larger scale human cooperation

Size: px
Start display at page:

Download "larger scale human cooperation"

Transcription

1 largerscalehumancooperation anddonot culturalevolutionaryprocesses

2 normpsychology Speciesdifferences: Scaleandintensitydifferences: Domaindifferences: Rapidintensification: Noncooperativeandmaladaptivesociality:

3 mismatchhypotheses 1 Prior work has referred to these mismatch hypotheses descriptively (not snarkily) as big mistake hypotheses (Fehr and Henrich 2003; Boyd and Richerson 2002; Henrich and Henrich 2007). We adopt mismatch here in deference to terminology used by proponents of this point of view.

4 thesethreefeatures these mechanismsalone ancientsocialpsychology tribalpsychology bcr givensomesimplifyingassumptions relatedness r=.5r=.25,r=.125

5 r thatcompetitionisglobal closerkinr less almostnever all more

6 =0 is approachingzero goestozeroevenfaster. broad largerscalecooperation, 2 Large-scale cooperation does not mean lots of cooperation in different dyads. Large-scale cooperation involves large groups cooperating, in for example, joint projects (armies, recycling, paying taxes, giving blood, etc.), not just the concurrent existence of many cooperating dyads.

7 reputation

8 cooperative irrelevant r r 3 Note that relatedness measures the genetic similarity between two individuals relative to the population average. Negative relatedness values imply that two individuals are less similar to each other than they are, on average, to a randomly selected member of their species (Grafen 1985).

9 4 As far as evolution is concerned, sparing someone when you could profit by exploiting them is just as much a cooperative act as providing them with costly aid, thus these are all examples of cooperation-relevant interactions with strangers.

10 Table1 Wordsfor secret infoursmallscalesocieties 6 Group Hadza Ju(!Kung) Tsimane Machiguenga 5 Much work in evolutionary psychology is based on the assumption that extra-pair copulations (adultery) and copulations with different males in relatively short periods of time were not only possible, but a serious factor in the lives of our foraging ancestors (Buss 2007). If true, secrets and anonymity (like about paternity) were indeed possible, and individuals could avoid reputational damage and revenge while engaging in clandestine activities. We think this fits well with ethnography, but not with the assumptions of some mismatch hypotheses. That is, mismatch assumptions about ancestral environments conflicts with much work by those studying mating psychology. 6 Thanks to Frank Marlowe, Glenn Sheppard, Mike Gurven, and Polly Weissner for this information, and to Rob Boyd for suggesting this approach.

11 learningstrategies culturalvariant content context Modelbiasedtransmission

12 Conformistbiasedtransmission context content ceterisparibus de novo 7 A lively debate persists on the evolutionary foundations of conformist transmission (e.g., Nakahashi 2007; Wakano and Aoki 2007; Guzman, Rodriguez-Sickert, and Rowthorn 2007).

13 immediately immediately socialnorms multiple stableequilibria Box1: Gametheoretic evolutionarymodelling tableequilibria equilibriumselection,

14 equilibriumselection culturalgroupselection can 8 Recent theoretical work by Lehman et. al. (2008; 2008; 2007) has sought to challenge some of these theoretical conclusions. Their models, however, make quite different assumptions (which are buried deep in the mathematics) about the (1) strength of cultural learning relative to the forces of mixing among group and (2) presence of multiple stable equilibria. Their efforts are not critiques but rather alternative hypotheses based on empirically difficult-tosupport assumptions about human cultural transmission and social norms (Boyd, Richerson, and Henrich forthcoming). 9 Note that this does not imply that groups are internally homogenous; equilibria can be polymorphic: stable mixes of different behaviours, where an increase in the frequency of any behavioural strategy makes the alternatives more advantageous. For example, consider a market in which everyone is selling peanut better. In such an environment, a jelly-seller will do well until there are too many jelly-sellers, and not enough peanut butter sellers.

15

16 withinasingleisolatedgroup

17 normpsychology: stable feelsright not feel rationally highly 10 Is this strong reciprocity? The term strong reciprocity emerged in the late 1990 s as a label for a set of social motivations that were implied by a large body of empirical findings mostly emerging from experimental economics laboratories. Some efforts were made to develop evolutionary theories to explain the origins of these motivations (Gintis 2000; Fehr, Fischbacher, and Gächter 2002). It s important to differentiate the empirical finding (and their implied proximate motivations) from efforts at evolutionary explanations (which are not strong reciprocity ). Our approach begins with culture-gene coevolution theory and derives predictions. These predictions regarding normspsychology can account for the patterns associated with strong reciprocity, as empirical phenomena, but also go well beyond them by seating such patterns in the context of both cultural and genetic evolution.

18 Humanshouldpossesscognitivemechanismsthatscaffoldtheacquisitionofnorms. Becauseculturalgroupselectionhasselectivelyfavouredmostlyprosocialnormsovertensof thousandsofyears,ourcognitiveintuitionsshouldincludeprosocialdefaultcontentandrelated learningbiases. Humansshouldreadilyacquirecostlysocialnormsviaculturallearning. Bothadheringtosocialnormsandpunishingnormviolatorsshouldbeintrinsicallyorinternally rewarding.

