Online Dictionary of Invertebrate Zoology: M

Similar documents
Introduction to Animals

Arthropoda ARTHRO JOINTED PODA FEET

Animals contain specialized cells

INVERTEBRATE DIVERSITY

Kingdom Animalia. Zoology the study of animals

2. Examine the external anatomy of the squid and identify the following structures: tentacles, arms, fins, siphon, mantle, eyes and collar.

Lab 2 Phylum Porifera and phylum Cnidaria. Grantia. Phylum Porifera. Kingdom :- Animalia. Phylum:- Porifera. Class:- Calcarea. Order:- Leucosolenida

Intro to Animals. Chapter 32

Subphylum Myriapoda and Insect External Morphology and Sensory Structures D. L. A. Underwood Biology General Entomology

09/12/2012. Classification. Characteristics. Learning Outcome G2. Student Achievement Indicators. Phylum Porifera The Sponges

Chapter 7. Marine Animals Without a Backbone

Life Science 7 th NOTES: Ch Animals Invertebrates

Name Class Date. Matching On the lines provided, write the letter of the description that best matches each term on the left. 1.

Resources. Visual Concepts. Chapter Presentation. Copyright by Holt, Rinehart and Winston. All rights reserved.

UNIT 8A MARINE SCIENCE: Lower Invertebrates

Chps : Animals. Characteristics of kingdom Animalia: Multicellular Heterotrophic Most are motile Possess sense organs

Characteristics of Echinoderms

BIOLOGY. An Introduction to Invertebrates CAMPBELL. Reece Urry Cain Wasserman Minorsky Jackson

3. Choanoflagellates resemble what? What is the significance of this resemblance?

World of Insects. Characteristics, Orders, and Collecting

BIOLOGY - CLUTCH CH.32 - OVERVIEW OF ANIMALS.

Name Class Date. After you read this section, you should be able to answer these questions:

Invertebrate Diversity

Superphylum Deuterostomia

Chapter 32. Objectives. Table of Contents. Characteristics. Characteristics, continued. Section 1 The Nature of Animals

28 3 Insects Slide 1 of 44

Chapter 24 Introduction to Animals

What is a Cnidarian?

Primitively there is a pair of ganglia per body segment but there has been progressive fusion of ganglia both within and between segments.

Unit 12 ~ Learning Guide

Chapter 8. Sponges Phylum Porifera Basic characteristics: simple asymmetric sessile

Arthropods. Ch. 13, pg

Introduction to Animal Diversity. Chapter 23.1, 23.2 and additional

KINGDOM ANIMALIA CHARACTERISTICS

Chapter 33: Invertebrates

What Is an Animal? Animals come in many shapes, forms, and sizes. About 98 percent of all animals are invertebrates. The Kingdom Animalia

Biology 224 Human Anatomy and Physiology - II Week 1; Lecture 1; Monday Dr. Stuart S. Sumida. Review of Early Development of Humans.

Introduction to Animal Kingdom. Invertebrates and Vertebrates

*Add to Science Notebook Name 1

Ph. Porifera and Ph. Cnidaria

By Sudha Vashisht P.G.G.C.G. -11 CHD

The Animal Kingdom: The Protostomes. Protostomes 4/16/2012. Chapter 30

Invertebrate Survey Lab

Notes - Porifera and Cnideria

Eukaryote Phylogeny. Glycogen. Kingdom Animalia. Amoebozoa Animalia. Plantae. Chromalveolata Rhizaria. Fungi. Excavata

LEARN 10 Insect Orders of the Wenatchee Watershed

Characteristics of Animals

Nonvascular Plants mosses, liverworts and hornworts are nonvascular plants. These lack vascular tissue which is a system of tubes that transport

Chapter 8-9 Intro to Animals. Image from:

23.1 Animal Characteristics EQ Although diverse, what common characteristics do all animal share?

EXTERNAL ANATOMY OF INSECTS

Developmental Zoology. Ectodermal derivatives (ZOO ) Developmental Stages. Developmental Stages

Sponges and Cnidarians

Today: Animal Body Plans. Animal Body Plans: The Gut. The Animal Kingdom- General Characteristics: Animal Body Plans: Symmetry

A. Incorrect! Sponges are mostly marine animals. This is a feature of sponges.

Chapter 32 Intro to Animals. Image from:

Classification. The three-domains. The six-kingdom system. The traditional five-kingdom system. Bacteria Archaea Eukarya

Animals. What are they? Where did they come from? What are their evolutionary novelties? What characterizes their diversification?

The cuticular structure

INVERTEBRATES. The Earth and Living Things. Carme Font Casanovas 1

Features of the Animal

Sponge and Cnidarian Review

BIOS1101 Lab Notes. Contents ANIMALS. Lab 1: Animal Diversity invertebrates. Lab 2: Animal Diversity 2 vertebrates

Outline. v Definition and major characteristics of animals v Dividing animals into groups based on: v Animal Phylogeny

Module 4: Marine Invertebrates I. Kingdom Animalia

Axis Specification in Drosophila

Biosc 41 9/10 Announcements

Marine Invertebrates

v Scientists have identified 1.3 million living species of animals v The definition of an animal

CHAPTER 14 Flatworms Phylum Platyhelminthes, Mesozoa, Nemertea

Paramecium. Sub-Order Peniculina. Genus Paramecium

Chapter 8. Sponges, Cnidarians, Comb Jellies, and Marine Worms

Revision Based on Chapter 25 Grade 11

Introduction to Animals

Sponges. What is the sponge s habitat. What level of organization do sponges have? Type of symmetry?

Invertebrates. Invertebrate Characteristics. Body Symmetry

4/5/15. Myriopods: myriad of legs. Myriapods and Insects CH 14 Subphylum Mandibulata. More on Myriapods:

Phylum Cnidaria (a stinging thread)

Architectural Pattern of an animal. Chapter 9

Folding of the embryo.. the embryo is becoming a tube like structure

An Introduction to the Invertebrates (part 4?!) Annelida & Nematoda. Reference: Chapter 33.3, 33.4

Mesoderm Divided into three main types - Paraxial (somite) - Intermediate - Lateral (somatic and splanchnic)

ANIMAL DIVERSITY AND THE EVOLUTION OF BODY PLANS

The Radiata-Bilateria split. Second branching in the evolutionary tree

Animals are in Domain Eukarya

The Mollusks. Phylum Mollusca

Animals. Chapters Exam November 22, 2011

What Is an Animal? Section 25.1 Typical Animal Characteristics. I. Characteristics of Animals. Biology II Mrs. Michaelsen

Biology 340 Comparative Embryology Lecture 4 Dr. Stuart Sumida. Overview of Pre-Metazoan. and Protostome Development (Insects)

Invertebrate Zoology. Unit 2: Phylums: Porifera, Cnidaria, and Ctenophora

Importance of Taxonomic Collections

Animal Diversity I: Porifera, Cnidaria, Ctenophora, Platyhelminthes, Rotifera, Annelida

Axis Specification in Drosophila

Kingdom Animalia. Special Features: Advanced nervous systems means cephalization (faces), brains, and efficient mobility (walk/run/swim/grab)

Learning Objectives. The Animal Kingdom: An Introduction to Animal Diversity. Sexual Reproduction

Biology: Get out your packet from yesterday! If you would like to use gloves on Mon and Tues for Dissection PLEASE BRING THEM!!!

Are these organisms. animals or not?

