On several new Glomeridae (Diplopoda) from Indochina

A N N A L E S H I S T O R I C O - N A T U R A L E S M U S E I NATIONAL1S H U N G A R I C I Tomus 75. Budapest, 1983 p. 107-116. On several new Glomeridae (Diplopoda) from Indochina b y S. I. G o L O V A T C H, M o s c o w Abstract Six new lndochinese species o f the millipede family Glomeridae are described: Peplomeris magna sp. n., Hyperglomeris maxima sp. n. and H. conspicua sp. n. from N o r t h V i e t n a m. Hyleoglomeris montana sp. n., H. albicollis sp. n. and H. cremea sp. n. f r o m N o r t h T h a i l a n d. Keys to all the four so far k n o w n species o f the genus Hyperglomeris S I L V. ( = Dinoglomeris S I L V., syn. n.k as well as to the nine lndochinese species o f Hyleoglomeris V E R H. are provided. W i t h 25 figures. Present k n o w l e d g e o f the G l o m e r i d a e o f I n d o c h i n a is based o n b u t a few p a p e r s ( V E R H O E F F 1915, 1920; S I L V E S T R I 1917; A T T E M S 1938, 1953), w i t h a t o t a l o f 12 species f r o m 6 genera o r subgenera r e c o g n i z e d. T h e r e f o r e t h e m o r e agreeable it was f o r m e t o have b e e n able r e c e n t l y t o a c c u m u l a t e f o r s t u d y several f r e s h c o l l e c t i n g s o f t h i s g r o u p f r o m N o r t h V i e t n a m a n d N o r t h T h a i l a n d w h i c h p r o v e d t o c o m p r i s e six i d e n t i f i a b l e species, a l l n e w t o science. T h e present p a p e r deals w i t h t h e i r d e s c r i p t i o n a n d a l l o c a t i o n a m o n g r e l a t e d f o r m s. F o r t h e o p p o r t u n i t y t o s t u d y the l o a n m a t e r i a l I a m m o s t g r a t e f u l t o D R S. Z. K A S Z A B a n d S. M A H U N K A o f the H u n g a r i a n N a t u r a l H i s t o r y M u s e u m, B u d a p e s t, a n d t o D R. H. E N G H O F F of the U n i v e r s i t e t s Z o o l o g i s k M u s e u m, K o b e n h a v n. Besides, I wish to thank particu l a r l y D R S. D. A. K R I V O L U T S K Y a n d L. B. R Y B A L O V o f the I n s t i t u t e o f E v o l u t i o n a r y A n i m a l M o r p h o l o g y a n d E c o l o g y, U S S R A c a d e m y o f Sciences, M o s c o w, f o r d o n a t i n g m e t w o nice series o f g l o m e r i d s f r o m V i e t n a m. Peplomeris magna sp.n. (Figs 1 5) L o c a l i t y : Vietnam, Prov. N i n h b i n h. Cue phuong, from pitfall traps i n forest, 1 0*, 5-18 V 1966 ( N o. 385), leg. G. T o p á i. M a t e r i a l examined: 1 specimen. H o l o t y p e : T h e above specimen, deposited i n the Hungarian N a t u r a l H i s t o r y Museum. Budapest. D i a g n o s i s Well distinguishable f r o m the only hitherto k n o w n species o f the genus, P. demangei (SILVESTRI, 1917) f r o m H a n o i, V i e t n a m, by a different colour pattern, longer a n t e n n á i j o i n t 6, larger hyposchism o f chest shield, several details o f structure i n male leg-pairs 17 to 19. D e s c r i p t i o n Length ca. 16 m m, w i d t h ca. 9.0 m m. C o l o u r yellowish, w i t h light b r o w n spots: a bit darker, subtriangular paramedian and a somewhat paler, transverse, marble lateral one f r o m each side o f terga 3 to 10 (Fig. 1). Chest shield w i t h a similar pattern, t h o u g h the lateral marble spots extending dorsad at anterior h a l f o f the tergum to contact the paramedian pair. C o l l u m w i t h o u t distinct spots, marble, a bit denser along anterior margin. Pygidium also w i t h o u t distinct spots, light marble. A n t e n n á i j o i n t s 3 t o 7 light brown, legs whitish. Antennae w i t h a l o n g and a little curved dorsad slender j o i n t 6, apically w