19 Onceinternalised,normadherenceshouldbeautomaticsuchthatnormviolationsrequireacognitive override. Speciesdifferences: mismatchhypotheses Scaleandintensitydifferences: Domaindifferences:

20 Rapidintensification: Noncooperativeandmaladaptivesociality:

21 BiologicalTheory Proc.Natl. Acad.Sci. TheEvolutionofCooperation encere AmericanPoliticalSci view Science Neuron ProceedingsoftheNationalAcademyof SciencesoftheUnitedStatesofAmerica AmericanAntiquity ChildDevelopment Cognition:InternationalJournalof CognitiveScience Science Animal Behaviour Science ProceedingsoftheNationalAcademyofSciencesoftheUnitedStatesofAmerica JournalofTheoreticalBiology JournalofTheoreticalBiology BehavioralEcologyandSociobiology Journalof TheoreticalBiology

22 Ethology&Sociobiology EvolutionandCultur e CultureandtheEvolutionaryProcess JournalofTheoreticalBiology ChildDevelopment ChildDevelopment AnalyseandKritik narypsychology:thenewscienceofthemind chology:thenewscienceofthemind Evolutio EvolutionaryPsy BehaviorGameTheory:ExperimentsinStrategicInteraction Journalof SocioEconomcis alof Journ EconomicPerspectives Science Nature Processes& Group AmericanAnthropologist IntergroupRelations Thetrutha ewofparentallove boutcinderalla:adarwinianvi TheGoddelusion Science ProceedingsoftheRoyalSocietyofLondon SeriesBBiologicalSciences Science Guns,Germs,andSteel:TheFatesofHumanSocieties Coevolution:genes,culture,andhumandiversity Sicksocieties:challengingthemythofprimitiveharmony Science

23 EvolutionandHumanBehavior TrendsinCognitiveSciences GeneticandCulturalEvolution ofcooperation HumanNature JournalofCognitionandCulture JournalofAnthropologicalArchaeology JournalofEvolutio narybiology JournalofTheoreticalBiology JournalofTh l eoretica Biology Oxfordsurveysinevolutionarybiology Neuron Science PopulationAndDevelopmentReview Evolution andhumanbehavior TheoreticalPopulationBiology JournalofTheoreticalBiology DevelopmentalScience Nature AGeneralTheoryofEquilibriumSelectioninGames RationalityandSociety JournalofPersonalityandSocialPsychology JournalofEconomicBehavior&Organization Science

24 ExplainingCultureScientifically EvolutionandHumanBehavior GeneticandCultural EvolutionofCooperation EvolutionandHumanBehavior JournalofTheoreticalBiology JournalofCognitionandCulture EvolutionandHuman Behavior ProceedingsoftheRoyalSocietyB Biological Sciences EvolutionaryAnthropology EvolutionandHuman Behavior BehavioralandBrainSciences CurrentAnthropology Science OxfordHandbookofEvolutionaryPsychology WhyHumansCooperate:ACulturalandEvolutionaryExplanation Science PsychologicalScience

25 JournalofTheoreticalBiology ChildDevelopment Science ProceedingsoftheRoyalSocietyBBiological Sciences FamiliesoftheForest:MatsigenkaIndiansofthePeruvianAmazon Nature Journalof Genetics WarBeforeCivilization TheNuerConquest Theoretical PopulationBiology Proceedings cademyofsciencesoftheunitedstatesofamerica ofthenationala GoodCompanyandViolence:SorceryandSocialActioninaLowlandNew GuineaSociety JournalofAnthropologicalArchaeology ProceedingsoftheRoyalSocietyofLondonSeriesBBiologicalSciences NatureReviewsGenetics ProceedingsoftheNationalAcademyofSciences TheoreticalPopulationBiology AmericanNaturalist AmericanNaturalist Proceedings:BiologicalSciences Superfreakonomics:globalcooling,patriotic prostitutesandwhysuicidebombersshouldbuylifeinsurance American SociologicalReview

26 urrentanthropology Cross CulturalResearch C MathematicalModelsofSocialEvolution:AGuideforthe Perplexed CurrentAnthropology EvolutionandHumanBehavior CurrentAnthropology EvolutionandHumanBehavior BritishJournalofPsychology JournalofPersonalityandSocial Psychology TheoreticalPopulationBiology FoliaPrimatologica Science DevelopmentalScience EvolutionandHumanBehavior JournalofTheoreticalBiology Nature Nature OrganizationalBehaviorandHumanDecisionProcesses EvolutionandHumanBehavior DevelopmentalPsychology American Antiquity Indoctrinability, IdeologyandWarfare

27 International JournalofNeuropsychopharmacology ValuesandKnowledge JournalofPersonalityandSocialPsychology EuropeanJournalofSocial Psychology AmericanAnthropologist Science Nature Nature CurrentAnthropology JournalofAnthropologicalArchaeology Games andeconomicbehavior PsychologicalScience Evolution;internationaljournalof organicevolution Ethology&Sociobiology TheAdaptedMind:Evolutionary PsychologyandtheGenerationofCulture Science TheQuarterlyReviewofBiology SocialCognition:KeyReadings ProceedingsoftheNationalAcademyof SciencesoftheUnitedStatesofAmerica AnimalBehaviour

28 TheoreticalPopulationBiology Americanjournalofprimatology Current Biology InternationalJournalofPrimatology American Antiquity Individualstrategyandsocialstructure:anevolutionarytheoryof institutions

ABSTRACT: Dissolving the evolutionary puzzle of human cooperation.

ABSTRACT: Dissolving the evolutionary puzzle of human cooperation. ABSTRACT: Dissolving the evolutionary puzzle of human cooperation. Researchers of human behaviour often approach cooperation as an evolutionary puzzle, viewing it as analogous or even equivalent to the

More information

The Coevolution of Warfare, Punishment, and Culture in Homo sapiens

The Coevolution of Warfare, Punishment, and Culture in Homo sapiens The Coevolution of Warfare, Punishment, and Culture in Homo sapiens Richard McElreath Department of Anthropology and Center for Population Biology, University of California Davis, Davis, CA 95616 This

More information

Anthropology 562: Introduction to Evolutionary Anthropology

Anthropology 562: Introduction to Evolutionary Anthropology Anthropology 562: Introduction to Evolutionary Anthropology Instructor: Ed Hagen Office: VMMC 102E Phone: 360-546-9257 Email: edhagen@vancouver.wsu.edu Office hours: Anytime Course website: Lecture day

More information

On the evolution of reciprocal cooperation

On the evolution of reciprocal cooperation On the evolution of reciprocal cooperation Jean-Baptiste André Ecologie & Evolution - CNRS - ENS, Paris, France Biodiversity and Environment: Viability and Dynamic Games Perspectives Montreal, November

More information

Divided We Stand: The Evolution of Altruism. Darwin 101. The Paradox of Altruism. Altruism: Costly behavior that helps others

