Evolution and Darwin
Evolution : what does it mean? Evolution : change over time Biodiversity : observation of similar or different structures or functions in animals and vegetables
1831 : He begun his five- year sailing on the Beagle in the south seas (South America, Galapagos, ecc) 1859 He wrote : On the Origin of Species by Means of Natural Selection CHARLES DARWIN 1809-1882
LINNEO ANASSIMANDRO ARISTOTELE History of evolutionary thought Which theories before Darwin? LECLERC DE BUFFON LAMARK JAMES HUTTON CUVIER
ANASSIMANDRO(610-546 B.C.) homo comes from animals of different species as he cannot provide food himself until he has grown up
ARISTOTLE (384-322 B.C.) The great chain of being (Latin: scala naturae ) Hierarchical structure of all matter and life. Everything was supposed to be created by God
Creationism Religious belief that comes from the creation narrative of the Genesis Since then to nowadays nothing has changed Carl Von Linnè(1707-1778) Fixity of species
CATASTROPHISM Earth has been affected in the past by sudden, short-lived, violent events (worldwide) CUVIER (1769-1832) Many of the geological features of the earth and the past history of life could be explained by catastrophic events that caused the extinction of several species of animals (fossils)
George BUFFON (1707-1788) l'histoire naturelle, générale et particulière He was the first who tried to explain life variability
Erasmus DARWIN (1731-1802) the grandfather Animals are subject to changes whenever the environment gets modified. Descendants will receive changes as a heritage Great influence on his nephew
The Study of Geology was important to develop modern evolutionary theories
James HUTTON (1726-1797) actualism Great importance of timeline in geological changes (key role) Changes are slow Earth is thousands of years old
William SMITH (1769-1839) He discovered guide fossils
JEAN BAPTIST LAMARK (1744-1829) First scientist to propose an evolutionary theory Evolution and progression (giraffe necks) Acquired characters heritage
Trip on the south seas (1831-1836)
MALTHUS economy Charles LYELL (1797-1875) geology DARWIN Artificial selection Galapagos Investigations
MALTHUS (1766-1834) Population growth (esponential) Limit of resources (linear growth) Result: crisis Solution: to stop the growth (epidemies, death, disaster, ecc)
Charles LYELL (1797-1875) Past is the key to get the present He wrote Principles of Geology. According to him, the earth was shaped by slow-moving forces which are still in operation today. Lyell was a close and influential friend of Charles Darwin.
Artificial selection Darwin considered how able the farmers and the gardeners were in getting several minor variations among subjetcs belonging to the same species They used to select the useful features by artificial selection Close relation between the artificial selection and the natural one, although the natural one is considerably more perfect and slower.
Breeds of domestic dogs obtained by artificial selection along 10.000 years
The beagle Darwin observed that each island of Galapagos archipelago has got specific kinds of turtles and birds which are at the same time - similar and different (aspects, diet.) Darwin analyzed different kinds of animal and vegetable samples - collected during sailing - and he observed similiarites between fossils and living species coming from the same area
Galapagos observations 14 Vulcanic islands (Ecuador) In the islands there are several species both animal and vegetable due to isolation large distance from mainland wide climatic and habitat variations Endemic - typical - species While visiting the Galapagos Islands, Darwin made several remarkable discoveries.
The Glyptodonts The armadillo.are they relatives?????
Similarity between the carapace of a giant fossil and smaller shell that covered living armadillos The armadillos are native to the Americas. Fossil "related" forms giant extinct, replaced, in the modern fauna, from the smallest species in the same basic group.