i t h numerous small cones (Fig. 2). Black convex ocelli 7 + 1 f r o m each side. Surface smooth, shining. C o l l u m w i t h 2 usual transverse striae. Chest shield w i t h hyposchism reaching, but not exceeding caudal tergal margin, w i t h 8 (left side) or 9 (right side) very fine striae o f w h i c h 1st and 2nd (counting f r o m anterior tergal margin) cross dorsum, while 2 or 3 subsequently reaching the dorsum but becoming interrupted and vague. Pygidium w i t h a pair o f small, r o u n d, paramedian, marginal projections divided by a gentle concavity. 0". Leg-pair 17 (Fig. 3) w i t h very high outer coxal lobes, telopodite 4-jointed. Leg-pair 18 ( F i g. 4) w i t h a r o u n d syncoxite notch, telopodite 4-jointed. Pair 19 (telopods) ( F i g. 5) greatly enlarged; syncoxite median lingual lamina high, trapeziform, emarginated; syncoxite lateral horns densely setose, only a little higher than lamina, slightly clavate; prefemur w i t h a short, finger-shaped, half-hidden, inner apical process; femur w i t h a large projecting inner lobe f r o m posterior side; tibia w i t h a small p a p i l l ä r r o u n d finger distally f r o m caudal side; tarsus relatively small, acute, quickly tapering t o w a r d end.

Hyperglomeris maxima sp.n. (Figs 6 10) Locality : Vietnam, Prov. Hoa binh, Mai tiao distr., Von mai, secondary tropical forest, 4 ^ (including holotype), 1 juv., 12 XII1981, leg. D. A. Krivolutsky et L. B. Rybalov. Material examined: 5 specimens. Holotype and 2 paratypes have been deposited in the Zoological Institute of the USSR Academy of Sciences, Leningrad; one male paratype (dissected) will be placed in the Zoological Museum of the Moscow State University, Moscow; one male paratype has been sent to the Hungarian Natural History Museum, Budapest. Description Body 13-14 mm long and 6.0-7.0 mm wide, 14 mm long and 7.0 mm wide in holotype. Colour yellowish-brown, with dark marble brown spots and markings from each side of a thin irregular axial stripe which sometimes (especially in several mid-body terga) coalescesing into large (sub)triangular median spots (Fig. 6). Pygidium with a dim darker central spot. Collum marble light brown, without distinct spots. Head and antennae brown, legs greyish-yellow. Antennae like in P. magna sp. n., but apically with 4 larger cones. Black convex ocelli 7 +1 from each side. Surface smooth, shining. Collum quite usual, with 2 transverse striae. Chest shield with hyposchism reaching, but not exceeding hind tergal margin, with about a dozen of fine striae of which only 2nd entirely crosses dorsum, whereas some 6 or 7 of subsequent ones, though reaching the dorsum, do so becoming interrupted, vague. Pygidium with a pair of very small round paramedian projections at caudal margin divided by a very gentle concavity. Leg-pair 17 (Fig. 7) with quite low outer coxal lobes, telopodite 4-jointed. Pair 18 (Fig. 8) with an extremely wide and shallow syncoxite notch, telopodite 4-jointed. Pair 19 (telopods) (Figs 9-10) Figs 1-5. Pepîomeris magna sp. n,, o holotype: 1 = habitus (dorsal view), 2 = distal part of antenna (drawn not to scale), 3 = leg-pair 17, 4 = leg-pair 18, 5 = telopods (caudal view)

Figs 6-10. Hyperglomeris maxima sp. n.. Ö holotype: 6 - habitus (dorsal view, drawn not to scale), d* paratype: 7 = leg-pair 17, 8 = leg-pair 18, 9-10 = telopods (frontal and caudal view, respectively)