Divided We Stand: The Evolution of Altruism. Darwin 101. The Paradox of Altruism. Altruism: Costly behavior that helps others Predator inspection Divided We Stand: The Evolution of Altruism Karthik Panchanathan buddha@ucla.edu Territorial defense Hunting Grooming Altruism: Costly behavior that helps others Food sharing Warfare

More information

Darwinian Evolution of Cooperation via Punishment in the Public Goods Game

Darwinian Evolution of Cooperation via Punishment in the Public Goods Game Darwinian Evolution of Cooperation via Punishment in the Public Goods Game Arend Hintze, Christoph Adami Keck Graduate Institute, 535 Watson Dr., Claremont CA 97 adami@kgi.edu Abstract The evolution of

More information

Play Locally, Learn Globally: The Structural Basis of Cooperation

Play Locally, Learn Globally: The Structural Basis of Cooperation Play Locally, Learn Globally: The Structural Basis of Cooperation Jung-Kyoo Choi 1 University of Massachusetts, Amherst, MA 01003 November 2002 1 I would like to thank the Santa Fe Institute for the Computational

More information

MATCHING STRUCTURE AND THE EVOLUTION OF COOPERATION IN THE PRISONER S DILEMMA

MATCHING STRUCTURE AND THE EVOLUTION OF COOPERATION IN THE PRISONER S DILEMMA MATCHING STRUCTURE AN THE EVOLUTION OF COOPERATION IN THE PRISONER S ILEMMA Noureddine Bouhmala 1 and Jon Reiersen 2 1 epartment of Technology, Vestfold University College, Norway noureddine.bouhmala@hive.no

More information

Joe Yuichiro Wakano. Research Accomplishments

Joe Yuichiro Wakano. Research Accomplishments Joe Yuichiro Wakano Research Accomplishments (May 22nd, 2013) Current Position Associate Professor School of Interdisciplinary Mathematical Sciences Meiji University Research Interests Mathematical Biology

More information

9.916 Ingroups and Outgroups

9.916 Ingroups and Outgroups 9.916 Ingroups and Outgroups Today s Lecture One mind thinking about another: - stable - universal Social relationships: - dynamic - co-dependent - context-bound Social relationships depend on a continuous

More information

Cooperation Achieved by Migration and Evolution in a Multilevel Selection Context

Cooperation Achieved by Migration and Evolution in a Multilevel Selection Context Proceedings of the 27 IEEE Symposium on Artificial Life (CI-ALife 27) Cooperation Achieved by Migration and Evolution in a Multilevel Selection Context Genki Ichinose Graduate School of Information Science

More information

The co-evolution of culturally inherited altruistic helping and cultural transmission under random group formation

The co-evolution of culturally inherited altruistic helping and cultural transmission under random group formation Theoretical Population Biology 73 (2008) 506 516 Contents lists available at ScienceDirect Theoretical Population Biology journal homepage: www.elsevier.com/locate/tpb The co-evolution of culturally inherited

More information

Exercise 3 Exploring Fitness and Population Change under Selection

Exercise 3 Exploring Fitness and Population Change under Selection Exercise 3 Exploring Fitness and Population Change under Selection Avidians descended from ancestors with different adaptations are competing in a selective environment. Can we predict how natural selection

More information

EVOLUTIONARY GAMES WITH GROUP SELECTION

EVOLUTIONARY GAMES WITH GROUP SELECTION EVOLUTIONARY GAMES WITH GROUP SELECTION Martin Kaae Jensen Alexandros Rigos Department of Economics University of Leicester Controversies in Game Theory: Homo Oeconomicus vs. Homo Socialis ETH Zurich 12/09/2014

More information

At Penn Museum Conference: Summary of Richerson

At Penn Museum Conference: Summary of Richerson At Penn Museum Conference: Summary of Richerson Steven O. Kimbrough kimbrough@wharton.upenn.edu 215-898-5133 29 September 2007 File: PennMuseum-foils-sok.tex/pdf. Two papers Conference paper: Rethinking

More information

Living in groups 1. What are three costs and three benefits of living in groups?

Living in groups 1. What are three costs and three benefits of living in groups? Study questions Living in groups 1. What are three costs and three benefits of living in groups? 2. What is the dilution effect? What is a key assumption of the dilution effect hypothesis? What is a biological

More information

3/30/2012. Two Contrasting but Complementary Evolutionary Perspectives on Human Behavior:

3/30/2012. Two Contrasting but Complementary Evolutionary Perspectives on Human Behavior: Two Contrasting but Complementary Perspectives on Human Behavior: Psychology (EP) derived from a synthesis of biology and psychology Human Behavioral Ecology (HEB) derived from a synthesis of biology and

More information

Journal of Economic Behavior & Organization

Journal of Economic Behavior & Organization Journal of Economic Behavior & Organization 72 (2009) 1 20 Contents lists available at ScienceDirect Journal of Economic Behavior & Organization journal homepage: www.elsevier.com/locate/jebo A group selection

More information

Anthro 101: Human Biological Evolution. Lecture 11: Cooperation, Intelligence, Communication, Culture, & Human Behavior. Prof.

Anthro 101: Human Biological Evolution. Lecture 11: Cooperation, Intelligence, Communication, Culture, & Human Behavior. Prof. Anthro 101: Human Biological Evolution Lecture 11: Cooperation, Intelligence, Communication, Culture, & Human Behavior Prof. Kenneth Feldmeier Reminders Fossil lecture for homework Quiz about material

More information

Evolution and Human Behavior

Evolution and Human Behavior Evolution and Human Behavior 34 (2013) 342 349 Contents lists available at ScienceDirect Evolution and Human Behavior journal homepage: www.ehbonline.org Original Article Cultural evolution of cooperation:

More information

Evolution and Epigenetics. Seminar: Social, Cognitive and Affective Neuroscience Speaker: Wolf-R. Brockhaus

Evolution and Epigenetics. Seminar: Social, Cognitive and Affective Neuroscience Speaker: Wolf-R. Brockhaus Evolution and Epigenetics Seminar: Social, Cognitive and Affective Neuroscience Speaker: Wolf-R. Brockhaus 1. History of evolutionary theory The history of evolutionary theory ~ 1800: Lamarck 1859: Darwin's

More information

Game Theory, Population Dynamics, Social Aggregation. Daniele Vilone (CSDC - Firenze) Namur

Game Theory, Population Dynamics, Social Aggregation. Daniele Vilone (CSDC - Firenze) Namur Game Theory, Population Dynamics, Social Aggregation Daniele Vilone (CSDC - Firenze) Namur - 18.12.2008 Summary Introduction ( GT ) General concepts of Game Theory Game Theory and Social Dynamics Application:

More information

Evolution of motivations and behavioral responses! Integrating the proximate and ultimate causes of behavior!