SOBA Bee School April, 2015

1 1:29 PM 2 1:29 PM. Porifera Placozoa Cnidaria Ctenophora Platyhelminthes Gastrotricha Gnathostomulida Cycliophora Rotifera Annelida

Transcription:

University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Armand R. Maggenti Online Dictionary of Invertebrate Zoology Parasitology, Harold W. Manter Laboratory of September 2005 Online Dictionary of Invertebrate Zoology: M Mary Ann Basinger Maggenti University of California-Davis Armand R. Maggenti University of California, Davis Scott Gardner slg@unl.edu Follow this and additional works at: http://digitalcommons.unl.edu/onlinedictinvertzoology Part of the Zoology Commons Maggenti, Mary Ann Basinger; Maggenti, Armand R.; and Gardner, Scott, "Online Dictionary of Invertebrate Zoology: M" (2005). Armand R. Maggenti Online Dictionary of Invertebrate Zoology. 14. http://digitalcommons.unl.edu/onlinedictinvertzoology/14 This Article is brought to you for free and open access by the Parasitology, Harold W. Manter Laboratory of at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Armand R. Maggenti Online Dictionary of Invertebrate Zoology by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln.

Online Dictionary of Invertebrate Zoology 528 M macerate v. [L. macerare, to soften] To waste away; to soften or wear away. machopolyp, machozooid see dactylozooid macraner n. [Gr. makros, large; aner, male] (ARTHRO: Insecta) A male ant of unusually large form. macrergate n. [Gr. makros, large; ergate, worker] (ARTHRO: Insecta) In Formicidae, an unusually large worker. macrobiota n. [Gr. makros, large; bios, life] Larger organisms in the soil, such as insects and earthworms. see mesobiota. macrocephalic female (ARTHRO: Insecta) In Hymenoptera, a large female of Halictidae, that possesses a disproportionately large head, usually the egg layers of the colony. macrocercous cercaria (PLATY: Trematoda) Cystophorous type cercaria with a long, simple, cylindrical tail. macrochaetae n.pl. [Gr. makros, large; chaite, hair] Large bristles. see chaetotaxy. macrocilia n.pl.; sing. -ium [Gr. makros, large; L. cilium, eyelash] (CTENO) In Beroida, 2,500-3,500 giant ciliary shafts interconnected and bound together on the mobile lips that function in food gathering. macrocnemes n.pl. [Gr. makros, large; kneme, lower leg] (CNID: Anthozoa) In Actinaria, complete and filamented mesenteries in the first one or two cycles of simple tentacles; maybe fertile or sterile and possess acontia and strong retractors. see microcnemes. macrocyte see plasmatocyte macroesthetes see megalaesthetes macroevolution n. [Gr. makros, large; L. evolvere, to unroll] Evolutionary processes that extend through geologic eras; large scale evolution of new species and genera due to mutations that result in marked changes in chromosomal patterns and reaction systems. see microevolution. macrofauna n. [Gr. makros, large; L. Faunus, diety of herds and fields] 1. Widely distributed; from a macrohabitat. 2. Animals measured in centimeters rather than microscopic units. macrogamete n. [Gr. makros, large; gamete, wife] A large, quiescent, female anisogamete. see microgamete. macrogametocyte n. [Gr. makros, large; gamein, to marry; kytos, container] The infected human red blood cell that contains the female form of the malarial parasite which upon transfer to the Culicidae becomes a macrogamete. macrogamy see hologamy macrogenesis n. [Gr. makros, large; genesis, origin] The sudden origin of new species by saltation. macrogyne n. [Gr. makros, large; gyne, woman] (ARTHRO: Insecta) In Formicidae, a female or queen of unusually large stature. macroic see meganephridia macrolecithal a. [Gr. makros, large; lekethos, egg yolk] With a large amount of yolk. see microlecithal. macromere n. [Gr. makros, large; meros, part] A distinctly large cell resulting from unequal cleavages during early embryology. macromesentery n. [Gr. makros, large; mesos, middle; enteron, gut] (CNID: Anthozoa) One of the larger complete mesenteries. macromitosome n. [Gr. makros, large; mitos, thread; soma, body] (ARTHRO: Insecta) The paranucleus as seen in Lepidoptera. macromolecule n. [Gr. makros, large; L. moles, mass] Very large molecules such as protein, cellulose, starch, etc. macromutation n. [Gr. makros, large; mutare, to change] Theory of instantaneous evolution of new taxa by a mutation that establishes reproductive isolation at once. macronotal a. [Gr. makros, large; notos, back] (ARTHRO: Insecta) Having a large thorax, as a queen ant.

529 Online Dictionary of Invertebrate Zoology 530 macronucleocyte see prohemocyte macrophage n. [Gr. makros, large; phagein, to eat] A large phagocytic cell of the body. macrophagous a. [Gr. makros, large; phagein, to eat] Feeding on large objects. see microphagous. macrophthalmic a. [Gr. makros, large; ophthalmos, eye] Having large eyes; having eyes larger than normal. macroplankton n. [Gr. makros, large; planktos, wandering] Large organisms such as jellyfish that drift with the currents. macropore see megalopore macropseudogyne see pseudogyne macropterous a. [Gr. makros, large; pteron, wing] (ARTHRO: Insecta) Having a long or large wing. see brachypterous. macrosclere see megasclere macroscopic, macroscopical a. [Gr. makros, large; skopein, to view] Capable of being studied with the unaided eye; megascopic. see microscopic. macroseptum n. [Gr. makros, large; L. septum, partition] 1. (CNID: Anthozoa) The variously functioning primary septum. 2. (NEMER) In asexual reproduction, a partition across the body marking the plane of subsequent fragmentation. macrosiphon n. [Gr. makros, large; siphon, tube] (MOLL: Cephalopoda) Internal siphon of certain cuttlefishes, and all octopuses. macrosymbiont n. [Gr. makros, large; symbios, living together] The larger of two symbiotic organisms. macrotaxonomy n. [Gr. makros, large; taxis, arrangement] The classification of higher taxa. macrotrichia n.pl.; sing. -ium [Gr. makros, large; thrix, hair] 1. The larger surface hairs. 2. (ARTHRO: Insecta) The large hairs on the wing membrane. macrotype n. [Gr. makros, large; typos, type] (CNID: Anthozoa) Modified arrangement of mesenteries consisting mainly of macromesenteries. see microtype. macrurous a. [Gr. makros, large; oura, tail] Long-tailed. macula n.; pl. maculae [L. macula, spot] 1. A colored spot of rather large size. 2. A spot level with surrounding surface. 3. (BRYO: Stenolaemata) Prominences, and less commonly flat or depressed areas on colony surfaces regularly spaced among feeding zooids caused by clusters of a few polymorphs, and/or extrazooidal skeleton. see monticule. 4. (MOLL: Cephalopoda) An oval spot on the wall of a statocyst to which a calcareous statolith is attached; gives information on position relative to gravity. maculate a. [L. macula, spot] Splashed or spotted; blotched. madrepore n. [L. mater, mother; Gr. poros, friable stone] (CNID) A stony, branched, reef building coral of the order Madreporia. madreporic plate (ECHINOD: Asteroidea) An enlarged interradial plate on the disc, that connects the water vascular system to the sea. madreporite n. [L. mater, mother; porus, pore] (ECHINOD) An oral or aboral perforated plate of the water-vascular system connecting with the stone canal; sieve plate. main bud (BRYO: phylactolaemata) The largest of three bud primordia which occurs on every mature zooid, and is the first to form a new polypide. major gene Controls production of qualitative phenotypic effects in contrast to its modifiers. major worker (ARTHRO: Insecta) The largest worker subcaste in social insects; in Formicidae, usually specialized for defense and referred to as a soldier. see media worker, minor worker. mala n.; pl. malae [L. mala, cheek, jaw] 1. A lobe; ridge or grinding surface. 2. (ARTHRO) a. Part of the maxilla of certain insects. b. Mandible of some myriapods. malacoid a. [Gr. malakos, soft] Soft textured. malacology n. [Gr. malakos, soft; logos, discourse] The branch of zoology dealing with mollusks, the animal inside the shell. malacophilous a. [Gr. malakos, soft; philios, loving] (MOLL: Gastropoda) Being pollinated by the action of gastropods.

