre existing community. Examples include: newly emerged coral atolls, volcanic islands newly formed glacial moraines islands where the previous community has been extinguished by a volcanic eruption A classical sequence of colonization begins with lichens, mosses, and liverworts, progresses to ferns, grasses, shrubs, and culminates in a climax community of mature forest. In reality, this scenario is rare. Hawaii: Local plants are able to rapidly recolonize 4

Mount St Helens Primary succession more typically follows a sequence similar to the revegetation of Mt St Helens, USA, following its eruption on May 18, 1980. The vegetation in some of the blast areas began recovering quickly, with fireweed growing through the ash within weeks of the eruption. Animals such as pocket gophers, mice, frogs, and insects were hibernating below ground and survived the blast. Their activities played an important role in spreading seed and mixing soil and ash. Secondary Succession Secondary succession occurs where an existing community has been cleared by a disturbance that does not involve complete soil loss. Such disturbance events include cyclone damage, forest fires and hillside slips. Because there is still soil present, the ecosystem recovery tends to be more rapid than primary succession, although the time scale depends on the species involved and on climatic and edaphic (soil) factors. 5

Deflected Successions Humans may deflect the natural course of succession, e.g. through controlled burning, mowing, or grazing livestock. The resulting climax community will differ from the natural (pre existing) community. A relatively stable plant community arising from a deflected (or arrested) succession is called a plagioclimax. Grassland and healthland in lowland Britain are plagioclimaxes. Gap Regeneration The reduced sunlight beneath large canopy trees impedes the growth of the saplings below. When a large tree falls, a crucial hole opens in the canopy, allowing sunlight to reach the saplings below. The forest regeneration following the loss of a predominant canopy tree is called gap regeneration. Gap regeneration is an example of secondary succession. 6

Gap Regeneration Cycle Gap regeneration is an important process in established forests in temperate and tropical regions. Gaps are the sites of greatest understorey regeneration and species recruitment. The creation of a gap allows more light to penetrate the canopy and alters other factors that affect regeneration, exposing mineral soils and altering nutrient and moisture regimes. Activity: Video on Niches key notes please! Gause's Competitive Exclusion Principle states that organisms occupying exactly the same niche cannot coexist because they will compete for the same resources. How do organisms with a similar habitat and feeding requirement can minimise niche overlap, How do they differentiate their niches to avoid competition? Jun 26 9:25 p.m. 7

Activity: Read page 10 in Pathfinder 12. Explain the difference between habitat, species and ecological niche Tolerance and limiting factors key points Think about: How do organisms with a similar habitat and feeding requirement can minimise niche overlap, How do they differentiate their niches to avoid competition? Jun 27 10:35 a.m. Climax Communities In contrast to early successional communities, climax communities typically show: Complex structure, with a large number of species interactions. Narrow niches. High species diversity. large number of species interaction 8