greatly enlarged; syncoxite median lingual lamina high, trapeziform, gently emarginated; syncoxite lateral horns setose, only a little higher than lobe, apically w i t h a tiny triangular o u t g r o w t h ; prefemur w i t h a short inner apical finger surmounted by a rudimentary seta; femur w i t h a large inner poster i o r lobe w i t h a small knob-like finger anteriorly at base, also surmounted by a rudimentary seta; tibia w i t h a good apical inner posterior swelling produced anteriad i n the f o r m o f a shelf; tarsus considerably curved, acute, gently tapering toward end. Hyperglomeris conspicua sp.n. (Figs 11 14) L o c a l i t y : Vietnam, Prov. H o a binh. M a i tiao distr.. V o n m a i, secondary tropical forest, 5 ^ (including holotype), 12 5?, 6 X I I 1981, leg. D. A. K r i v o l u t s k y et L. B. R y b a l o v. Same locality, 6??. 7 X I I 1981, leg. D. A. K r i v o l u t s k y et L. B. R y b a l o v. M a t e r i a l examined: 23 specimens. H o l o t y p e together w i t h several p a r a t y p e s have been deposited i n the Zoological Institute o f the USSR Academy o f Sciences, L e n i n g r a d ; several paratypes o f b o t h sex w i l l be placed i n the Zoological M u s e u m o f the Moscow State University, M o s c o w ; one male and one female paratypes have been sent to the Hungarian N a t u r a l H i s t o r y M u s e u m, Budapest; one female paratype w i l l go to the Universitets Zoologisk Museum, Kobenhavn. D e s c r i p t i o n Males up to 7 m m long and 3.0 m m wide (holotype), females up to 8 m m long and 3.5 m m wide. Colour yellow, w i t h dark brown to b r o w n transverse spots (Fig. 11) sometimes absent on chest shield. Pygidium w i t h o u t spots. Head and c o l l u m sometimes light b r o w n, legs pale, distal halves o f tarsi sometimes b r o w n. Antennae w i t h somewhat shorter j o i n t 6 which is ca. 2.0 times longer than wide, apically w i t h 4 larger cones. Black convex ocelli 7 + 1 from each side. Surface smooth, shining. C o l l u m w i t h usual 2 transverse striae. Chest shield w i t h a smaller hyposchism not reaching caudal tergal margin, from each side w i t h a deep sulcus (w) extending antero-dorsad f r o m the anteriormost part o f schism (Fig. 11), w i t h o u t further striae. P y g i d i u m with a pair o f very small marginal r o u n d paramedian projections divided by a very poor and gentle concavity i n males, regularly round-margined i n females. 0*. Leg-pair 17 (Fig. 12) w i t h moderately high outer coxal lobes, telopodite 4-jointed. Pair 18 (Fig. 13) w i t h a broadly r o u n d syncoxite notch, telopodite 4-jointed. Pair 19 (telopods) (Fig. 14) greatly enlarged; syncoxite median lingual lamina almost regularly semi-circular, rather h i g h ; syncoxite lateral horns a little higher than lamina, setose, subtriangular at the apex, disto-laterally with a good spine; prefemur w i t h a short, stout, setiferous disto-mesal process (finger); femur w i t h very long and slender posterior finger-shaped inner process w i t h setiferous anterior k n o b at base; tibia w i t h a very conspicuous, serrate, l o n g (ca. 2/3 as long as tarsus), posterior apical process, w i t h a good membraneous inner hook, and a small subterminal setiferous k n o b anteriorly; tarsus considerably curved, acute, gently tapering t o w a r d end. R e m a r k s Hyperglomeris dirupta E v e r since S I L V E S T R I ' S ( 1 9 1 7 ) e r e c t i o n o f t w o m o n o t y p i c a l genera, a n d Dinoglomeris, t o c o m p r i s e the n e w species H. lamellosa SILV. and D. S I L V. f r o m N o r t h V i e t n a m, b o t h the genera have b e e n c o n s i d e r e d g o o d ( M A U R I È S 1 9 7 1, HOFFMAN 1 9 7 9 ). H o w e v e r, the o n l y d i s t i g u i s h i n g c h a r a c t e r, as SILVESTRI ( 1 9 1 7 ) noted by h i m s e l f, was the presence o r absence at base o f the t e l o p