531 Online Dictionary of Invertebrate Zoology 532 malapophysis n.; pl. -ses [L. mala, cheek, jaw; Gr. apophysis, projection] (ARTHRO: Chelicerata) In Acari, the paired anterior region of the infracapitulum. malar a. [L. mala, cheek] Of or about the cheek region. malar cavity (ARTHRO: Chelicerata) In Acari, interior of the malapophysis that connects to the pharynx. male n. [L. mas, a man] An individual that produces sperm cells but not egg cells; designated by. male-cell receptacle (ARTHRO: Crustacea) In Rhizocephala, a pocket or pair of pockets within the mantle cavity of the female where cells of male cyprid undergo spermatogenesis. male ducts, male gonoducts see sperm ducts male funnel (ANN: Oligochaeta) A funnel or rosette-shaped enlargement of the ental end of the sperm duct for passage of sperm through the central aperture into lumen of the duct on their way to the exterior. malella n. [L. dim. mala, jaw] (ARTHRO: Symphyla) The distal toothed process on the outer stipes of the deutomala of some myriapods. male tube (ECHI) A sexually maturing area in which developing males are housed for one to two week inside the female. malleate mastax (ROTIF) Chewing apparatus; rami untoothed and unci are curved plates with prong type teeth. malleations n.pl. [L. dim. malleus, hammer] A hammered appearance. malleolus n.; pl. -li [L. dim. malleus, hammer] 1. (ARTHRO: Chelicerata) In Solpugida, 3 to 5 innervated appendages on the coxae and trochanter of the fourth leg; function uncertain; racket-organs. 2. (ARTHRO: Insecta) see haltere. malleoramate mastax (ROTIF: Monogononta) Chewing apparatus, variant of the ramate type of mastax occurring in the order Flosculariaceae. malleus n.; pl. malli [L. malleus, hammer] (ROTIF) Unci and manubria of the mastax, collectively. Malpighian tubules (ARTHRO) Long, thin excretory tubules extending into the body cavity from the posterior region of the gut in insects, arachnids and myriapods. maltha see mesogloea mamelon n. [F. mamelon, protuberance] 1. (ECHINOD: Echinoidea) Terminal knob on the boss that articulates with the spine on the test. 2. (NEMATA: Secernentea) Two or three ventral, serrated projections on the ventral surface of the male of the genus Syphacia; function unknown. mammillate a. [L. mamilla, small breast] Having rounded protuberances or wart-like projections; mammiform; mammose. manca n. [L. mancus, imperfect] (ARTHRO: Crustacea) In Peracarida, a juvenile or postlarva that lack the last thoracopod when released from the marsupium. manchette n. [F. manchette, cuff] (NEMATA) Line of demarcation between the cheilostome and esophastome; nema's collar; sleeve. mancoid stage (ARTHRO: Crustacea) Postlarval stage in Leptostraca, with rudimentary 4th pleopod. mandible n. [L. mandibula, jaw] 1. A jaw. 2. (ANN: Polychaeta) The ventral chitinous plates or rods, maybe dentate, against which the maxilla work. 3. (ARTHRO: Crustacea) One of the third pair of cephalic appendages. 4. (ARTHRO: Diplopoda) The first pair of jaws, consisting of cardo, stipes and gnathal lobe. 5. (ARTHRO: Insecta) One of the variously modified anterior pair of paired mouthpart structures. 6. (BRYO) A modified orificial wall of the avicularium. see operculum. mandibular fossa (ARTHRO: Insecta) The dorsal articulation of the mandible. mandibular gland (ARTHRO: Insecta) A sac-like reservoir usually paired, and partially or completely lined by secretory cells, opening in the mesal junction of the mandible with the head; functioning as alarm pheromone, fungistatic agent, aggregation pheromone, sex pheromone, territory defending secretion, location of food and/or the 'burning' secretion of the 'fire bees'; in larval Lepidoptera, large and secrete saliva (with the normal salivary glands specialized

533 Online Dictionary of Invertebrate Zoology 534 for silk production). mandibular palp 1. (ARTHRO: Crustacea) In Cirripedia, distally articulated part of the mandible functioning in feeding or cleaning; in Acrothoracica associated with the mandibular gnathobase; setose lobe on labrum. 2. (ARTHRO: Insecta) see prostheca. mandibular plates (ARTHRO: Insecta) In Hemiptera, plates between and attached to the mandibular stylets and the ventral surface of the sucking pump. mandibular pouch (ARTHRO: Insecta) In Thysanoptera, a ventrally oriented cone formed by the labrum and labium containing maxillary stylets and a single (left) functional mandible, emerging at the apex. mandibular ring (NEMATA: Adenophorea) In Enoplida, transverse extension of the mandibles in the buccal cavity. mandibular scar (ARTHRO: Insecta) In certain Coleoptera pupae, round or oval areas with raised margins that serve as supports for the deciduous provisional mandibles. mandibular sclerite (ARTHRO: Insecta) In some larval Diptera, mouth-hooks articulating basally with the intermediate (hypostomal) sclerites. Mandibulata n. [L. mandibula, jaw] Formerly a subphylum of arthropods including Myriapoda, Crustacea and Insecta. mandibulate a. [L. mandibula, jaw] Having jaws fitted for chewing; mandibuliform. manducate v. [L. manducare, to chew] To bite; eat. manica n. [L. manica, sleeve] (ARTHRO: Insecta) In Lepidoptera, the inner layer of the anellus, that fastens around the aedeagus. maniform a. [L. manus, hand; forma, shape] Hand-shaped. manitruncus see prothorax manna n. [Gr. manna, morsel or honey dew] (ARTHRO: Insecta) Honeydew (90-95% sugar) produced by certain coccids, used as human food. manometabola see hemimetabolous metamorphosis mantle n. [L. mantellum, cloak] 1. Something that enfolds, envelopes or covers. 2. (ARTHRO: Crustacea) In thoracic Cirripedia, membranous covering of the body, often strengthened by calcareous plates. 3. (BRACHIO) Prolongation of the body wall as fold of ectodermal epithelium. 4. (BRYO: Phylactolaemata) Ciliated fold of colony wall covering one to four small, sexually produced colony progenitor polypides. 5. (MOLL) A membranous covering that secretes the shell; the marginal glands produce the periostracum; pallium. mantle canal (BRACHIO) One of several flattened, tube-like extensions of the body cavity into each mantle lobe. mantle cavity Specialized cavity found in certain mollusks, brachiopods and crustaceans lined with epidermis and usually exposed to sea or fresh water, or air, due to habitat; may or may not contain part of the viscera; pallial chamber or cavity; mantle chamber. mantle cells (ARTHRO: Insecta) Corneagenous cells of the eye enclosing the retina. mantle fold (MOLL: Bivalvia) In oysters, one of 3 small folds at the edge of a mantle lobe. mantle groove (BRACHIO) Site of proliferation of the periostracum and bears the setae. mantle lobe 1. (BRACHIO) Lobe of the body wall that secretes and lines the valves; in some, cecae project into perforations (punctae) in the shell serving as food repositories of glycoproteins and mucroproteins, inhibitors of boring organisms, and accessory respiratory structures. 2. (MOLL: Bivalvia) In oysters, one of 2 thin epithelial extensions of the mantle adjoining the valve. mantle papilla see caecum mantle skirt (MOLL) Roof of the mantle cavity formed by the projection of the mantle from the edge of the visceral mass. manubrium n. [L. manubrium, handle] 1. (ARTHRO: Crustacea) In fish parasitic Copepoda, the handle or distal part of the attachment organ. 2. (ARTHRO: Insecta) The basal part of the furcula of collembolan springtails; part of the sternum associated with the cavity of the prothorax of Coleoptera.