Evolution of motivations and behavioral responses! Integrating the proximate and ultimate causes of behavior! Evolution of motivations and behavioral responses! Integrating the proximate and ultimate causes of behavior! Erol Akçay! National Institute for Mathematical! and Biological Synthesis (NIMBioS)! University

More information

Cooperation. Main points for today. How can altruism evolve? Group living vs. cooperation. Sociality-nocooperation. and cooperationno-sociality

Cooperation. Main points for today. How can altruism evolve? Group living vs. cooperation. Sociality-nocooperation. and cooperationno-sociality Cooperation Why is it surprising and how does it evolve Cooperation Main points for today Sociality, cooperation, mutualism, altruism - definitions Kin selection Hamilton s rule, how to calculate r Group

More information

Evolution of Diversity and Cooperation 2 / 3. Jorge M. Pacheco. Departamento de Matemática & Aplicações Universidade do Minho Portugal

Evolution of Diversity and Cooperation 2 / 3. Jorge M. Pacheco. Departamento de Matemática & Aplicações Universidade do Minho Portugal Evolution of Diversity and Cooperation 2 / 3 Jorge M. Pacheco Departamento de Matemática & Aplicações Universidade do Minho Portugal Luis Santaló School, 18 th of July, 2013 prisoner s dilemma C D C (

More information

Cultural transmission and the evolution of human behaviour: a general approach based on the Price equation

Cultural transmission and the evolution of human behaviour: a general approach based on the Price equation doi: 10.1111/jeb.12296 Cultural transmission and the evolution of human behaviour: a general approach based on the Price equation C. EL MOUDEN*, J.-B. ANDRÉ, O.MORIN &D.NETTLE *Department of Zoology &

More information

ALTRUISM AND REPUTATION cooperation within groups

ALTRUISM AND REPUTATION cooperation within groups ALTRUISM AND REPUTATION cooperation within groups Krzysztof Kułakowski Małgorzata Krawczyk Przemysław Gawroński Kraków, Poland 21-25 September, University of Warwick, UK Complex are: text, mind, community,

More information

Damon Centola.

Damon Centola. http://www.imedea.uib.es/physdept Konstantin Klemm Victor M. Eguíluz Raúl Toral Maxi San Miguel Damon Centola Nonequilibrium transitions in complex networks: a model of social interaction, Phys. Rev. E

More information

Sociobiological Approaches. What We Will Cover in This Section. Overview. Overview Evolutionary Theory Sociobiology Summary

Sociobiological Approaches. What We Will Cover in This Section. Overview. Overview Evolutionary Theory Sociobiology Summary Sociobiological Approaches What We Will Cover in This Section Overview Evolutionary Theory Sociobiology Summary 7/22/2007 Sociobiological Approaches.ppt 2 Overview Total focus on internal aspects of personality

More information

Assignment #3 Genetic Material LO: To develop a model and explain inheritance and skin color variation. EQ: Do your genes determine who you are?

Assignment #3 Genetic Material LO: To develop a model and explain inheritance and skin color variation. EQ: Do your genes determine who you are? Assignment #3 Genetic Material LO: To develop a model and explain inheritance and skin color variation. EQ: Do your genes determine who you are? AGENDA 10/20-10/21 1. Variation Model 2. Notes HOMEWORK

More information

AN EVOLUTIONARY MODEL OF RECIPROCITY

AN EVOLUTIONARY MODEL OF RECIPROCITY AN EVOLUTIONARY MODEL OF RECIPROCITY SUREN BASOV 1 Department of Economics, The University of Melbourne, Melbourne, Victoria 3010, Australia. 1 I am grateful to Peter Bardsley, Hsueh-Ling Huynh, Bob Rosenthal,

More information

Organizing Diversity Taxonomy is the discipline of biology that identifies, names, and classifies organisms according to certain rules.

Organizing Diversity Taxonomy is the discipline of biology that identifies, names, and classifies organisms according to certain rules. 1 2 3 4 5 6 7 8 9 10 Outline 1.1 Introduction to AP Biology 1.2 Big Idea 1: Evolution 1.3 Big Idea 2: Energy and Molecular Building Blocks 1.4 Big Idea 3: Information Storage, Transmission, and Response

More information

The evolutionary puzzle of human cooperation is a red herring. An influential line of theoretical modeling and analysis is directly motivated by

The evolutionary puzzle of human cooperation is a red herring. An influential line of theoretical modeling and analysis is directly motivated by The evolutionary puzzle of human cooperation is a red herring Abstract An influential line of theoretical modeling and analysis is directly motivated by describing human cooperation as an evolutionary

More information

1 low Humans Evolved

1 low Humans Evolved 1 low Humans Evolved Robert Howl IOIIIB Silk UNIVERS1. i 1 \..UK I..1 I \ Nv I Technische Unive-^itdt Darmstadt FACHDCRLICH 10 BIOLOGIE B i!. I i o t h p k -_ ScLninspilinstiafiG 10 D-6 4287 Darmstadt

More information

Major questions of evolutionary genetics. Experimental tools of evolutionary genetics. Theoretical population genetics.

Major questions of evolutionary genetics. Experimental tools of evolutionary genetics. Theoretical population genetics. Evolutionary Genetics (for Encyclopedia of Biodiversity) Sergey Gavrilets Departments of Ecology and Evolutionary Biology and Mathematics, University of Tennessee, Knoxville, TN 37996-6 USA Evolutionary

More information

ALTRUISM OR JUST SHOWING OFF?