o d i n n e r f e m o r a l process o f a s m a l l a n t e r i o r finger t o s u p p o r t a seta: Dinoglomeris w i t h such finger, Hyperglomeris without it. T h i s d i s t i n c t i o n, suspicious e n o u g h i n itself, i n the l i g h t o f t h e n e w discoveries s h o u l d be a b a n d o n e s n o w a l t o g e t h e r, f o r i n H. maxima l y r e m i n d s the p o o r k n o b i n H. conspicua glomeris sp. n. the finger is a l r e a d y q u i t e s m a l l a n d r e a d i sp. n. T o f o r m a l i z e t h e issue, I c o n s i d e r S I L V. a subjective senior s y n o n y m o f Dinoglomeris Hyper- S I L V., a l l the f o u r so far k n o w n species o f this genus b e i n g easily d i s t i n g u i s h a b l e b y the k e y b e l o w. Perhaps i t is neccessary t o r e m i n d that Hyperglomeris m o s t closely r e l a t e d t o the l a r g e genus Hyleoglomeris i n its present b r o a d e r concept is also o f O r i e n t a l o r i g i n. H o w e v e r, the l a t t e r possesses t h e p o s t e r i o r t e l o p o d f e m o r a l process m u c h l a r g e r a n d w i t h an a p i c a l m e m b r a n e o u s sack, a n d t h e t i b i a o f the t e l o p o d s w i t h o u t g o o d p o s t e r i o r processes o r swellings.

0-3 M M Figs 11-14. Hyperglomeris conspicua sp. n., o* paratype: 11 = habitus (dorsal view, drawn not to scale), 12 = leg-pair 17, 13 = leg-pair 18, 14 = leg-pair 19 (frontal view). Fig. 15. Hyleoglomeris albicollis sp. n., <J holotype, habitus (dorsal view, drawn not to scale). Figs 16-19. Hyleoglomeris mont ana sp. n., d* paratype: 16 = habitus (dorsal view, drawn not to scale). 17 = leg-pair 17, 18 = leg-pair 18, 19 = telopods (frontal view)

A KEY TO THE KNOWN SPECIES OF HYPERGLOMERIS SILVESTRI 1 (2) Pygidium entirely, chest shield almost or entirely depigmented ; latter without fine striae, but with a good sulcus antero-dorsad of the schism; syncoxite 18 notch broadly round, telopod syncoxite lateral horns with an outer distal spine each; telepod caudal tibial process very long, slender, serrate, reaching ca. 2/3 tarsal length H. conspicua sp. n. 2 (1) Both pygidium and chest shield with dark pigment; chest shield with a number of fine striae, but without deep sulci; notch of male syncoxite 18 not round; lateral horns of telopod syncoxite without outer spines; telepod caudal tibial process or swelling far from that long 3 3 (4) A thin dark axial stripe upon lighter background of dorsum, chest hyposchism larger, reaching caudal tergal margin; lateral lobes of male coxite 17 quite low; male syncoxite 18 notch extremely broadly triangular; median lingual lamina of telopod syncoxite trapeziform, gently, but evidently emarginated H. maxima sp. n. 4 (3) No dark axial stripe along dorsum, dorsal background black to red-yellow; chest hyposchism smaller, not reaching tergal caudal margin; lateral lobes of male coxite 17 higher; notch of male syncoxite 18 ogive; telopod syncoxite median lamina not emarginated 5 5 (6) Chest hyposchism only a little not reaching tergal caudal margin; telopod syncoxite median lamina trapeziform, with evidently truncate ventral margin H. dirupta (SILV.) comb. n. 6 (5) Chest hyposchism almost wanting; telopod syncoxite median lamina regularly jounded, semi-circular H. lamellosa SILV. Hyleoglomeris montana sp.n. (Figs 16-19) Locality : Thailand. Prov. Chieng Mai, Doi Inthanon, summit, 2500 m, 2 dv (including holotype), 3 $9, 1 juv. 7X 1981, leg. Zool. Museum Kobenhavn. Material examined: 6 specimens. Holotype, 1 female and the juvenile paratypes are deposited in the Universitets Zoologisk Museum, Kobenhavn ; 1 female paratype