535 Online Dictionary of Invertebrate Zoology 536 3. (CNID: Hydrozoa) The extension between the stomach cavity and the mouth of a medusae or polyp; also called gullet or esophagus. 4. (NEMATA) The proximal portion of a spicule; a capitulum. 5. (ROTIF) One of two paired trophi in the mastax. manus n. [L. manus, hand] 1. The hand. 2. (ARTHRO: Crustacea) Broad proximal part of a propodal cheliped of a Decapoda. 3. (ARTHRO: Insecta) Formerly applied to the anterior tarsus. 4. (MOLL: Cephalopoda) see tentacle. manuscript name An unpublished scientific name. see nomen nudum. marble gall (ARTHRO: Insecta) The hard spherical gall of Cynipidae Adleria kollari, usually on oak that produce the agamic generation of that species. margaritaceous a. [Gr. margarites, a pearl] Pearly in texture; nacreous. margin n. [L. margo, border] 1. A border or an edge. 2. (AR- THRO: Insecta) The edge of a wing. 3. (CNID: Anthozoa) The junction of the oral disk and collum of a sea anemone. 4. (MOLL) The edge of a shell. marginal a. marginal bodies (PLATY: Trematoda) In Aspidogastrea, sensory pits or short tentacles between marginal loculi of the opisthaptor. marginal bristles (ARTHRO: Insecta) In Diptera, abdominal bristles inserted dorsally on the margins of the segment. marginal cell (ARTHRO: Insecta) A cell in the distal part of the wing bordering the costal margin. marginalia n. [L. margo, border] (PORIF) Spicules protruding upward around an oscule. marginal nuclei (NEMATA) Nuclei of marginal cells in the nematode esophagus; believed to lay down the fibers from the apex of the lateral arm to the basement membrane of the esophagus, or to secrete the cuticular lining of the esophagus, or both. marginal tubes (NEMATA) The distal cylindric endings of some esophageal radii, other forms have convergent terminals. marginal carina (MOLL: Bivalvia) A ridge running from umbo to posteroventral angle of the shell delimiting the posterior area in Trigoniacea. marginal veins (ARTHRO: Insecta) A vein running along the front margin of a wing that gives off a stigmal vein. marginate a. [L. margo, border] Having a distinct margin in appearance or structure. marine a. [L. mare, sea] Pertaining to or inhabiting the sea, ocean, or other salt waters. marita n. [L. maritus, conjugal] (PLATY: Trematoda) A sexually mature fluke. marker gene Genetic marker; gene of known position and conspicuous in its action. marmorate a. [L. marmor, marble] Having color or veined like marble. marsh n. [A.S. mersc, marsh] An area of wet soil. marsupium n. [L. marsupium, bag] 1. Brood pouch. 2. (AR- THRO: Crustacea) The oostegite in Peracarida, or brood chamber or pouch in others. 3. (ARTHRO: Insecta) In marsupial coccids, a waxen ovisac, longer than the body, filled with eggs that hatch while the insect is still mobile. 4. (BRYO) The ovicell. 5. (CNID: Hydrozoa) In some, the internal pouch borne by the blastostyle. 6. (ECHINOD) a. In some, the cardiac stomach. b. In Viviparous Crinoidea, present on the base of the pinnules adjacent to the gonad and having an external opening; a brood chamber. mask n. [Ar. maskhara, buffoon] (ARTHRO: Insecta) In dragonfly nymphs, the prehensile labium that conceals the other mouthparts. masked a. masked pupa see pupa larvata mass communication (ARTHRO: Insecta) In Hymenoptera, transfer of information among large groups of individuals which cannot be accounted for by one to one communication. mass provisioning (ARTHRO: Insecta) Social behavior of solitary bees and wasps by storing cells with sufficient food to satisfy their developing offspring and closing them down before the eggs hatch. see progressive provisioning.

537 Online Dictionary of Invertebrate Zoology 538 mastax n. [Gr. mastax, jaws] (ROTIF) A muscular rounded, trilobed, or elongate organ containing trophi; in suspension feeders adapted for grinding, in carnivores modified as forceps that can be projected from the mouth to seize prey; the pharynx. mastication n. [L. mastecare, to chew] The act of chewing; to grind or crush. masticatory process see gnathal lobe masticatory stomach see gastric mill masticomorphic a. [L. mastecare, to chew; Gr. morphos, form] Designed for chewing. mastidia n.pl.; sing. mastidion [Gr. mastos, breast] (ARTHRO: Chelicerata) Small, conical, nipple-like tubercles on the front of the chelicerae of small spiders. mastigobranch, mastigobranchia n. [Gr. mastix, whip; branchos, gill] (ARTHRO: Crustacea) A slender respiratory process at the base of the epipod. mastigophore n. [Gr. mastix, whip; pherein, to bear] (CNID: Anthozoa) A nematocyst with a cylindrical hempe and tube extending beyond the hempe; microbasic with hempe not more than 3 times the capsule length; macrobasic with hempe 4 or more times the capsule length. mastigopus larva (ARTHRO: Crustacea) Larva in the megalopa stage found among some Decapoda. maternal inheritance Inheritance controlled by maternal extra chromosomal determinants. maternal zooid (BRYO: Gymnolaemata) An autozooid that extrudes eggs. mating plug (ARTHRO: Insecta) A plug formed from the accessory gland secretions of the male, deposited in the genital chamber of the female, thought to prevent loss of sperm in some Culicidae and Lepidoptera; also called spermatophragma. see sphragis. mating spines (ARTHRO: Insecta) In female mayflies, compound conical spines covering the lower surface of the egg valve. matricidal hatching (NEMATA) Intrauterine larval development leading to the destruction of the female by the larvae or juveniles; eclosion intrauterine. matrifilial a. [L. mater, mother; filia, daughter] (ARTHRO: Insecta) In Apis, having colonies made up of mothers and daughters. matrix n.; pl. matrices [L. mater, mother] 1. That which gives form, origin or foundation to something enclosed or embedded in it. 2. (NEMATA: Secernentea) The gelatinous substance secreted by some female nematodes into which eggs are deposited to form an egg mass. 3. (PLATY) In Cestoda and Trematoda, a living interface of interacting photoplasmic layer; part of the syncytium of the tegument. 4. (PORIF) The non-cellular ground material of a sponge in which the cellular elements are dispersed. matrix glands (NEMATA: Secernentea) An excretory cell or modified rectal glands that secrete the gelatinous matrix through the anus or excretory pore and into which eggs maybe imbedded. matrix layer (NEMATA) Historically, a cuticular stratum of spongy material between the fibrillar layer and the boundary layer. matrone n. [L. mater, mother] (ARTHRO: Insecta) Macromolecular components (proteins) contained in the seminal fluid of some male Culicidae that inhibits further insemination of the female. maturation n. [L. maturus, ripe] The act or process pertaining to the developmental steps leading to reproducing adults. maturation divisions A series of nuclear divisions in the formation of the gametes in which the chromosome number, through meiosis is reduced from diploid to haploid. maturation feeding (ARTHRO: Insecta) Feeding required by some insects before their gonads can mature to produce eggs. maturation zone In males of many invertebrates, that part of the genital follicle below the germarium in which each spermatocyte undergoes the two meiotic divisions to produce spermatids. see transformation zones. mature region see exozone