ALTRUISM OR JUST SHOWING OFF? ALTRUISM OR JUST SHOWING OFF? Soha Sabeti ISCI 330 April 12/07 Altruism or Just Showing Off? Among the many debates regarding the evolution of altruism are suggested theories such as group selection, kin

More information

[in press, July, 2006: In W. A. Darity (Ed.), International Encyclopedia of Social Sciences (2 nd ed.). New. Genetic Determinism

[in press, July, 2006: In W. A. Darity (Ed.), International Encyclopedia of Social Sciences (2 nd ed.). New. Genetic Determinism p. 1 [in press, July, 2006: In W. A. Darity (Ed.), International Encyclopedia of Social Sciences (2 nd ed.). New York: Thomson Gale.] Genetic Determinism Lucas D. Schipper and Todd K. Shackelford Florida

More information

N-Player Prisoner s Dilemma

N-Player Prisoner s Dilemma ALTRUISM, THE PRISONER S DILEMMA, AND THE COMPONENTS OF SELECTION Abstract The n-player prisoner s dilemma (PD) is a useful model of multilevel selection for altruistic traits. It highlights the non zero-sum

More information

People talk about Fairness and Trust Stigler (1981) self-interest theory will win. Results = price tag on negative reciprocity

People talk about Fairness and Trust Stigler (1981) self-interest theory will win. Results = price tag on negative reciprocity Social Preferences Dictator, Ultimatum and Trust Games Joseph Tao-yi Wang 0/6/009 Fairness, Reciprocity, and Trust Example: Ultimatum Game Proposer: makes a take-it-or-leave-it offer Responder: accepts

More information

Study of Life. Intro to AP Biology

Study of Life. Intro to AP Biology Study of Life Intro to 2007-2008 Big Ideas Big Idea 1: The process of evolution drives the diversity and unity of life. Big Idea 2: Biological systems utilize free energy and molecular building blocks

More information

Culture Outsmarts Nature in the Evolution of Cooperation

Culture Outsmarts Nature in the Evolution of Cooperation Copyright JASSS Klaus Jaffe and Roberto Cipriani (2007) Culture Outsmarts Nature in the Evolution of Cooperation Journal of Artificial Societies and Social Simulation vol. 10, no. 1

More information

Unit of Study: Genetics, Evolution and Classification

Unit of Study: Genetics, Evolution and Classification Biology 3 rd Nine Weeks TEKS Unit of Study: Genetics, Evolution and Classification B.1) Scientific Processes. The student, for at least 40% of instructional time, conducts laboratory and field investigations

More information

Understanding and Solving Societal Problems with Modeling and Simulation

Understanding and Solving Societal Problems with Modeling and Simulation Understanding and Solving Societal Problems with Modeling and Simulation Lecture 8: The Breakdown of Cooperation ETH Zurich April 15, 2013 Dr. Thomas Chadefaux Why Cooperation is Hard The Tragedy of the

More information

Altruism, altruistic punishment and social investment

Altruism, altruistic punishment and social investment Altruism, altruistic punishment and social investment Klaus Jaffe Universidad Simón Bolívar (kjaffe@usb.ve), Caracas, Venezuela. Abstract The concept of altruism is used in very different forms by computer

More information

Essential knowledge 1.A.2: Natural selection

Essential knowledge 1.A.2: Natural selection Appendix C AP Biology Concepts at a Glance Big Idea 1: The process of evolution drives the diversity and unity of life. Enduring understanding 1.A: Change in the genetic makeup of a population over time

More information

Alana Schick , ISCI 330 Apr. 12, The Evolution of Cooperation: Putting gtheory to the Test

Alana Schick , ISCI 330 Apr. 12, The Evolution of Cooperation: Putting gtheory to the Test Alana Schick 43320027, ISCI 330 Apr. 12, 2007 The Evolution of Cooperation: Putting gtheory to the Test Evolution by natural selection implies that individuals with a better chance of surviving and reproducing

More information

Chapter 16: Evolutionary Theory

Chapter 16: Evolutionary Theory Chapter 16: Evolutionary Theory Section 1: Developing a Theory Evolution: Artificial Selection: Evolution: I. A Theory to Explain Change Over Time B. Charles Darwin C. Theory: D. Modern evolutionary theory

More information

932 Yang Wei-Song et al Vol. 12 Table 1. An example of two strategies hold by an agent in a minority game with m=3 and S=2. History Strategy 1 Strateg

932 Yang Wei-Song et al Vol. 12 Table 1. An example of two strategies hold by an agent in a minority game with m=3 and S=2. History Strategy 1 Strateg Vol 12 No 9, September 2003 cfl 2003 Chin. Phys. Soc. 1009-1963/2003/12(09)/0931-05 Chinese Physics and IOP Publishing Ltd Sub-strategy updating evolution in minority game * Yang Wei-Song(fflffΦ) a), Wang

More information

Piecing It Together. 1) The envelope contains puzzle pieces for 5 vertebrate embryos in 3 different stages of

Piecing It Together. 1) The envelope contains puzzle pieces for 5 vertebrate embryos in 3 different stages of Piecing It Together 1) The envelope contains puzzle pieces for 5 vertebrate embryos in 3 different stages of development. Lay out the pieces so that you have matched up each animal name card with its 3

More information

Understanding Emergence of Cooperation using tools from Thermodynamics

Understanding Emergence of Cooperation using tools from Thermodynamics Understanding Emergence of Cooperation using tools from Thermodynamics Pranjal Ralegankar May 13, 2018 Abstract Understanding how cooperation emerged in evolutionary biological systems is a long studied

More information

Biological institutions: an institutional perspective on biological cooperation

Biological institutions: an institutional perspective on biological cooperation Biological institutions: an institutional perspective on biological cooperation Erol Akçay Proximate mechanisms and the evolution of cooperation Department of Biology University of Pennsylvania Santa Fe

More information

AP Biology Curriculum Framework

AP Biology Curriculum Framework AP Biology Curriculum Framework This chart correlates the College Board s Advanced Placement Biology Curriculum Framework to the corresponding chapters and Key Concept numbers in Campbell BIOLOGY IN FOCUS,

More information

FAIRNESS IS A VIRTUE IN COOPERATION

FAIRNESS IS A VIRTUE IN COOPERATION European Journal of Science and Theology, February 2013, Vol.9, No.1, 61-70 FAIRNESS IS A VIRTUE IN COOPERATION Abstract Wang Yihong 127 Xinwenhua Jie Xicheng District Beijing, 100031 China (Received 19

More information

Perplexing Observations. Today: Thinking About Darwinian Evolution. We owe much of our understanding of EVOLUTION to CHARLES DARWIN.