even is in the Hungarian Natural History Mseum Budapest, and Coll. Golovatch, Moscow. Description Males 7.0 mm long and ca. 4.0 mm wide (holotype), females 9.5-12.0 mm long and 4.6-4.8 mm wide. Colour black-brown, with yellow-whitish thin axial stripe, pairs of large marble brown lateral spots, sometimes (in adult paratypes) with background irregularly interrupted by small light markings (Fig. 16). Collum dark brown, with a large lighter marble central spot. Pygidium with a caudal widening of the light axial stripe. Head and antennae brown, latter ones especially dark distally. Legs whitish. Antennae usual, apically with 4 larger cones, joint 6 ca. 2.0 times longer than wide. Black convex ocelli 6 + 1 from each side. Surface smooth, shining. Collum with 2 usual transverse striae. Chest shield with a large hyposchism reaching, but not protruding beyond hind tergal margin, with 10-13 fine striae of which up to 7 lying above schism and only four entirely crossing dorsum. Pygidium delicately sinuate mediocaudally in males, regularly margined in females. d*. Leg-pair 17 (Fig. 17) with very high outer coxal lobes, telopodite 4-jointed. Leg-pair 18 (Fig. 18) with an ogive syncoxite notch, telopodite 4-jointed. Telopods greatly enlarged (Fig. 19); syncoxite median lingual lamina subtrapeziform, with rounded ventral angles; syncoxite lateral horns very high, slender, apically with a long hyaline bifid seta each, densely setose; prefemur with a long inner apical finger crowned by a long flagelloid ; femur anteriorly with a similar apical inner digitoid process, but surmounted by a shorter flagelloid, posteriorly with a very large lobe apically membraneous and slightly curved; tibia anteriorly with a long distal inner seta, posteriorly with a large membraneous hook-like inner process; tarsus quite slightly curved, acute, very gently tapering toward end.

Hyleoglomeris albicollis sp.n. (Figs 15, 20-22) Locality : Thailand, Prov. Chieng Mai. Doi Inthanon N. P., Mae Chaem, 1700 m, road, 1 <f, 15 X 1981, leg. Zool. Museum Kobenhavn. Material examined: 1 specimen. Holotype is deposited in the Universitets Zoologisk Museum. Kobenhavn. Description Body 11 mm long and 5.0 mm wide. Colour black, with yellowish spots: a small median at hind margin and a pair of larger lateral marble ones on each tergum but collum and pygidium. Chest shield with yellowish postero-lateral corners, collum entirely yellow, without darker pigment, pygidium with a large median subtriangular spot at hind margin. Head and antennae dark brown, legs whitish, only distal halves of tarsi a bit brownish (Fig. 15). Antenna! joint 6 long, ca. 2.2 times longer than wide. Black convex ocelli 6 +1 from each side. Surface smooth, shining. Collum with 2 usual transverse striae. Chest shield with a large hyposchism reaching, but not projecting beyond hind tergal margin, with 8 fine striae of which (counting from anterior tergal margin) 1st. 4th and 7th entirely crossing dorsum. Pygidium regularly margined, without peculiarities. Figs 20-22. Hyleoglomeris albicollis sp. n., o* holotype: 20 = leg-pair 17, 21 = leg-pair 18, 22 = telopods (frontal view) 8 Természettudományi Múzeum Évkönyve 1983