the grandchildren look like their grandparents Fossils testify a relationship
Darwin uses the branching structure of a tree. The affinities of all the beings of the same class have sometimes been represented by a great tree...as buds give rise by growth to fresh buds, and these, if vigorous, branch out and overtop on all sides many a feebler branch, so by generation I believe it has been with the great Tree of Life, which fills with its dead and broken branches the crust of the earth, and covers the surface with its ever branching and beautiful ramifications." «the grandchildren look like their grandparents I think.. phylogenetic tree
Phylogenesis = evolution history THE TREE MAKES THE RELATIONSHIP BETWEEN THE DIFFERENT SPECIES High level= living species Low level = ancient species Fork = birth of new species
Birth of new species Two populations of the same species, which have been separated by a geographic barrier, differ very much in characters, so that they keep isolated in terms of reproductivity. The split populations over time may distinctly evolve different characteristics. If the geographical barriers are eventually removed, members of the two populations may be unable to mate each other successfully : the genetically isolated groups have emerged as different species
Adaptive radiation: is a consequence of allopatric speciation among island populations QUICK BIRTH OF NEW SPECIES e.g. mammalian 65 billions years ago (mass extinction ) e.g. finches of Galapagos
Finches of Galapagos On islands, Darwin met several species of finches that - although being biologically related - displayed a variety of different features. Some of them were marked by larger beaks, while some other presented more brightly coloured tails. Darwin stated that those features were adaptive, meaning that over time the finches had genetically adapted to suit environment; he named that process as natural selection.
Natural selection can lead to formation of new species Patterns of evolution Adaptive radiation: many related species evolved from a single ancestral species. Adaptive radiation most commonly occurs when a species of organism successfully invades an isolated region, where a few competing species already exist. If new habitats get available, new species will evolve. Adaptive radiation is an example of divergent evolution. Divergent evolution : two or more related species becoming more and more dissimilar (different environments) Convergent evolution: unrelated species become more and more similar in appearance as they adapt to the same kind of environment
Consequently divergent evolution Remarkable variety of forms : common ancestor. As they adapted to different environments, the appearance of the species diverged.
Convergent evolution Different species adopt similar solutions of evolution (either morphological, or physiological) in response to similar environmental conditions e.g.: fish, cetaceans, ichthyosaurs
NATURAL SELECTION ENVIRONMENT TO FIX POPULATION SIZE CONTROL Struggle for existence WHO IS MORE SUITABLE? Individuals in possession of suitable traits to struggle for local resources, will be more contributing offspring to the next generation. FITNESS = reproductive efficiency
Natural selection is the force acting in nature, to promote the survival of some individuals with certain variants of characters Adaptation is the result of the selection of variants that accidentally are more favorable
Adaptation to abyssal environment Large mouth Photophores: structures suitable for light generation ( bioluminescence)
Angraecum sesquipedale - also called Darwin's orchid - is an orchid of the Madagascar. In 1822 it is noteworthy for its long spur and Charles Darwin suggested that the flower was pollinated (by a lately discovered moth) by a proboscis whose length was then unprecedented. His prediction had gone unverified for 21 years (after his death), when the moth was discovered so his hypotesis found a confirmation.
BISTON BETULARIA and industrial revolution Originally, the vast majority of peppered moths had light colouration, which effectively camouflaged them against the light-coloured trees and lichens which they rested upon. However, because of widespread pollution during the Industrial Revolution in England, many of the lichens died out, and the trees that peppered moths rested on became blackened by soot, causing most of the light-coloured moths, or typical, to die off from predation. At the same time, the dark-coloured, or melanic, moths, carbonaria, flourished because of their ability to hide on the darkened trees.
Darwin Theory Summary 1. Species (populations of interbreeding organisms) change over time and places. 2. All organisms share common ancestors with other ones 3. Evolutionary change is gradual and slow The primary mechanism of change over time is natural selection
The Process of Natural Selection Darwin s process of natural selection is made of four components. 1. Variation. Organisms (within populations) show individual variation in appearance and behaviour. 2. Inheritance. Some traits are consistently passed on from parent to offspring. 3. High rate of population growth. Most populations have more offspring - each year compared to local resources, leading to a struggle for resources. Each generation experiences substantial mortality. 4. Differential survival and reproduction. Individuals possessing traits - well suited for struggling on local resources - will more contribute to next offspring
During the twentieth century, genetics studies were fully integrated with Darwin s mechanism, allowing us to consider the natural selection as the differential survival of genotypes, corresponding to particular phenotypes. Natural selection can only work on existing variations within a population. Such variations arise by mutation, i.e. change in some part of the genetic code for a trait.