Perplexing Observations. Today: Thinking About Darwinian Evolution. We owe much of our understanding of EVOLUTION to CHARLES DARWIN. Today: Thinking About Darwinian Evolution Part 1: Darwin s Theory Perplexing Observations Mystery of the Black Death?? What is evolution?? And what is this finch doing?!? We owe much of our understanding

More information

On the Origin of Altruism. desire that compels humans to help is called altruism by psychologists. Through this feeling,

On the Origin of Altruism. desire that compels humans to help is called altruism by psychologists. Through this feeling, Ding 1 Chunyang Ding Mrs.Boness Gifted Freshmen English 12 February 2012 On the Origin of Altruism How do humans actually behave when faced with the decision to help others? The innate desire that compels

More information

Chapters AP Biology Objectives. Objectives: You should know...

Chapters AP Biology Objectives. Objectives: You should know... Objectives: You should know... Notes 1. Scientific evidence supports the idea that evolution has occurred in all species. 2. Scientific evidence supports the idea that evolution continues to occur. 3.

More information

Unit of Study: Viruses & Microbes and Living Systems

Unit of Study: Viruses & Microbes and Living Systems Biology 4 th Nine Weeks TEKS Unit of Study: Viruses & Microbes and Living Systems B.1) Scientific processes. The student, for at least 40% of instructional time, conducts laboratory and field investigations

More information

Social semantics: altruism, cooperation, mutualism, strong reciprocity and group selection

Social semantics: altruism, cooperation, mutualism, strong reciprocity and group selection doi:10.1111/j.1420-9101.2006.01258.x MINI REVIEW Social semantics: altruism, cooperation, mutualism, strong reciprocity and group selection S. A. WEST,* A. S. GRIFFIN* & A. GARDNER* *Institute of Evolutionary

More information

EVOLUTION OF LEARNING IN FLUCTUATING ENVIRONMENTS: WHEN SELECTION FAVORS BOTH SOCIAL AND EXPLORATORY INDIVIDUAL LEARNING

EVOLUTION OF LEARNING IN FLUCTUATING ENVIRONMENTS: WHEN SELECTION FAVORS BOTH SOCIAL AND EXPLORATORY INDIVIDUAL LEARNING ORIGINAL ARTICLE doi:10.1111/j.1558-5646.2007.00313.x EVOLUTION OF LEARNING IN FLUCTUATING ENVIRONMENTS: WHEN SELECTION FAVORS BOTH SOCIAL AND EXPLORATORY INDIVIDUAL LEARNING Elhanan Borenstein, 1,2 Marcus

More information

These are my slides and notes introducing the Red Queen Game to the National Association of Biology Teachers meeting in Denver in 2016.

These are my slides and notes introducing the Red Queen Game to the National Association of Biology Teachers meeting in Denver in 2016. These are my slides and notes introducing the Red Queen Game to the National Association of Biology Teachers meeting in Denver in 2016. Thank you SSE and the Huxley Award for sending me to NABT 2016! I

More information

Other-Regarding Preferences: Theory and Evidence

Other-Regarding Preferences: Theory and Evidence Other-Regarding Preferences: Theory and Evidence June 9, 2009 GENERAL OUTLINE Economic Rationality is Individual Optimization and Group Equilibrium Narrow version: Restrictive Assumptions about Objective

More information

Mathematical Games and Random Walks

Mathematical Games and Random Walks Mathematical Games and Random Walks Alexander Engau, Ph.D. Department of Mathematical and Statistical Sciences College of Liberal Arts and Sciences / Intl. College at Beijing University of Colorado Denver

More information

Enduring understanding 1.A: Change in the genetic makeup of a population over time is evolution.

Enduring understanding 1.A: Change in the genetic makeup of a population over time is evolution. The AP Biology course is designed to enable you to develop advanced inquiry and reasoning skills, such as designing a plan for collecting data, analyzing data, applying mathematical routines, and connecting

More information

Regents Review Assignment #8-A08 Living Environment: Comet Part A Questions

Regents Review Assignment #8-A08 Living Environment: Comet Part A Questions Part A Questions 1. A student notices that fruit flies with the curlywing trait develop straight wings if kept at a temperature of 16 C, but develop curly wings if kept at 25 C. The best explanation for

More information

Brief history of The Prisoner s Dilemma (From Harman s The Price of Altruism)

Brief history of The Prisoner s Dilemma (From Harman s The Price of Altruism) Brief history of The Prisoner s Dilemma (From Harman s The Price of Altruism) 1948 The RAND Corporation. a civilian nonprofit think tank of an elite cadre of physicists, mathematicians, economists, and

More information

The Mechanisms of Evolution

The Mechanisms of Evolution The Mechanisms of Evolution Figure.1 Darwin and the Voyage of the Beagle (Part 1) 2/8/2006 Dr. Michod Intro Biology 182 (PP 3) 4 The Mechanisms of Evolution Charles Darwin s Theory of Evolution Genetic

More information

Hamilton s Rule Applied to Reciprocal Altruism

Hamilton s Rule Applied to Reciprocal Altruism Hamilton s Rule pplied to Reciprocal ltruism Jeffrey. Fletcher Systems Science Ph.D. Program Portland State University Portland, OR 9727 Email: jeff@pdx.edu bstract Reciprocal altruism and inclusive fitness

More information

Selfishness is rarely described as a group-beneficial strategy.