539 Online Dictionary of Invertebrate Zoology 540 maxaponta n. [L. maxilla, jaw; pons, bridge] (ARTHRO: Insecta) A bridge formed by a midline fusion of the lower maxillariae and the postgenae. maxapontal a. maxilla n.; pl. -llae [L. maxilla, jaw] 1. (ANN: Polychaeta) The large, hook-shaped, dorsal chitinous jaw plate. 2. (ARTHRO) In most arthropods, one of the paired mouth-part structures posterior to the mandibles or jaws; third pair of head appendages. 3. (ARTHRO: Diplopoda) One of the paired second jaws to form the gnathochilarium. maxillary a. [L. maxilla, jaw] (ARTHRO) Of or pertaining to the maxilla. maxillary carrier (ANN: Polychaeta) A posterior support structure for the maxilla. maxillary glands (ARTHRO) Glands belonging to the maxillary segment, possibly functioning in the lubrication of the mouthparts. maxillary guides (ARTHRO: Insecta) In Anoplura, paired structures of wrinkled sucking lice arising from the maxillary appendages that function as guides for the dorsal stylet. maxillary palp/palpus; pl. -pi (ARTHRO: Insecta) Small sensory organ arising from the maxilla, used to test quality of food. see labial palp. maxillary segment see maxillulae maxillary stylets (ARTHRO: Insecta) In Hemiptera, the inner pair of stylets of the trophic sac. maxillary tentacle (ARTHRO: Insecta) In female Tegeticula Lepidoptera, an inner elongate lobe of the maxilla, adapted for holding a large mass of pollen; perhaps palpifers. maxilliped, maxillipede, maxillipe a. [L. maxilla, jaw; pes, foot] (ARTHRO: Crustacea) The paired appendages on thoracic somites 1-3 posterior to the maxillae, that usually function in feeding; sometimes adapted for other functions such as prehension in parasitic forms. maxillulae n.pl.; sing. -ula [L. dim. maxilla, jaw] 1. (ARTHRO: Crustacea) The first maxillae having more than one pair of maxillae; paragnath. 2. (ARTHRO: Insecta) In primitive forms, the non-functional appendages between mandibles and first maxillae. see superlinguae. maxim n. [L. maximus, greatest] (ARTHRO: Insecta) In Formicidae, a major worker or one of the soldier caste. mayrian furrow see notaulix meatal cleft (ARTHRO: Insecta) A slit or line on the trumpet of some mosquito pupae extending into the meatus from the spiracular opening, facilitating enlargement of the opening at the surface of the water. meatus n. [L. metus, passage] A channel or duct. mechanical isolation Reproductive isolation due to mechanical incompatibility of male and female genitalia. mechanoreceptor n. [Gr. mechane, contrivance; L. recipere, to receive] Specialized structures that perceive any mechanical distortion of the body, i.e., touch, vibrations, altitude and gravity. meconida n.pl.; sing. -ium [Gr. mekon, poppy] 1. (ARTHRO: Insecta) Waste products of pupal metabolism that are discharged shortly after adult emergence. 2. (CNID: Hydrozoa) Medusoid gonophores, sessile or pedicellate, which upon emergence from the gonangium act as external brood sacs. meconium n. media n. [L. medius, middle] 1. The middle structure. 2. (AR- THRO: Insecta) The longitudinal vein between the cubitus and the radius of the wing. medial a. mediad adv. [L. medius, middle; ad, toward] Toward the median plane or line; mesad; admedial. medial cross vein (ARTHRO: Insecta) A cross vein connecting two branches of the media of the wing. medial-cubital cross vein (ARTHRO: Insecta) A cross vein of an insect wing between the posterior medial vein and the anterior cubital vein. median a. [L. medius, middle] In the middle; along the midline of the body; middle variate when variates are arranged in order of magnitude. median bulb see metacorpus

541 Online Dictionary of Invertebrate Zoology 542 median caudal nerve (NEMATA) A nerve extending from the dorsorectal ganglion to the tail. median cercus see urogomphus median cord (ARTHRO: Insecta) An embryonic chain of cells derived from the ectoderm lining the neural groove. median dorsal plate (ARTHRO: Crustacea) In some Peracardia, an elongate plate separating carapace valves posterodorsally. median esophageal bulb see metacorpus median eye see nauplius eye median lamina see median wall median latus (ARTHRO: Crustacea) In Lepadomorpha, a plate between the rostral and carinal latera in forms with paired latera in one whorl. see lateral. median ligament (ARTHRO: Insecta) A common thread formed by the ovarioles of opposite sides that help maintain the ovaries in position and is attached to the body-wall, the fat-body or the pericardial diaphragm. median oviduct see common oviduct median segment (ARTHRO: Insecta) The basal segment of the abdomen when it is fused with the metathorax during the change from larva to pupa. see propodeum, epinotum. median tubuli (BRYO: Stenolaemata) Aligned pustules or mural lacunae in a laminated skeleton. median valve see intermediate valve median wall (BRYO: Stenolaemata) Erect colony wall parallel to the growth direction from which zooids bud to form a bifoliate colony. mediator n. [L. medius, middle] Association, internuncial, neuron; chemical such as a hormone that controls or modifies a metabolic process. media worker (ARTHRO: Insecta) In Formicidae, an individual belonging to the medium-size subcaste in a polymorphic series of three or more worker subcastes. see minor worker, major worker. medio-cubital cross vein see medial-cubital cross vein mediolateral nerve cord (NEMATA) Several nerves extending from the median and posterior externolateral ganglia and the posterior internolateral ganglia to the lumbar ganglia. mediotergite n. [L. medius, middle; tergum, back] (ARTHRO: Insecta) In Diptera, the median region of the mesopostnotum. medioventral a. [L. medius, middle; venter, belly] In the middle ventral line. mediproboscis see haustellum medulla n. [L. medulla, marrow, pith] 1. Central portion of an organ. 2. (ARTHRO) Apical lobes of the brain; the central synaptic region; epiopticon; medulla externa; external medullary mass. medulla externa see medulla medulla interna (ARTHRO: Insecta) The lobular complex. medulla X-organ, medulla terminalis ganglionic X-organ, MTGX (ARTHRO: Crustacea) A group of neurosecretory cells in the medulla terminalis; the main source of eyestalk hormones. medusa n.; pl. -sae [L. Medusa, a gorgon of mythology] (CNID) The free swimming umbrella-like forms. megabenthos see abyssobenthos megacephalic a. [Gr. megas, large; kephale, head] Having an abnormally large head. see microcephalic, mesocephalic. megaclad n. [Gr. megas, large; klados, branch] (PORIF) In megascleres, a relatively large smooth desma; megaclone. megaclone see megaclad megadrile n. [Gr. megas, large; drilos, worm] (ANN: Oligochaeta) Terrestrial forms; not used systematically, although recognized as a general term. see microdrile. megalaesthetes n.pl. [Gr. megas, large; aisthesis, sensation] (MOLL: Polyplacophora) Large sensory organs terminating in the tegmentum in the forms of eyes with cornea, lens, pigment layers, iris and retina. see micraesthetes. megalolecithal see macrolecithal