3. Leg-pair 17 (Fig. 20) w i t h rather high outer coxal lobes, telopodite 4-jointed. Pair 18 ( F i g. 21) with a r o u n d syncoxite notch, telopodite 4-jointed. Leg-pair 19 (telopods) (Fig. 22) very massive; median lingual lamina o f syncoxite subtrapeziform, emarginated. w i t h rounded ventro-lateral cor ners; syncoxite lateral horns very high, slender, setose, apically w i t h a very short setoid i n front o f a tiny r o u n d l o b e ; prefemur w i t h a very long inner apical finger crowned w i t h a long flagelloid; femur w i t h a similar smaller antero-mesal finger also crowned by a long flagelloid, posteriorly w i t h a large inner lobe surmounted by a membraneous sack curved f o r w a r d ; tibia w i t h a long antero-mesal seta, posteriorly w i t h an indistinct membraneous process; tarsus considerably curved, gently tapering toward end. Hyleoglomeris cremea sp.n. (Figs 23-25) 1 L o c a l i t y : N o r t h T h a i l a n d, Chiang D a o. 1800 m, 3 o d (including holotype), 5 $$, 1958-59, leg. B. Degerbol. M a t e r i a l examined : 8 specimens. H o l o t y p e, together with 1 male and 4 female p a r a t y p e s are deposited i n the Universitets Zoologisk M u s e u m, K o b e n h a v n ; 1 female paratype is i n the Hungarian N a t u r a l History M u s e u m. Budapest; 1 male paratype is retained i n C o l l. Golovatch, M o s c o w. D e s c r i p t i o n B o d y 8-12 m m long and 3.6-5.5 m m wide i n males (10 and 4.2 m m i n holotype). 10-15 m m long and 4.5-7.0 m m i n females. C o l o u r y e l l o w i s h - b r o w n ; c o l l u m, chest shield and almost all pygidium (except for the antero-lateral corners) yellowish, other terga almost all dar ker, brownish, w i t h a more o r less wide axial light stripe. Head and antennae yellowish-brown t o b r o w n, legs light yellowish-brown. Antenna! j o i n t 6 ca. 2.0 times as l o n g as wide. B l a c k - b r o w n convex ocelli 7 + 1 from each side. Surface smooth, shining. C o l l u m w i t h 2 usual transverse striae. Chest shield w i t h a rather large hyposchism almost reaching h i n d tergal margin, w i t h 9-11 fine striae o f which 6 crossing the dorsum (always 1st and 2nd, but never 3rd). M a l e pygidium very slightly sinuate medio-caudally, w i t h o u t modifications i n females. d*. Leg-pair 17 (Fig. 23) w i t h rather high outer coxal lobes, telopodite 4-jointed. Pair 18 ( F i g. 24) w i t h syncoxite notch ogive, telopodite 4-jointed. Telopods (Fig. 25) greatly enlarged; syncoxite medial lamina prominent, subovoid ; syncoxite lateral h o r n high, slender, setose, apically w i t h a good setoid ; prefemur w i t h usual l o n g flagelliferous finger ; femur w i t h a similar, but stout antero-mesal finger crowned by a flagelloid and posteriorly w i t h a big lobe apically bearing a membraneous sack curved f o r w a r d ; tibia w i t h a l o n g antero-apical inner seta, posteriorly w i t h an indistinct apical inner membraneous process; tarsus rather gently curved, very moderately tapering t o w a r d end. R e m a r k s Six species o f Hyleoglomeris have h i t h e r t o b e e n k n o w n f r o m I n d o c h i n a : H. tricmgulifera A T T., H. robusta A T T., H. maior S I L V. a n d H. siamensis S I L V. T h e l a t t e r is t h e o n l y o n e k n o w n f r o m c o n t i n e n t a l T h a i l a n d. T h e d i s c o v e r y o f the t h r e e n e w Hyleoglomeris A T T., H. pulchra A T T., H. electa i n N o r t h T h a i l a n d n o t o n l y enriches t h e l o c a l f a u n a, b u t also p r o v i d e s t h e o p p o r t u n i t y t o a r r a n g e a l l the l n d o c h i n e s e species o f t h i s g e n u s i n a t e n t a t i v e k e y. A s several o f these f o r m s h a v e b e e n described b u t b y females, t h e k e y b e l o w c h i e f l y deals w i t h c o l o u r p a t t e r n s. I t s h o u l d be r e m e m b e r e d, h o w e v e r, t h a t the c o l o u r p a t t e r n s described a b o v e a n d h e r e i n a f t e r d i s r e g a r d t h e u s u a l l y l i g h t l a t e r a l a n d posterior tergal margins. A K E Y T O T H