Selfishness is rarely described as a group-beneficial strategy. Selfishness as second-order altruism Omar Tonsi Eldakar* and David Sloan Wilson* Departments of *Biological Sciences and Anthropology, Binghamton University, Binghamton, NY 13902-6000 Edited by Brian Skyrms,

More information

Learning objectives. Evolution in Action. Chapter 8: Evolution and Natural Selection. By the end of today s topic students should be able to:

Learning objectives. Evolution in Action. Chapter 8: Evolution and Natural Selection. By the end of today s topic students should be able to: ANNOUNCEMENTS Thursday February 13, 2014 Chapter 8: Evolution and Natural Selection Dr. Traver returns next week. Movie on Tuesday afternoon What Darwin Never Knew. Midterm 2 is next week 2/20/2014 Use

More information

and just what is science? how about this biology stuff?

and just what is science? how about this biology stuff? Welcome to Life on Earth! Rob Lewis 512.775.6940 rlewis3@austincc.edu 1 The Science of Biology Themes and just what is science? how about this biology stuff? 2 1 The Process Of Science No absolute truths

More information

NGSS Example Bundles. Page 1 of 13

NGSS Example Bundles. Page 1 of 13 High School Modified Domains Model Course III Life Sciences Bundle 4: Life Diversifies Over Time This is the fourth bundle of the High School Domains Model Course III Life Sciences. Each bundle has connections

More information

A A A A B B1

A A A A B B1 LEARNING OBJECTIVES FOR EACH BIG IDEA WITH ASSOCIATED SCIENCE PRACTICES AND ESSENTIAL KNOWLEDGE Learning Objectives will be the target for AP Biology exam questions Learning Objectives Sci Prac Es Knowl

More information

Selfishness and Cooperation: Challenge for Social Life

Selfishness and Cooperation: Challenge for Social Life Studia Humana Volume 6:3 (2017), pp. 15 23 DOI: 10.1515/sh-2017-0019 Konrad Szocik Selfishness and Cooperation: Challenge for Social Life University of Information Technology and Management in Rzeszow,

More information

Journal of Artificial Societies and Social Simulation vol. 10, no

Journal of Artificial Societies and Social Simulation vol. 10, no Copyright JASSS András Németh and Károly Takács (2007) The Evolution of Altruism in Spatially Structured Populations Journal of Artificial Societies and Social Simulation vol. 10, no. 3 4 For information

More information

Chapter 1. Introduction: Biology Today. Lectures by Chris C. Romero, updated by Edward J. Zalisko

Chapter 1. Introduction: Biology Today. Lectures by Chris C. Romero, updated by Edward J. Zalisko Chapter 1 Introduction: Biology Today Lectures by Chris C. Romero, updated by Edward J. Zalisko PowerPoint Lectures for Campbell Essential Biology, Fourth Edition Eric Simon, Jane Reece, and Jean Dickey

More information

:: Slide 1 :: :: Slide 2 :: :: Slide 3 :: :: Slide 3 continued :: :: Slide 4 :: :: Slide 5 ::

:: Slide 1 :: :: Slide 2 :: :: Slide 3 :: :: Slide 3 continued :: :: Slide 4 :: :: Slide 5 :: :: Slide 1 :: :: Slide 2 :: Why Do We Say Nature and Nurture Are Intertwined? What Are the Building Blocks of Behavior? What Is the Field of Behavioral Genetics? How Does Evolution Occur? How Does Evolution

More information

Inference of Cultural Transmission Modes Based on Incomplete Information

Inference of Cultural Transmission Modes Based on Incomplete Information Wayne State University Human Biology Open Access Pre-Prints WSU Press 9-1-2015 Inference of Cultural Transmission Modes Based on Incomplete Information Bryan Wilder University of Central Florida, wilders.cfl@gmail.com

More information

Computation of Efficient Nash Equilibria for experimental economic games

Computation of Efficient Nash Equilibria for experimental economic games International Journal of Mathematics and Soft Computing Vol.5, No.2 (2015), 197-212. ISSN Print : 2249-3328 ISSN Online: 2319-5215 Computation of Efficient Nash Equilibria for experimental economic games

More information

Sorting via Screening vs. Signaling

Sorting via Screening vs. Signaling Sorting via Screening vs. Signaling - Static Frame and Experimental Evidence - Werner Güth and Fabian Winter 1. Introduction 2. Static Frame 3. Experimental Design 4. Main Findings 5. Conclusions 1. Introduction

More information

Evolutionary Dynamics and Extensive Form Games by Ross Cressman. Reviewed by William H. Sandholm *

Evolutionary Dynamics and Extensive Form Games by Ross Cressman. Reviewed by William H. Sandholm * Evolutionary Dynamics and Extensive Form Games by Ross Cressman Reviewed by William H. Sandholm * Noncooperative game theory is one of a handful of fundamental frameworks used for economic modeling. It

More information

MCDB 1111 corebio 2017 Midterm I

MCDB 1111 corebio 2017 Midterm I MCDB 1111 corebio 2017 Midterm I Name: There are 18 questions, each worth a maximum of 6 points. Multiple choice questions are worth 3 points, while explanations are worth 3 points. If you need to, use

More information

Chapter 7. Evolutionary Game Theory

Chapter 7. Evolutionary Game Theory From the book Networks, Crowds, and Markets: Reasoning about a Highly Connected World. By David Easley and Jon Kleinberg. Cambridge University Press, 2010. Complete preprint on-line at http://www.cs.cornell.edu/home/kleinber/networks-book/

More information

(Richard Dawkins) 20 Desmond Morris

(Richard Dawkins) 20 Desmond Morris 2005135694 703 Advances in Psychological Science 310028 90 B84-09 1 (Richard Dawkins) 20 Desmond Morris Konrad Lorenz Niko Tinbergen 80 20 50 (Evolutionary psychologyep) On Aggression Jerome (1966) Barkow