543 Online Dictionary of Invertebrate Zoology 544 megalopa stage (ARTHRO: Crustacea) 1. Larvae of Malacostraca with functional pleopods; also referred to as glaucothoe. 2. First postlarval stage in development of Eucarida, not present in other crustaceans. 3. Originally applied to Brachyura larvae with large stalked eyes and functional pleopods. megalopore, megapore n. [Gr. megas, large; poros, pore] (MOLL: Polyplacophora) Large pore in the dorsal plate; associated with aesthete. megalops see megalopa stage megamere see macromere megameric a. [Gr. megas, large; meros, part] With relatively large parts; autosomes with large heterochromatic segments. meganephridia n.pl.; sing. -ium [Gr. megas, large; dim. nephros, kidney] (ANN: Oligochaeta) A pair of large nephridia in each segment of the body except, for first and last segments; holonephridia; holoic; macroic. see nephridium, micronephridia. megaplankton see macroplankton megasclere n. [Gr. megas, large; skleros, hard] (PORIF) A large structural spicule. see microsclere. megascolecin n. [Gr. megas, large; skolex, worm] (ANN: Oligochaeta) The single pair of prostates, tubular or racemose, opened to the exterior, along side of or together with the sperm ducts. megascopic see macroscopic megetic a. [Gr. megas, large] Pertaining to size variations in polymorphic forms. see epimegetic, eumegetic, hypomegetic. Mehlis' glands (PLATY: Trematoda) Unicellular mucous and serous glands surrounding the ootype in the reproductive system. meiocyte n. [Gr. meion, smaller; kytos, container] Primary oocytes and spermatocytes. meiofauna n. [Gr. meion, smaller; L. Faunus, diety of herds and fields] Microscopic and small macroscopic fauna on the sea bottom. meiolecithal a. [Gr. meion, smaller; lekithos, egg yolk] Having little yolk. meiomery n. [Gr. meion, smaller; meros, part] The condition of possessing fewer than the normal number of parts. meiosis n.; pl. meioses [Gr. meiosis, to make smaller] Two successive cell divisions in the developing germ cells characterized by the pairing and segregation of homologous chromosomes, resulting in reduction from a diploid number to a haploid one. meiotic a. meiotic drive A meiotic mechanism of cell division resulting in two kinds of gametes produced by a heterozygote with unequal recovery. meiotrichy n. [Gr. meion, smaller; thrix, hair] Loss of setae in ontogenetic development or of homologous setae in natural groups. melania n. [Gr. melas, black] Blackness. melanin n. [Gr. melas, black] A term for a group of chemically ill-defined pigments, often found associated with protein, produced by insects and marine animals responsible for colors from brown to black. melanoid a. melanism n. [Gr. melas, black; ismos, denoting condition] 1. An excessive darkening of color owing to increased amounts of black pigment. 2. A certain percentage of individuals in a population that give rise to polymorphism. see industrial melanism, albinism. melanoid a. [Gr. melas, black; eidos, like] Looking black or dark. meliphagous a. [Gr. meli, honey; phagein, to eat] Honeyeating; melivourous. melittology n. [Gr. melitta, honeybee; logus, discourse] The study of bees. melittophily n. [Gr. melitta, honeybee; philos, love] (ARTHRO: Insecta) 1. A symbiont of social bees. 2. Any organism that must spend a portion of its life cycle in a bee colony. melivorous see meliphagous

545 Online Dictionary of Invertebrate Zoology 546 mellifera n. [L. mel, honey; ferre, to bear] (ARTHRO: Insecta) Honey-makers; bees as a whole. melliferous a. [L. mel, honey; ferre, to bear] Honey producing. mellisugent a. [L. mel, honey; sugere, to suck] Honey-sucking. member n. [L. membrum, part] A limb or organ. membrane n. [L. membrana, skin] A thin film of tissue. membranization n. [L. membrana, skin] Changed into a membrane. membranous a. [L. membrana, skin] Consisting of membranes; soft and pliable; membranaceous. membranous sac (BRYO: Stenolaemata) Membrane surrounding digestive and reproductive systems of zooid; the entosaccal and exosaccal cavity. membranule, membranula n. [L. dim. membrana, skin] (AR- THRO: Insecta) A small semi-opaque membrane on the base of the hind wing of certain Odonata. Mendelian character Character formed under the control of chromosomal genes. Mendelian inheritance Mode of inheritance from chromosomal genes. Mendelian mutation True gene mutation and recombination. Mendelian population A population with unrestricted interbreeding of organisms sharing a common gene pool. Mendelism n. [Gregor Mendel] Particulate inheritance of chromosomal genes. Mendel's laws of inheritance Genetic principles proposed by Mendel; law of segregation and law of independent assortment. meniscoidal a. [Gr. meniskos, a crescent] 1. Crescent-shaped lens; concavo-convex lens; one side convex and the other concave. 2. With one side concave, crescent shape of water in a tube, or convex, crescent-shaped as with mercury in a tube. menognath n. [Gr. menein, to remain; gnathos, jaw] (AR- THRO: Insecta) Having biting mandibles in both larval and adult stages. menognathous a. see Menorhyncha, metagnath. Menorhyncha n.pl. [Gr. menein, to remain; rhynchos, snout] (ARTHRO: Insecta) A former division of insects composed of those who ingest by suction in both larval and adult stages. see menognath and metagnath. menotaxis n. [Gr. menein, to remain; taxis, arrangement] Orientation in a fixed direction with respect to the stimulus. mental a. [L. mentum, chin] Of or pertaining to the mentum. mental plate (ARTHRO: Insecta) In Lepidoptera, representing the basal sclerites of the labium. see labial plate. mental setae (ARTHRO: Insecta) Setae located on the mentum. mental suture (ARTHRO: Insecta) A distinct sclerite defined by a suture intervening between the mentum and the gula. mentasuture see mental suture mentigerous a. [L. mentum, chin; gerere, to bear] (ARTHRO: Insecta) Having a mentum. mentum n. [L. mentum, chin] 1. (ARTHRO: Diplopoda) A median, slightly triangular sclerite in the gnathochilarium. 2. (ARTHRO: Insecta) a. The distal sclerite of a typical insect labium, bearing the palps and the ligula. b. In bees, the second joint bearing the palps, paraglossa and ligula. meraspis larva (ARTHRO: Trilobita) The second larval stage with the pygidium located behind the cephalon; the thoracic region will appear during succeeding molts. see holaspis larva. merdivorous see scatophagous mereopodite see merus meridional canal (CTENO) One of the eight canals extending in an oral-aboral direction under the external surface; part of the gastrovascular system; in Pleurobranchia it emits a greenish-blue luminescence. meristal annuli (ARTHRO: Insecta) In Orthoptera and Odonata, annuli derived from, and adjacent to, the meriston that di-