E I N D O C H I N E S E S P E C I E S O F HYLEOGLOMERIS 1(2) VERHOEFF C o l l u m entirely depigmented (yellow), w e l l contrasting w i t h m u c h darker (black) pigmentation o f the other terga H. 2 a l b i c o l l i s sp. n. (1) C o l l u m at least p a r t l y w i t h d a r k p i g m e n t, l i g h t b r o w n t o b l a c k, u s u a l l y m a r b l e 3 3 (10) C h e s t s h i e l d a n d p y g i d i u m e n t i r e l y o r m a i n l y l i g h t, w h i t i s h t o b r o w n ; spots, w h e n present, darker t h a n b a c k g r o u n d 4 ( 7 ) C h e s t shield a n d p y g i d i u m w i t h d i s t i n c t spots o r m a r k i n g s 4 5

5 (6) Chest sh,eld yellowish-brown, pygidium whitish-yellow, both with darker snots stiu well contrasting with black background of terga 3 to 11 prov dedi with3 r 1 3 r of large yellowish spots; larger: 6.5 mm wide W S H. triangulifera ATT. Figs 23-25. Hyleoglomeris cremea sp. n., rf paratype: 23 - leg-pair 17, 24 = leg-pair 18 25 = telopods (frontal view)

6 (5) Chest shield and pygidium cream-brown, each with a pair of darker spots, not contrasting with cream-brown terga 3 to 11; smaller: 5 mm wide H. siamensis SILV. 7 (4) Chest shield and pygidium without distinct spots 8 8 (9) Collum, chest shield and pygidium only slightly lighter than and not contrasting with light brown or brown terga 3 to 11 which have a more or less wide, regular, lighter axial stripe; larger: 4.5 to 7.0 mm wide H. cremea sp. n. 9 (8) Collum darker, chest shield and pygidium orange-yellowish, well contrasting with black background of terga 3 to 11 each provided with a large, median, yellowish spot; smaller: 3 mm wide H. pulchra ATT. 10 (3) Chest shield and pygidium mainly dark, dark brown to black; spots, when present, lighter than background 11 11 (12) Chest shield to segment 4 completely dark, black, without lighter spots or markings; larger: 7-9 mm wide H. maior ATT. 12(11) Chest shield to segment 4 also with several more or less distinct lighter spots or markings 13 13 (14) A thin, more or less regular, whitish axial stripe from chest shield to pygidium H. montana sp. n. 14 (13) From chest shield to pygidium no axial stripe, but separate small spots at best 15 15 (16) From chest shield to pygidium 3 rows of brown-yellowish spots: one median and a pair of large marble lateral ones; pygidium with a large median spot, but without lateral ones H. electa SILV. 16 (15) From chest shield to pygidium no axial row of spots; pygidium with a large median and a pair of marble brown-yellowish lateral spots H. robustus ATT. References ATTEMS, C. (1938): Die von Dr. C. Dawydoff in Französisch Indochina gesammelten Myriopoden. Mém. Mus. natn. Hist. nat. Paris, N. S., 6 (2): 187-353. ATTEMS, C. (1953): Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938-1939). Ibid., 5A (3): 133-230. HOFFMAN, R. L. (1979): Classification of the Diplopoda. Mus. Hist. nat. Geneve, p. 1-237. MAURIÈS, J.-P. (1971): Diplopodes épigés et cavernicoles des Pyrénées espagnoles et des Monts Cantabriques. VII. Glomérides. Essai de classification de Glomeroidea. Bull. Soc. Hist.nat. Toulouse, 107 (3-4): 423-436. SILVESTRI, F. (1917): Contributions to a knowledge of the Oriental Diplopoda Oniscomorpha. Ree. Ind. Mus. Calcutta, 13: 103-151. VERHOEFF, K. W. (1915): Zur Kenntnis der Plesiocerata. (Über Diplopoden, 82. Aufsatz). Zool. Anz., 46: 16-29. 43-62. VERHOEFF, K.W. (1920): Chilognathen-Studien (91. Diplopoden-Aufsatz). Arch. Naturg., 86A (12): 23-80. Author's address: S. I. GOLOVATCH Institute of Evolutionary Animal Morphology and Ecology, USSR Academy of Sciences Leninsky prospekt 33 Moscow V 71 USSR