Central Principle of Biology. Evolution by Natural Selection. Aristotle BCE. Anaximander BCE. Charles Darwin

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Central Principle of Biology Evolution by Natural Selection Charles Darwin 1809-1882 Evolution of Species Fixed Species Anaximander 610-545 BCE Aristotle 384-322 BCE 1

Jean-Baptiste Lamarck 1744-1829 Theory of Evolution by Descent -Comparing fossils with current species Adaptation to Environment Inheritance of Acquired Traits Charles Lyell Geologists 1797-1875 Gradualism profound change is the result of cumulative slow but continuous processes Uniformitarianist - The same forces that gradually shaped the geological structures have not changed and act the same today as in the past. 2

Theoretical Population Growth Actual Population Growth Thomas Malthus 1766-1834 3

Darwin s Observations peccaries Armadillos Darwin s Two Ideas 1. Evolution (Descent with Modification) 2. Natural Selection (Mechanism of Evolution) 4

Natural Selection 1. Populations have the potential to increase exponentially. 2. Populations are fairly constant size. 3. Natural resources are limited. Deduction I. Only some organisms survive. There is a struggle for existence among individuals. 4. There is variation within a species which is heritable. Deduction II. Individuals with favorable variation are more likely to survive and reproduce Deduction III. Populations accumulate favorable variation over time. Evidence for Evolution (Descent with Modification 1. Fossil Evidence 2. Biogeography 3. Comparative Anatomy 4. Comparative Embryology 5. Molecular Biology 5

Fossil Record Striated Gradualism Punctuated Equilibrium Gradualism In Fossil Record 6

Biogeography Study of the geographical distribution of species Species from the same area tend to be more closely related Endemic species from islands closest relative come from near-by mainland or neighboring island Comparative Anatomy Anatomical Similarities see vertebrate forelimb Anatomical Homologies similarity due to common ancestor Vestigial Structures Remnant homologous structures 7

Analogous Structures Vestigial Structures 8

Comparative Embryology Embryonic Homologies Gill Slits Eustachian Tubes Molecular Biology Molecular Clock - Rare but steady mutation - Separate evolutionary lines with common ancestor - Number of DNA difference proportional to time since divergence from last common ancestor Molecular evidence suggests a common ancestor to all life Darwins theory doesn t explain the origin of life, simply its diversity 9

Evidence for Natural Selection 1. Artificial Selection 2. Evolving Populations 3. Adaptation is Common Artificial Selection Natural Selection unequal ability to reproduce due to interaction of phenotypes and environment. Artificial selection unequal ability to reproduce due to human intervention Artificial selection results in huge modifications over short periods of time, natural selection should be able to result in even greater modifications over vast evolutionary time periods. 10

Evolving Populations Daphne s Finches on the Galapagos Islands Seed Eaters with specialized beaks 1970 s drought changed the plant population fewer small seed, more large seed. Evolution of beak size in population of finches in result to change in seed size. Numerous Evolving Populations Trinidadian guppies Pesticide Resistance Roach resistance to Combat. 11

Adaptation is Common Skin Pigmentations Skin pigmentation (melanin ) absorbs UV light Different human populations differ in pigmentation UV light needed to make Vitamin D. Too much UV light destroys folic acid Differences in skin pigmentation represents an adaptation to the amount of UV radiation in an environments Darwin Doubted A. False Notion of Theory - Set of principles, supported by evidence that explains some aspect of the natural world. B. Use of Historical Evidence C. No Appreciation of Abundance of Evidence D. Human s special status in the universe. 12