547 Online Dictionary of Invertebrate Zoology 548 vides. meriston n. [L. merizein, to divide] (ARTHRO: Insecta) The most basal annulus of the antennal flagellum. meritrichy a. [Gr. meros, part; trichos, hair] (ARTHRO: Chelicerata) In Acari, chaetotaxy characterized by a reduction in number and size of setae from the holotrichous form. mermithaner n. [Gr. mermis, cord; aner, male] (ARTHRO: Insecta) Male Formicidae parasitized by the nematode Mermis; a mermithophore. mermithergate n. [Gr. mermis, cord; ergates, worker] (AR- THRO: Insecta) In Formicidae, a worker parasitized by the nematode Mermis; a mermithophore. mermithized a. [Gr. mermis, cord] (NEMATA) Pertaining to parasitism by nematodes of the genus Mermis. mermithodinergate see mermithostratiote mermithogyne n. [Gr. mermis, cord; gyne, woman] (ARTHRO: Insecta) A female Formicidae parasitized by the nematode Mermis; a mermithophore. mermithophore n. [Gr. mermis, cord; pherein, to carry] (NEMATA) An anomalous form resulting from parasitism by the nematode Mermis; a mermithaner, mermithergate, mermithogyne, mermithostratiote. mermithostratiote n. [Gr. mermis, cord; stratiotes, soldier] (ARTHRO: Insecta) Soldier Formicidae parasitized by the nematode Mermis; a mermithophore. mermitoid esophagus see stichosome meroandry n. [Gr. meros, part; aner, male] The condition of possessing less than the normal number of testes. meroandric a. see holandry. meroblastic cleavage Cleavage of a heavily yolked egg in which only the egg cell divides, leaving the yolk undivided. merocerite n. [Gr. meros, part; keras, horn] (ARTHRO: Crustacea) The 4th segment of an antenna. merocrine a. [Gr. meros, part; krinein, to separate] The passing of a secretion by a gland in which the nucleus remains intact and thereby can recover. see holocrine. merognathite see merus meroic a. [Gr. meros, part] (ANN: Oligochaeta) Pertaining to the excretory system with nephridial tubules formed by longitudinal or transverse fragmentation of the original single pair of embryonic rudiments of each segment. meroistic ovariole (ARTHRO: Insecta) An ovariole in which nurse cells, or trophocytes are present; telotrophic (acrotrophic) and polytrophic types; panoistic; meroistic egg tube. meromyarian a. [Gr. meros, part; mys, muscle] (NEMATA) Muscle arrangement with only a few, frequently only two, flat muscle cells seen in each quadrant of a cross section of the animal. meron n. [Gr. meros, upper thigh] (ARTHRO: Insecta) The posterior part of the basicoxite; in higher Diptera, separated from the coxa and forms part of the thoracic wall (mesomeron, metameron). meronephridium see micronephridia meropleuron n.; pl. -ura [Gr. meros, part; pleuron, side] (AR- THRO: Insecta) A sclerite composed of the meron of the coxa and the lower region of the epimeron. meropodite n. [Gr. meros, part; pous foot] (ARTHRO) 1. The fourth segment of a generalized limb. 2. The femur in Chelicerata and Insecta. 3. For Crustacea see merus. merosome n. [Gr. meros, part; soma, body] A body segment; a somite or metamere. merospermy n. [Gr. meros, part; sperma, seed] Fusion of an egg cell with a sperm that has lost its nucleus; therefore, it cannot take part in karyogamy with the egg nucleus. merus n. [Gr. meros, part] (ARTHRO: Crustacea) The fourth segment of the mouth part, articulating with the ischium anteriorly and carpus posteriorly; a meropodite. mesad, mesiad adv. [Gr. mesos, middle; ad, toward] Toward the midline of the body. mesadenia n.pl.; sing. mesadene [Gr. mesos, middle; aden, gland] (ARTHRO: Insecta) 1. Mesodermal accessory glands of male genitalia. 2. In some male Heteroptera, paired,

549 Online Dictionary of Invertebrate Zoology 550 highly coiled tubules that run side by side to enter anteriorly the bulbus ejaculatorius or closely associated with the short vasa deferentia. mesal, mesial a. [Gr. mesos, middle] At or near the midline of the body. mesal penellipse (ARTHRO: Insecta) In larvae, a series of crochets covering at least the mesal half of the proleg, incomplete laterally. see penellipse. mesanapleural suture (ARTHRO: Insecta) In Diptera, a suture between the mesanepisternum and the meskatepisternum. mesanepisternum n. [Gr. mesos, middle; ana, up; epi, on; sternon, chest] (ARTHRO: Insecta) 1. In Diptera, the upper area of the mesepisternum separated from the meskatepisternum by the mesanapleural suture; sometimes divided into anterior and posterior by anepisternal cleft; the anepisternum. 2. In Odonata, the anepisternum. mesaxon n. [Gr. mesos, middle; axon, axel] (ARTHRO: Insecta) The spiral arrangement of a Schwann cell around an axon; suspensory fold. mesenchymatous cell see hemocyte mesenchyme, mesenchyma n. [Gr. mesos, middle; enchyma, infusion] Embryonic connective tissue derived primarily from mesoderm and consisting of a diffuse network of loosely connected or scattered cells not segregated into layers or blocks. mesenteron n. [Gr. mesos, middle; enteron, gut] The midgut or midportion of the alimentary tract, endodermal in origin; ventriculus; midintestine. mesenteron rudiments (ARTHRO: Insecta) Groups of embryologic endodermal cells that regenerate the midgut (stomach) including the anterior and posterior; becomes the epithelium of the adult mesenteron. mesentery n. [Gr. mesos, middle; enteron, gut] A supporting membrane or one that forms a partition. mesepimeral scale or seta (ARTHRO: Insecta) Any scale or seta borne on the mesepimeron. mesepimeral suture see mesopleural suture mesepimeron n.; pl. -mera [Gr. mesos, middle; epi, on; meros, part] (ARTHRO: Insecta) 1. The area of the mesopleuron posterior to the mesopleural suture; the epimeron of the mesothorax. 2. In Odonata, the area between the humeral and first lateral suture. mesepisternum n.; pl. -sterna [Gr. meros, middle; epi, on; sternum, chest] (ARTHRO: Insecta) 1. The area of the mesopleuron anterior to the mesopleural suture; sometimes divided into an upper mesanepisternum and a lower meskatepisternum; the episternum of the mesothorax. 2. In Diptera, horizontally divided into a large mesanepimeron but with a minute meskatepimeron below. 3. In Hymenoptera Ichneumonidae, usually termed mesopleurum. mesiad see mesad mesial see mesal mesial sclerite see labellar mesial sclerite mesic a. [Gr. mesos, middle] Climate characterized by a moderate amount of water. mesinfraepisternum n. [Gr. mesos, middle; L. infra, below; Gr. epi, on; sternon, chest] (ARTHRO: Insecta) A ventral subdivision of the mesepisternum. meskatepimeron n. [Gr. mesos, middle; kata, inferior; epi, on; meros, part] (ARTHRO: Insecta) The lower division of the mesepimeron. meskatepisternum n. [Gr. mesos, middle; kata, inferior; epi, on; sternon, chest] (ARTHRO: Insecta) In Diptera, the lower area of the mesepisternum. mesobasisternum n. [Gr. mesos, middle; basis, bottom; sternon, chest] (ARTHRO: Insecta) 1. The basisternum of the mesothorax. 2. In Diptera, maybe separated from the mesofurcasternum by a secondary line of inflection. see furcasternum. mesobiota n. [Gr. mesos, middle; bios, life] Organisms in the soil ranging in size from nematodes to microannelids, microarthropods and mites; mesofauna. mesoblast n. [Gr. mesos, middle; blastos, bud] Embryonic mesoderm; the middle germ layer. mesoblastic a.