More information

Levels of Analysis in Ethology

Levels of Analysis in Ethology Levels of Analysis in Ethology Ethology: the study of animal behavior Example: Why do male monkeys fight? Niko Tinbergen 1963 On Aims and Methods in Ethology Answer questions about WHY behaviors occur

More information

Policing and group cohesion when resources vary

Policing and group cohesion when resources vary Anim. Behav., 996, 52, 63 69 Policing and group cohesion when resources vary STEVEN A. FRANK Department of Ecology and Evolutionary Biology, University of California at Irvine (Received 5 January 996;

More information

Globalization-polarization transition, cultural drift, co-evolution and group formation

Globalization-polarization transition, cultural drift, co-evolution and group formation - Mallorca - Spain CABDyN, Reference Saïdof Business research School, project Oxford, &/or name November of conference 7 Globalization-polarization transition, cultural drift, co-evolution and group formation

More information

NGSS Example Bundles. Page 1 of 23

NGSS Example Bundles. Page 1 of 23 High School Conceptual Progressions Model III Bundle 2 Evolution of Life This is the second bundle of the High School Conceptual Progressions Model Course III. Each bundle has connections to the other

More information

Revisiting The evolution of reciprocity in sizable groups : Continuous reciprocity in the repeated -person prisoner s dilemma

Revisiting The evolution of reciprocity in sizable groups : Continuous reciprocity in the repeated -person prisoner s dilemma Revisiting The evolution of reciprocity in sizable groups : Continuous reciprocity in the repeated -person prisoner s dilemma Masanori Takezawa, Michael E. Price To cite this version: Masanori Takezawa,

More information

The Effects of Extra-Somatic Weapons on the Evolution of Human Cooperation towards Non-Kin

The Effects of Extra-Somatic Weapons on the Evolution of Human Cooperation towards Non-Kin The Effects of Extra-Somatic Weapons on the Evolution of Human Cooperation towards Non-Kin Tim Phillips 1 *, Jiawei Li 2, Graham Kendall 2,3 1 Independent Researcher, Birmingham, United Kingdom, 2 Automated

More information

Population Games and Evolutionary Dynamics

Population Games and Evolutionary Dynamics Population Games and Evolutionary Dynamics (MIT Press, 200x; draft posted on my website) 1. Population games 2. Revision protocols and evolutionary dynamics 3. Potential games and their applications 4.

More information

The Ways of Altruism 1

The Ways of Altruism 1 The Ways of Altruism 1 Gualtiero Piccinini, 1 University Blvd, Department of Philosophy, University of Missouri St. Louis, St. Louis, MO 6121, piccininig@umsl.edu, http://www.umsl.edu/~piccininig/ Armin

More information

AIS 2016 AIS 2014 AIS 2015

AIS 2016 AIS 2014 AIS 2015 Interne score ASMF journal subject category Economics or Business, Finance on ISI Web of Knowledge TI selected Marketing or OR journal Journal 2013 2014 2015 2016 AVERAGE Advances in Water Resources 1,246

More information

The Evolution of Sex Chromosomes through the. Baldwin Effect

The Evolution of Sex Chromosomes through the. Baldwin Effect The Evolution of Sex Chromosomes through the Baldwin Effect Larry Bull Computer Science Research Centre Department of Computer Science & Creative Technologies University of the West of England, Bristol

More information

Nature s Art Village

Nature s Art Village Nature s Art Village Educational Field Trip Programs Guide To: College, Career & Civic Life C3 Framework For Social Studies State Standards Grades 3 through 5 All That Glitters Children journey back in

More information

Social Studies 3 Vocabulary Cards. century. History 1. period of 100 years

Social Studies 3 Vocabulary Cards. century. History 1. period of 100 years century History 1 period of 100 years chronological History 1 in order of time decade History 1 period of 10 years timeline History 1 list of important events in the order in which they happened year History

More information

Jei. Balancing Inertia, Innovation, and Imitation in Complex Environments. Geoffrey M. Hodgson and Thorbjørn Knudsen

Jei. Balancing Inertia, Innovation, and Imitation in Complex Environments. Geoffrey M. Hodgson and Thorbjørn Knudsen Jei JOURNAL OF ECONOMIC ISSUES Vol. XL No. 2 June 2006 Balancing Inertia, Innovation, and Imitation in Complex Environments Geoffrey M. Hodgson and Thorbjørn Knudsen Since Thorstein Veblen, perennial themes

More information

Introduction to Biological Anthropology: Notes 18 The evolution of cooperation: Altruism and kin selection Copyright Bruce Owen 2011 It was not

Introduction to Biological Anthropology: Notes 18 The evolution of cooperation: Altruism and kin selection Copyright Bruce Owen 2011 It was not Introduction to Biological Anthropology: Notes 18 The evolution of cooperation: Altruism and kin selection Copyright Bruce Owen 2011 It was not difficult to understand how selection can affect mating and

More information

Introduction to Biological Anthropology: Notes 17 The evolution of cooperation: Altruism and kin selection Copyright Bruce Owen 2010 It was not

Introduction to Biological Anthropology: Notes 17 The evolution of cooperation: Altruism and kin selection Copyright Bruce Owen 2010 It was not Introduction to Biological Anthropology: Notes 17 The evolution of cooperation: Altruism and kin selection Copyright Bruce Owen 2010 It was not difficult to understand how selection can affect mating and

More information

The evolution of norms

The evolution of norms ARTICLE IN PRESS Journal of Theoretical Biology 241 (2006) 233 240 www.elsevier.com/locate/yjtbi The evolution of norms F.A.C.C. Chalub a,, F.C. Santos b, J.M. Pacheco c a Centro de Matemática e Aplicac-ões

More information

Applying evolutionary theory to human behaviour: past differences and current debates. Citation:

Applying evolutionary theory to human behaviour: past differences and current debates. Citation: Applying evolutionary theory to human behaviour: past differences and current debates Gillian R. Brown 1 and Peter J. Richerson 2 1 School of Psychology and Neuroscience, University of St Andrews, U.K.

More information