551 Online Dictionary of Invertebrate Zoology 552 mesoblastic somites Segmental divisions of embryonic mesoderm. mesobranchial lobe or area (ARTHRO: Crustacea) In Decapoda, an intermediate part of the branchial region of the carapace. mesocardiac ossicle (ARTHRO: Crustacea) In Decapoda, a triangular or oblong plate, the apex pointing forward maybe more or less truncated; forming the keystone of the anterior arch of the gastric mill. mesocephalic a. [Gr. mesos, middle; kephale, head] Having a medium size head. mesocephalic pillars (ARTHRO: Insecta) In bees, two oblique chitinous bars forming a brace between the anterior and posterior walls of the head. mesocercaria n. [Gr. mesos, middle; kerkos, tail] (PLATY: Trematoda) A juvenile stage occurring in digenetic trematodes, an unencysted stage between the cercaria and the metacercaria. mesocerebrum n. [Gr. mesos, middle; L. cerebrum, brain] (ARTHRO: Crustacea) Ganglion of antennular somite; deuterocerebrum. mesocoel n. [Gr. mesos, middle; koilos, hollow] 1. The body cavity of the second division of the deuterostome body. 2. (BRYO) Assumed to be the cavity within and at the base of the tentacles. 3. (MOLL) Second or middle division of the coelom. mesocole a. [Gr. mesos, middle; L. colere, to inhabit] Living conditions with neither too much nor too little water. mesoconch n. [Gr. mesos, middle; konche, shell] (MOLL: Bivalvia) An intermediate stage in formation of the dissoconch; separated from other stages by pronounced discontinuities. mesocuticle n. [Gr. mesos, middle; L. cutis, skin] A layer with distinctive staining properties between the exocuticle and endocuticle. mesoderm n. [Gr. mesos, middle; derma, skin] The cell layer between ectoderm and endoderm in the embryonic cells of all animals above the Cnidaria. mesodermal tube The dorsal blood vessel; heart. mesodont a. [Gr. mesos, middle; odous, tooth] (ARTHRO: Insecta) Pertaining to male Lucanidae bearing mandibles intermediate in size; amphiodont. see teleodont, priodont. meso-epinotal suture (ARTHRO: Insecta) In Formicidae, the transverse seam separating the mesonotum from the epinotum. mesoepisternum see mesepisternum mesofacial plate see face mesofauna see mesobiota mesofurca see furca mesogastric lobe/area (ARTHRO: Crustacea) In Decapoda, the medial division of the gastric region of the carapace; usually five-sided in outline with a long narrow forward projection. mesogloea n. [Gr. mesos, middle; gloios, glutinous] A thin to very thick, acellular to rather cellular, gelatinous connective tissue between the inner and the outer layers of a two-layered animal. mesohyl n. [Gr. mesos, middle; hyle, matter] (PORIF) The space lying between the pinacoderm and the choanoderm. mesolamella n. [Gr. mesos, middle; L. dim. lamina, layer] A thin mesogloeal layer between epidermis and gastrodermis in Cnidaria and Porifera. mesolecithal egg Eggs with moderate yolk content. see centrolecithal egg. mesology see ecology mesomere n. [Gr. mesos, middle; meros, part] 1. A blastomere of medium size. 2. A mesoblastic somite. 3. Central zone of coelomic pouches in an embryo. 4. (ARTHRO: Insecta) The inner divisions of the phallic lobes that unite to form the aedeagus (the intromittent organ). mesomerites n.pl. [Gr. mesos, middle; meros, part; -ites, having nature of] (ARTHRO: Diplopoda) Modified 9 pair of limbs; together with 8 pair (promerites) of limbs function-

553 Online Dictionary of Invertebrate Zoology 554 ing as pincers to pull out female vulvae. mesomeron n. [Gr. mesos, middle; meros, thigh] (ARTHRO: Insecta) In Diptera, the meron of the mesothorax behind the midcoxa below the mesepimeron. see metameron, meron. meson n. [Gr. mesos, middle] The central plane; the midline of the body; an imaginary plane dividing the body into right and left halves; the saggital plane. mesonephridium n. [Gr. mesos, middle; nephros, kidney] Nephridium of mesodermal origin. mesonotum n. [Gr. mesos, middle; notos, back] (ARTHRO: Insecta) The back or upper side of the mesothorax. mesopelagic a. [Gr. mesos, middle; pelagos, sea] Pertaining to the pelagic zone of intermediate depth of 200-1000 m; between the epipelagic and bathypelagic zones. mesopeltidium see schizopeltid mesophragma n. [Gr. mesos, middle; phragma, fence] (AR- THRO: Insecta) A chitinous piece that descends into the interior of an insect body with the postscutellum as the base. mesoplankton n. [Gr. mesos, middle; plankton, wandering] Floating life below euphotic zone; plankton organisms retained by a plankton net. mesoplax n. [Gr. mesos, middle; plax, plate] (MOLL: Bivalvia) A calcareous transverse plate straddling the two valves on their dorsal margins. see protoplax, metaplax. mesopleural bristles (ARTHRO: Insecta) In Diptera, a row of bristles on the posterior margin of the mesopleura. mesopleural fovea (ARTHRO: Insecta) In Hymenoptera Ichneumonidae, a pit or short horizontal groove on the mesopleurum, anterior to the mesopleural suture and below the speculum. mesopleural ridge (ARTHRO: Insecta) In Diptera, the pleural ridge marked externally by the mesopleural suture, between pleural apophyseal pit above the midcoxal articulation to the base of the wing. mesopleural sulcus (ARTHRO: Insecta) In Diptera, passing downwards from the wing base to the middle coxa. mesopleural suture (ARTHRO: Insecta) The external groove of the mesopleural ridge, between the base of the wing to the midcoxal articulation. mesopleuron n.; pl. -ra [Gr. mesos, middle; pleuron, side] (ARTHRO: Insecta) 1. The pleuron of the mesothorax; in winged insects, composed of basalare, subalare, mesepisternum, mesepimeron and mesotrochantin. 2. In Diptera, the dorsal part of the mesepisternum; area in front of the root of the wing between the noto- and sternopleural sutures. mesopleurosternal ridge (ARTHRO: Insecta) In Diptera, a large, posteriorly curved invagination of the mesopleurosternal suture. mesopleurosternal suture (ARTHRO: Insecta) In Diptera, the external groove between the meskatepisternum and the mesobasisternum, or mesosternum when the mesobasisternum is not distinguishable; the pleurosternal suture. mesopleurum n. [Gr. mesos, middle; pleuron, side] (ARTHRO: Insecta) In Hymenoptera Ichneumonidae, the mesepisternum. mesopostnotum n. [Gr. mesos, middle; L. post, after; Gr. notos, back] (ARTHRO: Insecta) The postnotum of the mesothorax. see metapostnotum. mesopostscutellum n. [Gr. mesos, middle; L. post, after; scutellum, small shield] (ARTHRO: Insecta) The postscutellum of an insect's mesothorax. mesopraescutum n. [Gr. mesos, middle; L. prae, before; scutum, shield] (ARTHRO: Insecta) The praescutum of the mesothorax. see prescutum. mesopsammic a. [Gr. mesos, middle; psammos, sand] Pertaining to organisms living interstitially in sand or a material in the form of rounded grains; psammous. mesopsammon n. see sabulous. mesopseudogyne see pseudogyne mesorhabdions n.pl. [Gr. mesos, middle; dim. rhabdos, rod] (NEMATA) The walls of the mesostome. see rhabdion. mesoscutellum n. [Gr. mesos, middle; L. scutellum, little